Mountain nyala: Difference between revisions

Mountain nyala
	Male
	Female; both Bale Mountains National Park, Ethiopia
Conservation status
	; Endangered (IUCN 3.1)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Artiodactyla
Family:	Bovidae
Subfamily:	Bovinae
Genus:	Tragelaphus
Species:	T. buxtoni
Binomial name
	Tragelaphus buxtoni; (Lydekker, 1910)

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The mountain nyala (Amharic: የተራራ ኒዮላ) (Tragelaphus buxtoni) or balbok, is a large antelope found in high altitude woodlands in a small part of central Ethiopia. It is a monotypic species (without any identified subspecies) first described by English naturalist Richard Lydekker in 1910. The males are typically 120–135 cm (47–53 in) tall while females stand 90–100 cm (35–39 in) at the shoulder. Males weigh 180–300 kg (400–660 lb) and females weigh 150–200 kg (330–440 lb). The coat is grey to brown, marked with two to five poorly defined white strips extending from the back to the underside, and a row of six to ten white spots. White markings are present on the face, throat and legs as well. Males have a short dark erect crest, about 10 cm (3.9 in) high, running along the middle of the back. Only males possess horns.

The mountain nyala are shy and elusive towards human beings. Four to five individuals may congregate for short intervals of time to form small herds. Males are not territorial. Primarily a browser, the mountain nyala may switch to grazing occasionally. Females start mating at two years of age, and males too become sexually mature by that time. Gestation lasts for eight to nine months, after which a single calf is born. The lifespan of a mountain nyala is around 15 to 20 years.

The typical habitat for the mountain nyala is composed of montane woodlands at an altitude of 3,000–3,400 m (9,800–11,200 ft). Human settlement and large livestock population have forced the animal to occupy heath forests at an altitude of above 3,400 m (11,200 ft). Mountain nyala are endemic to the Ethiopian highlands east of the Rift Valley, between 6°N and 10°N. Up to half of the total population of the mountain nyala occurs in the 200 km² (77 sq mi) area of Gaysay, in the northern part of the Bale Mountains National Park. The mountain nyala has been classified under the Endangered category of the International Union for Conservation of Nature and Natural Resources (IUCN). Their influence on Ethiopian culture is notable, with the mountain nyala being featured on the obverse of Ethiopian ten cents coins.

Taxonomy

	Giant eland

	Common eland

Greater kudu

Mountain nyala

	Bongo

	Sitatunga

	Cape bushbuck

	Harnessed bushbuck

Nyala

Lesser kudu

Phylogenetic relationships of the mountain nyala from combined analysis of all molecular data (Willows-Munro et.al. 2005)

The mountain nyala was first described by English naturalist Richard Lydekker in 1910. Its scientific name is Tragelaphus buxtoni. The species has been classified under the genus Tragelaphus of the family Bovidae.^[2] The mountain nyala was first brought to England in 1908 by Major Ivor Buxton, after whom the species has been named. Major Buxton, who had returned from a hunting trip in Ethiopia, first presented a specimen he had shot to British taxidermist Rowland Ward, who later on notified Lydekker about the discovery. It was the last large antelope to be discovered in Africa.^[3]^[4] The spiral horns and white markings on the face and across the flanks established it as a species of Tragelaphus.^[5] Lydekker believed that the species was related more closely to the nyala (T. angasii) than to the two kudu species (then placed under the genus Strepsiceros). In fact, mountain nyala's similarity to both Tragelaphus and Strepsiceros led to the merger of the latter into the former genus.^[6]^[7]

In 2005, Sandi Willows-Munro of the University of KwaZulu-Natal (Durban) carried out a mitochondrial analysis of the nine Tragelaphus species. mtDNA and nDNA data were compared. The results showed that the mountain nyala is sister taxon to bushbuck (T. scriptus), sitatunga (T. spekii) and bongo (T. eurycerus) in the mitochondrial tree (phylogenetic relationships obtained using mitochondrial data). However, in the nuclear tree, bushbuck, mountain nyala and sitatunga plus the bongo are three equal branches. Hence the mountain nyala forms a monophyletic clade with these three species.^[8]^[9] The greater kudu clade split from the clade formed by mountain nyala, bongo, sitatunga and bushbuck approximately 8.6 million years ago.^[10]^[11]

Description

The mountain nyala is a large sexually dimorphic bovid. The head-and-body length is approximately 240–260 cm (94–102 in) in males and 190–200 cm (75–79 in) in females. The males are typically 120–135 cm (47–53 in) tall while females stand 90–100 cm (35–39 in) at the shoulder. Males weigh 180–300 kg (400–660 lb) and females weigh 150–200 kg (330–440 lb). The bushy tail reaches the heel, and is 20–25 cm (7.9–9.8 in) long.^[12] The sensitive ears are large and lined with white hair.^[5]

