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{{Short description|Species of gastropod}}
{{Speciesbox
| image = SeaangelClione limacina by NOAA.jpg
| image_caption = ''Clione limacina''
| taxon = Clione limacina
| authority = ([[Constantine Phipps, 2nd Baron Mulgrave|Phipps]], 1774) <ref>Phipps, C.J., 1774. ''A voyage towards the North Pole undertaken by His Majesty's Command 1773'' : i–viii, 1–253</ref>
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== Distribution ==
''Clione limacina'' is found in cold waters of the Arctic Ocean and North Atlantic Ocean, ranging south at least to the [[Sargasso Sea]].<ref>Mileikovsky S.A. (1970) Breeding and larval distribution of the pteropod ''Clione limacina'' in the North Atlantic, Subarctic and North Pacific Oceans. ''Marine Biology'' 6(4): 317–334.</ref><ref name=Yamazaki2017>{{cite journal | last1 = Yamazaki | first1 = Tomoyasu | last2 = Kuwahara | first2 = Takashi | year = 2017 | title = A new species of Clione distinguished from sympatric C. limacina (Gastropoda: Gymnosomata) in the southern Okhotsk Sea, Japan, with remarks on the taxonomy of the genus | journal = Journal of Molluscan Studies | volume = 83 | issue = 1 | pages = 19–26 | issn=0260-1230 | doi=10.1093/mollus/eyw032| doi-access = free }}</ref> There are three other species in the genus, which formerly were included in ''C. limacina'' (either as subspecies, variants or subpopulations). These are ''[[Clione elegantissima|C. elegantissima]]'' of the cold North Pacific (at least north to the [[Gulf of Alaska]]; the [[Beaufort Sea]] is inhabited by ''C. limacina''), ''[[Clione okhotensis|C. okhotensis]]'' of the [[Okhotsk Sea]] (where it overlaps with ''C. elegantissima''), and ''[[Clione antarctica|C. antarctica]]'' of Antarctic waters.<ref name=Yamazaki2017/>
 
[[File:Clione limacina by NOAA.jpg|thumb|left|''Clione limacina'']]
 
== Description ==
There are two [[subspecies]] that differentiate in body length.<ref name="Böer 2005"/> The northern subspecies lives in colder water, matures at {{convert|3|cm|in|abbr=on}} and can reach a size of {{convert|7|-|8.5|cm|in|abbr=on}}.<ref name="Böer 2005"/><ref name=Kallevik>Kallevik, I.H.F. (2013). ''[http://munin.uit.no/bitstream/handle/10037/5858/thesis.pdf Alternative prey choice in the pteropod ''Clione limacina'' (Gastropoda) studied by DNA-based methods.] {{Webarchive|url=https://web.archive.org/web/20160304023256/http://munin.uit.no/bitstream/handle/10037/5858/thesis.pdf |date=2016-03-04 }}'' Biology Field of study - Arctic Marine Ecology and Resource Biology. Bio-3950 (60 ECT). The University Center in Svalbard.</ref> This makes it by far the largest [[sea angel]].<ref name=Yamazaki2017/> In comparison, the size of the southern subspecies is {{convert|1.2|cm|in|abbr=on|1}},<ref name="Böer 2005"/> ''C. elegantissima'' is up to {{convert|3|cm|in|abbr=on}}, ''C. okhotensis'' up to {{convert|0.8|cm|in|abbr=on|1}},<ref name=Yamazaki2017/> and ''C. antarctica'' up to {{convert|3|cm|in|abbr=on}}.<ref name=McClintock1997>{{cite book | editor1=McClintock, J.B. | editor2=B.J. Baker | year=1997 | title=Marine Chemical Ecology | url=https://archive.org/details/marinechemicalec00mccl | url-access=limited | publisher=CRC Press | page=[https://archive.org/details/marinechemicalec00mccl/page/n227 215] | isbn=0-8493-9064-8 }}</ref>
 
''C. limacina'' swims by beating its two wings to move upwards or maintain itself at a constant depth. To keep itself upright during swimming, it uses two [[statocyst]] gravity-sensing organs that correct it to an upright posture using its tail.<ref name="Deliagina Arshavsky Orlovsky 1998 pp. 172–175">{{cite journal |last1=Deliagina |first1=Tatiana G. |last2=Arshavsky |first2=Yuri I. |last3=Orlovsky |first3=Grigori N. |date=1998 |title=Control of spatial orientation in a mollusc |journal=Nature |volume=393 |issue=6681 |pages=172–175 |doi=10.1038/30251 |pmid=9603520 |bibcode=1998Natur.393..172D |issn=0028-0836}}</ref>
The [[Neuroscience|neurobiology]] of this [[pteropod]] has been studied in detail.
 
== Ecology ==
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== Cultural significance ==
While sea angels are relatively obscure in Western countries, they are extremely well-known in [[Japanese culture]]. As a result, two creatures from the Pokémon franchise, Manaphy and Phione, are based on the clione.<ref name=phione>{{cite web|url=https://bulbanews.bulbagarden.net/wiki/On_the_Origin_of_Species:_Manaphy_and_Phione|title=On the Origin of Species: Manaphy and Phione|date=9 February 2018 }}</ref>
 
==References==