The coat is grey to brown, marked with two to five poorly defined white strips extending from the back to the underside, and a row of six to ten white spots. White markings are present on the face, throat and legs as well. The white chevron between the eyes and the white patch on the throat are among the most conspicuous markings. The chest and the rump are white. The lower part of the legs are pale on the inside and clear white spots are present just above the hooves. The coat is fawn brown in male juveniles, and grows darker with age, eventually becoming charcoal in old bulls. Males have a short dark erect crest, about 10 cm (3.9 in) high, running along the middle of the back, from the neck to the tail.^[5] Coat texture may vary from smooth to rough, probably by season. Females resemble the red deer hind in size and proportions. Female juveniles are a bright rufous, and old females are as grey as young males. Females have two pairs of inguinal nipples.^[12]

Only males possess horns, whose maximum recorded length is 188 cm (74 in). Horns appear as cream-coloured nubs at around six months and start growing in a spiral pattern, reaching full growth by two years.^[5] The horns have only one or two spirals, though a few males have been reported to have two-and-a-half turns. The final form might vary in different males – the horns could form well-defined spirals, or could diverge in a structure similar to a lyre, resembling the horns of an impala, but with the final spiral incomplete. Growth rings are visible on the horn sheaths, but the annual patterns might be difficult to comprehend. Though the horns might wear out with age, the cream colour of the tips persists.^[5]^[13]

The mountain nyala resembles the greater kudu in that both have an array of white spots along their flanks and possess spiral horns. However, the greater kudu can be told apart from the mountain nyala due to the former's greater height and paler colour. Moreover, the horns in greater kudu have two to three spirals, and the tips are farther apart. Another species similar to the mountain nyala is the nyala, but the latter can be easily distinguished from the former due to its smaller size and a fringe of long hair along its throat and neck. The horns of nyala, though very similar, are slender and narrower.^[12]

Ecology and behaviour

The mountain nyala are shy and elusive towards human beings. They spend the night on the edges of forests, feeding part of the time. To avoid human disturbance, they choose to navigate at night. They come out in morning and late afternoon to browse in grasslands. They generally seek cover in woodlands and heather thickets when it is very hot or cold. Feeding may occur in midday, but is interspersed with resting intervals. They tend to come out when it is overcast or raining. In the dry season (November to March), given the dilapidated state of grasslands, the mountain nyala travel up to the wooden areas rich in ericaceous heath.^[14]

Four to five individuals may congregate for short intervals of time to form small herds. However, groups containing as many as 100 individuals have been reported from the Bale Zone, composed of several family units moving in and out periodically. Size and longevity of such large herds depends upon the season, habitat type and the time of the day. Female and juvenile groups have adult females accompanied by a calf of her previous year and another of the current year. These groups are led by adult males, depending on the presence of oestrus. Bachelor herds are formed by non-dominant adult bulls and young males, consisting of up to 13 individuals. Mixed sex groups may also be formed. Old bulls tend to lead a solitary life, though they may occasionally visit female herds looking for females in oestrus. Dominance hierarchies are observed in both sexes.^[15]

Males are not territorial, and have home ranges covering 15–20 km² (5.8–7.7 sq mi) in the wet season. Females and juveniles, on the other hand, occupy much smaller ranges of about 5 km² (1.9 sq mi) in the wet season. Dry season ranges are much larger than the wet season ones for both sexes.^[16] Males often wrestle using their horns.^[13] Though usually silent, the mountain nyala may "cough" noticing a potential threat, or utter a low bark if the threat is more serious.^[3] The leopard is the main predator of the mountain nyala.^[13]

Diet

Primarily a browser, the mountain nyala may switch to grazing occasionally. It feeds on low-height herbs, bushes, shrubs and general foliage. They might even eat lichens, ferns and aquatic plants. Grasses are specially preferred during the early wet season. They pick up fallen leaves and use their horns to reach higher branches. Favoured species include Artemisia afra (African wormwood), Hypericum revolutum, Kniphofia foliosa, Solanum sessilistellatum and Hagenia abyssinica leaves. They prefer Alchemilla rotti, Helichrysum splendidum and lower foliage of Lobelia rhynchopetalum.^[14]^[16]

Reproduction

Females start mating at two years of age, and males too become sexually mature by that time. Sexual dimorphism, larger home ranges for males, and a female-biased population indicate polygyny in the species.^[13] The mountain nyala breed throughout the year, but the peak occurs in December. Prior to and during the breeding season the adult males tend to get associated with mixed sex herds. Bulls continuously follow receptive females and test their vulvas. Three to four males may seek a single female, and, if equally ranked in hierarchy, may engage in circling displays. In these displays the males move very slowly and stiffly, with the crest on the back erect and the tail raised. They are less likely to engage in aggressive fights, which, if they happen, last only for a brief period. Flehmen follows vulva-testing.^[3]^[12]

Gestation lasts for eight to nine months, after which a single calf is born. In the Bale Mountains National Park, births occur throughout the year, but peak from September to November. Calves remain in cover for the first few weeks after birth. The calves remain close to their mothers for nearly two years. The young females may get pregnant by then.^[3] The young males, as they mature by two years, are challenged by other males and driven out of their herds.^[5] The lifespan of a mountain nyala is around 15 to 20 years.^[13]

Habitat and distribution

The typical habitat for the mountain nyala is composed of montane woodlands at an altitude of 3,000–3,400 m (9,800–11,200 ft). The vegetation generally includes African juniper (Juniperus procera), Afrocarpus gracilior, and Olea at lower levels and Hagenia abyssinica, juniper, and Hypericum revolutum in the upper reaches. They often visit the edges of montane grasslands at 2,800–3,100 m (9,200–10,200 ft), which is covered with Artemisia afra, Kniphofia, and evergreen Hypericum species.^[12]

Nowadays the continuous blocks of woodland habitat have been reduced to a series of pockets, interspersed in large cultivated lands. Human settlement and large livestock population have forced the animal to occupy heath forests at an altitude of above 3,400 m (11,200 ft) rich in Erica arborea, Erica trimera, Hypericum, Euphorbia, and Helichrysum species. They may even be forced into afroalpine grasslands containing Alchemilla and Festuca species, at an altitude of up to 4,300 m (14,100 ft). In the eastern borders of its range, the mountain nyala has usually been observed to occupy areas at lower altitudes of about 1,800–2,400 m (5,900–7,900 ft).^[12]

Mountain nyala are endemic to the Ethiopian Highlands east of the Rift Valley, between 6°N and 10°N. Their former range was from Mount Gara Muleta in the east to Shashamene and the northern Bale Zone to the south. Up to half of the total population of the mountain nyala occurs in the 200 km² (77 sq mi) area of Gaysay, in the northern part of the Bale Mountains National Park. Smaller relict populations occur in Chercher, mountains such as Chilalo in Arsi Zone, and the western Bale.^[17] A study identified an area of 39.38 km² (15.20 sq mi) on the southern escarpment of the Bale Mountains as the most sustainable habitat for the mountain nyala.^[18]

Threats and conservation

Major threats to the survival of the mountain nyala include illegal hunting, habitat destruction, encroachment by livestock, predation of calves by dogs, expansion of montane cultivation and construction at high altitudes.^[19]^[20] The animal is extensively hunted for its horns and meat. The meat is utilised in local medicine and for making nipples for traditional milk bottles. Impact of trophy hunting programs is obscure, and current trophy hunting quotas may be unsustainable in the long term although, if well regulated, trophy hunting could play an important role in the long term management of this species.^[1]^[17]

The mountain nyala has been classified under the Endangered category of the International Union for Conservation of Nature and Natural Resources (IUCN). It has not been listed under the Washington Convention (CITES). Though legal protection has been fully ensured for the species, the implementation has not been so effective. In 1991, there was widespread unrest in Ethiopia during which several mountain nyala were killed and the population in the Bale Mountains National Park fell to 150.^[21]^[22]^[23] After the Bale Mountains National Park, the small Kuni-Muktar Mountain Nyala Sanctuary was established in 1990 was established as a second protected area. But by 1996 the sanctuary had been so severely affected by deforestation, poaching, cultivation and gully erosion, that the populations soon disappeared from there.^[1]^[17]

In the 1960s, the population was estimated at 7,000 to 8,000 (and perhaps up to 12,500). This declined sharply to 2,000 to 4,000 individuals in the 1980s. The population is decreasing even now. The mountain nyala is feared to be extinct in the eastern and southern extremes of its range, though a few probably survive in Chiro (Asebe Teferi) and the border between Bale and Sidamo, south of Kofele. Apart from Gaysay about 80 to 120 mountain nyala occur in the rest of the Bale Mountains National Park, and a few hundred occur in the areas in the vicinity of the national park.^[1]^[17]

Relationship with humans

The influence of the mountain nyala, as well as the nyala, on Ethiopian culture can not be ignored. Although nyala is a South African word, several prominent businesses such as Nyala Motors use it in their names. The nyala and mountain nyala often regularly appear on walls, commercial products and tourist curios.^[5] The mountain nyala is featured on the obverse of the Ethiopian ten cents coin.^[5]^[24]^[25]

Mountain nyala are hunted by humans, for instance, by the Oromo people. The Oromo do not hunt these animals in general, except for a day close to Easter, when all able-bodied men set out on horses for a hunt.^[5]

References

^ ^a ^b ^c ^d IUCN SSC Antelope Specialist Group (2017) [errata version of 2016 assessment]. "Tragelaphus buxtoni". IUCN Red List of Threatened Species. 2016: e.T22046A115164345.
^ Grubb, P. (2005). "Order Artiodactyla". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 697–698. ISBN 978-0-8018-8221-0. OCLC 62265494.
^ ^a ^b ^c ^d Brown, L. H. (1969). "Observations on the status, habitat and behaviour of the mountain nyala Tragelaphus buxtoni in Ethiopia". Mammalia. 33 (4): 545–97. doi:10.1515/mamm.1969.33.4.545. S2CID 84219997.
^ Briggs, P. (2015). Ethiopia. USA: Bradt Travel Guides. p. 16. ISBN 978-1-78477-132-4.
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ Malcolm, J.; Evangelista, P.H. (2005). "The range and status of the mountain nyala" (PDF): 1–43. Archived from the original (PDF) on 2016-01-28. {{cite journal}}: Cite journal requires |journal= (help)
^ Lydekker, R. (June 1911). "On the Mountain Nyala, Tragelaphus buxtoni*". Proceedings of the Zoological Society of London. 81 (2): 348–53. doi:10.1111/j.1096-3642.1911.tb01934.x.
^ Evangelista, P.; Waltermire, R.; Swartzinski, P. (2007). "A profile of the mountain nyala (Tragelaphus buxtoni)". African Indaba: 1–47.
^ Willows-Munro, S.; Robinson, T. J.; Matthee, C. A. (June 2005). "Utility of nuclear DNA intron markers at lower taxonomic levels: Phylogenetic resolution among nine Tragelaphus spp". Molecular Phylogenetics and Evolution. 35 (3): 624–36. Bibcode:2005MolPE..35..624W. doi:10.1016/j.ympev.2005.01.018. PMID 15878131.
^ Groves, C.; Grubb, P. (2011). Ungulate Taxonomy. Baltimore, Maryland: Johns Hopkins University Press. pp. 126, 137. ISBN 978-1-4214-0093-8.
^ Ropiquet, A. (2006). "Etude des radiations adaptatives au sein des Antilopinae (Mammalia, Bovidae)". Ph.D. Thesis, Université Paris. 6 (1–247).
^ Hassanin, A.; Delsuc, F.; Ropiquet, A.; Hammer, C.; Jansen van Vuuren, B.; Matthee, C.; Ruiz-Garcia, M.; Catzeflis, F.; Areskoug, V.; Nguyen, T.T.; Couloux, A. (2012). "Pattern and timing of diversification of Cetartiodactyla (Mammalia, Laurasiatheria), as revealed by a comprehensive analysis of mitochondrial genomes". Comptes Rendus Biologies. 335 (1): 32–50. doi:10.1016/j.crvi.2011.11.002. PMID 22226162.
^ ^a ^b ^c ^d ^e ^f Kingdon, J. (2013). Mammals of Africa. London: Bloomsbury. pp. 159–62. ISBN 978-1-4081-2257-0.
^ ^a ^b ^c ^d ^e Aleman, M. "Tragelaphus buxtoni Mountain nyala". Animal Diversity Web. University of Michigan Museum of Zoology. Retrieved 21 January 2016.
^ ^a ^b Gagnon, M.; Chew, A. E. (May 2000). "Dietary preferences in extant African Bovidae". Journal of Mammalogy. 81 (2): 490–511. doi:10.1644/1545-1542(2000)081<0490:DPIEAB>2.0.CO;2.
^ Hillman, J.C.; Hillman, S.M. (1987). "The mountain nyala Tragelaphus buxtoni and the Simien fox Canis simensis in the Bale Mountains National Park". Walia. 10: 3–6.
^ ^a ^b Hillman, J. C. (24 April 2009). "Conservation in Bale Mountains National Park, Ethiopia". Oryx. 20 (2): 89–94. doi:10.1017/S0030605300026314.
^ ^a ^b ^c ^d East, R.; IUCN/SSC Antelope Specialist Group (1999). African Antelope Database 1998. Gland, Switzerland: The IUCN Species Survival Commission. pp. 130–1. ISBN 978-2-8317-0477-7.
^ Evangelista, P. H.; Norman, J.; Berhanu, L.; Kumar, S.; Alley, Nathaniel (2008). "Predicting habitat suitability for the endemic mountain nyala in Ethiopia". Wildlife Research. 35 (5): 409–16. CiteSeerX 10.1.1.570.6821. doi:10.1071/WR07173.
^ Mamo, Y.; Bekele, A. (2011). "Human and livestock encroachments into the habitat of Mountain Nyala (Tragelaphus buxtoni) in the Bale Mountains National Park, Ethiopia". Tropical Ecology. 52 (3): 265–73.
^ Bussmann, R.W.; Worede, A.; Swartzinsky, P.; Evangelista, P. (2011). "Plant use in Odo-Bulu and Demaro, Bale region, Ethiopia". Journal of Ethnobiology and Ethnomedicine. 7 (1): 28. doi:10.1186/1746-4269-7-28. PMC 3196683. PMID 21943288.
^ Woldegebriel, G.K. (1996). "The status of mountain nyala (Tragelaphus buxtoni) in Bale Mountains National Park 1986–1994". Walia. 17: 27–37.
^ Mamo, Y.; Pinard, M.A.; Bekele, A. (2010). "Demography and dynamics of mountain nyala Tragelaphus buxtoni in the Bale Mountains National Park, Ethiopia". Current Zoology. 56 (6): 660–9. doi:10.1093/czoolo/56.6.660.
^ Refera, B.; Bekele, A. (2004). "Population status and structure of mountain nyala in the Bale Mountains National Park, Ethiopia". African Journal of Ecology. 42 (1): 1–7. Bibcode:2004AfJEc..42....1R. doi:10.1111/j.0141-6707.2004.00437.x.
^ Cuhaj, G.S. (2015). 2016 Standard Catalog of World Coins 2001-Date. Iola: Krause Publications. p. 495. ISBN 978-1-4402-4410-0.
^ Balletto, B.L. (1995). Spectrum guide to Ethiopia. Camerapix Publishers. p. 295.

External links

ITIS entry on Tragelaphus buxtoni

[iucn-1] IUCN SSC Antelope Specialist Group (2017) [errata version of 2016 assessment]. "Tragelaphus buxtoni". IUCN Red List of Threatened Species. 2016: e.T22046A115164345.

[MSW3-2] Grubb, P. (2005). "Order Artiodactyla". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 697–698. ISBN 978-0-8018-8221-0. OCLC 62265494.

[brown-3] Brown, L. H. (1969). "Observations on the status, habitat and behaviour of the mountain nyala Tragelaphus buxtoni in Ethiopia". Mammalia. 33 (4): 545–97. doi:10.1515/mamm.1969.33.4.545. S2CID 84219997.

[briggs-4] Briggs, P. (2015). Ethiopia. USA: Bradt Travel Guides. p. 16. ISBN 978-1-78477-132-4.

[malcolm-5] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ Malcolm, J.; Evangelista, P.H. (2005). "The range and status of the mountain nyala" (PDF): 1–43. Archived from the original (PDF) on 2016-01-28. {{cite journal}}: Cite journal requires |journal= (help)

[lydekker-6] Lydekker, R. (June 1911). "On the Mountain Nyala, Tragelaphus buxtoni*". Proceedings of the Zoological Society of London. 81 (2): 348–53. doi:10.1111/j.1096-3642.1911.tb01934.x.

[evangelista2-7] Evangelista, P.; Waltermire, R.; Swartzinski, P. (2007). "A profile of the mountain nyala (Tragelaphus buxtoni)". African Indaba: 1–47.

[cladogram-8] Willows-Munro, S.; Robinson, T. J.; Matthee, C. A. (June 2005). "Utility of nuclear DNA intron markers at lower taxonomic levels: Phylogenetic resolution among nine Tragelaphus spp". Molecular Phylogenetics and Evolution. 35 (3): 624–36. Bibcode:2005MolPE..35..624W. doi:10.1016/j.ympev.2005.01.018. PMID 15878131.

[groves-9] Groves, C.; Grubb, P. (2011). Ungulate Taxonomy. Baltimore, Maryland: Johns Hopkins University Press. pp. 126, 137. ISBN 978-1-4214-0093-8.

[ropiquet-10] Ropiquet, A. (2006). "Etude des radiations adaptatives au sein des Antilopinae (Mammalia, Bovidae)". Ph.D. Thesis, Université Paris. 6 (1–247).

[hassanin-11] Hassanin, A.; Delsuc, F.; Ropiquet, A.; Hammer, C.; Jansen van Vuuren, B.; Matthee, C.; Ruiz-Garcia, M.; Catzeflis, F.; Areskoug, V.; Nguyen, T.T.; Couloux, A. (2012). "Pattern and timing of diversification of Cetartiodactyla (Mammalia, Laurasiatheria), as revealed by a comprehensive analysis of mitochondrial genomes". Comptes Rendus Biologies. 335 (1): 32–50. doi:10.1016/j.crvi.2011.11.002. PMID 22226162.

[kingdon-12] ^ ^a ^b ^c ^d ^e ^f Kingdon, J. (2013). Mammals of Africa. London: Bloomsbury. pp. 159–62. ISBN 978-1-4081-2257-0.

[adw-13] Aleman, M. "Tragelaphus buxtoni Mountain nyala". Animal Diversity Web. University of Michigan Museum of Zoology. Retrieved 21 January 2016.

[gagnon-14] Gagnon, M.; Chew, A. E. (May 2000). "Dietary preferences in extant African Bovidae". Journal of Mammalogy. 81 (2): 490–511. doi:10.1644/1545-1542(2000)081<0490:DPIEAB>2.0.CO;2.

[hillman2-15] Hillman, J.C.; Hillman, S.M. (1987). "The mountain nyala Tragelaphus buxtoni and the Simien fox Canis simensis in the Bale Mountains National Park". Walia. 10: 3–6.

[hillman-16] Hillman, J. C. (24 April 2009). "Conservation in Bale Mountains National Park, Ethiopia". Oryx. 20 (2): 89–94. doi:10.1017/S0030605300026314.

[east-17] East, R.; IUCN/SSC Antelope Specialist Group (1999). African Antelope Database 1998. Gland, Switzerland: The IUCN Species Survival Commission. pp. 130–1. ISBN 978-2-8317-0477-7.

[evangelista-18] Evangelista, P. H.; Norman, J.; Berhanu, L.; Kumar, S.; Alley, Nathaniel (2008). "Predicting habitat suitability for the endemic mountain nyala in Ethiopia". Wildlife Research. 35 (5): 409–16. CiteSeerX 10.1.1.570.6821. doi:10.1071/WR07173.

[mamo-19] Mamo, Y.; Bekele, A. (2011). "Human and livestock encroachments into the habitat of Mountain Nyala (Tragelaphus buxtoni) in the Bale Mountains National Park, Ethiopia". Tropical Ecology. 52 (3): 265–73.

[bussmann-20] Bussmann, R.W.; Worede, A.; Swartzinsky, P.; Evangelista, P. (2011). "Plant use in Odo-Bulu and Demaro, Bale region, Ethiopia". Journal of Ethnobiology and Ethnomedicine. 7 (1): 28. doi:10.1186/1746-4269-7-28. PMC 3196683. PMID 21943288.

[21] Woldegebriel, G.K. (1996). "The status of mountain nyala (Tragelaphus buxtoni) in Bale Mountains National Park 1986–1994". Walia. 17: 27–37.

[mamo2-22] Mamo, Y.; Pinard, M.A.; Bekele, A. (2010). "Demography and dynamics of mountain nyala Tragelaphus buxtoni in the Bale Mountains National Park, Ethiopia". Current Zoology. 56 (6): 660–9. doi:10.1093/czoolo/56.6.660.

[refera-23] Refera, B.; Bekele, A. (2004). "Population status and structure of mountain nyala in the Bale Mountains National Park, Ethiopia". African Journal of Ecology. 42 (1): 1–7. Bibcode:2004AfJEc..42....1R. doi:10.1111/j.0141-6707.2004.00437.x.

[cuhaj-24] Cuhaj, G.S. (2015). 2016 Standard Catalog of World Coins 2001-Date. Iola: Krause Publications. p. 495. ISBN 978-1-4402-4410-0.

[balletto-25] Balletto, B.L. (1995). Spectrum guide to Ethiopia. Camerapix Publishers. p. 295.

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@@ Line 1: / Line 1: @@
+{{Short description|Species of antelope}}
 {{good article}}
 {{Speciesbox
@@ Line 8: / Line 9: @@
 | status = EN
 | status_system = IUCN3.1
-| status_ref = <ref name=iucn>{{cite iucn | authors = Sillero-Zubiri, C. | title = ''Tragelaphus buxtoni'' | volume= 20016 | page = e.T22046A9351382 | year = 2016 | doi = 10.2305/IUCN.UK.2008.RLTS.T22046A9351382.en }}</ref>
+| status_ref = <ref name=iucn>{{cite iucn | author = IUCN SSC Antelope Specialist Group | title = ''Tragelaphus buxtoni'' |errata=2017 | volume= 2016 | page = e.T22046A115164345 | year = 2016}}</ref>
 | taxon = Tragelaphus buxtoni
 | authority = ([[Richard Lydekker|Lydekker]], 1910)
@@ Line 14: / Line 15: @@
 [[File:Mountain nyala (Tragelaphus buxtoni) juvenile.jpg|thumb|juvenile]]
 [[File:Mountain nyala (Tragelaphus buxtoni) female.jpg|thumb|young male]]
-The '''mountain nyala''' (''Tragelaphus buxtoni'') or '''balbok''' is an [[antelope]] found in high altitude [[woodland]] in a small part of central [[Ethiopia]]. It is a [[monotypic]] species (without any identified [[subspecies]]) first [[Scientific description|described]] by English naturalist [[Richard Lydekker]] in 1910. The males are typically {{convert|120|-|135|cm|in|abbr=on}} tall while females stand {{convert|90|-|100|cm|in|abbr=on}} at the shoulder. Males weigh {{convert|180|-|300|kg|lb|abbr=on}} and females weigh {{convert|150|-|200|kg|lb|abbr=on}}. The coat is grey to brown, marked with two to five poorly defined white strips extending from the back to the underside, and a row of six to ten white spots. White markings are present on the face, throat and legs as well. Males have a short dark erect crest, about {{convert|10|cm|in|abbr=on}} high, running along the middle of the back. Only males possess [[Horn (anatomy)|horns]].
+The '''mountain nyala''' ([[Amharic]]: የተራራ ኒዮላ) ('''''Tragelaphus buxtoni''''') or '''balbok''', is a large [[antelope]] found in high altitude [[woodland]]s in a small part of central [[Ethiopia]]. It is a [[monotypic]] species (without any identified [[subspecies]]) first [[Scientific description|described]] by English naturalist [[Richard Lydekker]] in 1910. The males are typically {{cvt|120|-|135|cm|in}} tall while females stand {{cvt|90|-|100|cm|in}} at the shoulder. Males weigh {{cvt|180|-|300|kg|lb}} and females weigh {{cvt|150|-|200|kg|lb}}. The coat is grey to brown, marked with two to five poorly defined white strips extending from the back to the underside, and a row of six to ten white spots. White markings are present on the face, throat and legs as well. Males have a short dark erect crest, about {{cvt|10|cm|in}} high, running along the middle of the back. Only males possess [[Horn (anatomy)|horn]]s.
 The mountain nyala are shy and elusive towards human beings. Four to five individuals may congregate for short intervals of time to form small herds. Males are not [[Territory (animal)|territorial]]. Primarily a browser, the mountain nyala may switch to grazing occasionally. Females start mating at two years of age, and males too become [[sexually mature]] by that time. [[Gestation]] lasts for eight to nine months, after which a single calf is born. The lifespan of a mountain nyala is around 15 to 20 years.
-The typical habitat for the mountain nyala is composed of montane woodlands at an altitude of {{convert|3000|-|3400|m|ft|abbr=on}}. Human settlement and large livestock population have forced the animal to occupy heath forests at an altitude of above {{convert|3400|m|ft|abbr=on}}. Mountain nyala are endemic to the [[Ethiopian highlands]] east of the [[Main Ethiopian Rift|Rift Valley]], between 6°N and 10°N. Up to half of the total population of the mountain nyala occurs in the {{convert|200|sqkm|sqmi|abbr=on}} area of Gaysay, in the northern part of the [[Bale Mountains National Park]]. The mountain nyala has been classified under the [[Endangered]] category of the [[International Union for Conservation of Nature and Natural Resources]] (IUCN). Their influence on Ethiopian culture is notable, with the mountain nyala being featured on the obverse of Ethiopian ten cents coins.
+The typical habitat for the mountain nyala is composed of montane woodlands at an altitude of {{cvt|3000|-|3400|m|ft}}. Human settlement and large livestock population have forced the animal to occupy heath forests at an altitude of above {{cvt|3400|m|ft}}. Mountain nyala are endemic to the [[Ethiopian highlands]] east of the [[Main Ethiopian Rift|Rift Valley]], between 6°N and 10°N. Up to half of the total population of the mountain nyala occurs in the {{cvt|200|sqkm|sqmi}} area of Gaysay, in the northern part of the [[Bale Mountains National Park]]. The mountain nyala has been classified under the [[Endangered]] category of the [[International Union for Conservation of Nature and Natural Resources]] (IUCN). Their influence on Ethiopian culture is notable, with the mountain nyala being featured on the obverse of Ethiopian ten cents coins.
 ==Taxonomy==
@@ Line 44: / Line 45: @@
                                   |1=[[Bongo (antelope)|Bongo]]
                                   |2=[[Sitatunga]]}}
-                           |2=[[Cape bushbuck]] }} }} }} }} }} }} }} }}
+                           |2={{Clade
+                           |1={{clade
+                              |1={{clade
+                                  |1=[[Cape bushbuck]]
+|2=[[Harnessed bushbuck]] }} }} }} }} }} }} }} }}
          |2=[[Nyala]]}}
-|2=[[Lesser kudu]] }} }} }}
+|2=[[Lesser kudu]] }} }} }} }} }} }}
-The mountain nyala was first [[Scientific description|described]] by English naturalist [[Richard Lydekker]] in 1910. Its scientific name is ''Tragelaphus buxtoni''. The species has been classified under the genus ''[[Tragelaphus]]'' of the family [[Bovidae]].<ref name=MSW3>{{MSW3 Artiodactyla|pages=697–698|id=14200728}}</ref>  The mountain nyala was first brought to [[England]] in 1908 by Major Ivor Buxton, after whom the species has been named. Major Buxton, who had returned from a hunting trip in [[Ethiopia]], first presented a specimen he had shot to British taxidermist [[Rowland Ward]], who later on notified Lydekker about the discovery. It was the last large antelope to be discovered in Africa.<ref name=brown>{{cite journal|last1=Brown|first1=L. H.|title=Observations on the status, habitat and behaviour of the mountain nyala ''Tragelaphus buxtoni'' in Ethiopia|journal=Mammalia|date=1969|volume=33|issue=4|pages=545–97|doi=10.1515/mamm.1969.33.4.545|s2cid=84219997}}</ref><ref name=briggs>{{cite book|last1=Briggs|first1=P.|title=Ethiopia|date=2015|publisher=Bradt Travel Guides|location=USA|isbn=978-1-78477-132-4|page=16}}</ref> The spiral horns and white markings on the face and across the flanks established it as a species of ''Tragelaphus''.<ref name=malcolm>{{cite journal|last1=Malcolm|first1=J.|last2=Evangelista|first2=P.H.|title=The range and status of the mountain nyala|date=2005|pages=1–43|url=http://w.ewca.gov.et/sites/default/files/MalcolmEvangelistaMountainNyalaReport_0.pdf|url-status=dead|archive-url=https://web.archive.org/web/20160128041346/http://w.ewca.gov.et/sites/default/files/MalcolmEvangelistaMountainNyalaReport_0.pdf|archive-date=2016-01-28}}</ref> Lydekker believed that the species was related more closely to the [[nyala]] (''T. angasii'') than to the two kudu species (then placed under the genus ''Strepsiceros''). In fact, mountain nyala's similarity to both ''Tragelaphus'' and ''Strepsiceros'' led to the merger of the latter into the former genus.<ref name=lydekker>{{cite journal|last1=Lydekker|first1=R.|title=On the Mountain Nyala, ''Tragelaphus buxtoni''*|journal=Proceedings of the Zoological Society of London|date=June 1911|volume=81|issue=2|pages=348–53|doi=10.1111/j.1096-3642.1911.tb01934.x}}</ref><ref name=evangelista2>{{cite journal|last1=Evangelista|first1=P.|last2=Waltermire|first2=R.|last3=Swartzinski|first3=P.|title=A profile of the mountain nyala (''Tragelaphus buxtoni'')|journal=African Indaba|date=2007|pages=1–47}}</ref>
+The mountain nyala was first [[Scientific description|described]] by English naturalist [[Richard Lydekker]] in 1910. Its scientific name is ''Tragelaphus buxtoni''. The species has been classified under the genus ''[[Tragelaphus]]'' of the family [[Bovidae]].<ref name=MSW3>{{MSW3 Artiodactyla|pages=697–698|id=14200728}}</ref>  The mountain nyala was first brought to [[England]] in 1908 by Major Ivor Buxton, after whom the species has been named. Major Buxton, who had returned from a hunting trip in [[Ethiopia]], first presented a specimen he had shot to British taxidermist [[Rowland Ward]], who later on notified Lydekker about the discovery. It was the last large antelope to be discovered in Africa.<ref name=brown>{{cite journal|last1=Brown|first1=L. H.|title=Observations on the status, habitat and behaviour of the mountain nyala ''Tragelaphus buxtoni'' in Ethiopia|journal=Mammalia|date=1969|volume=33|issue=4|pages=545–97|doi=10.1515/mamm.1969.33.4.545|s2cid=84219997}}</ref><ref name=briggs>{{cite book|last1=Briggs|first1=P.|title=Ethiopia|date=2015|publisher=Bradt Travel Guides|location=USA|isbn=978-1-78477-132-4|page=16}}</ref> The spiral horns and white markings on the face and across the flanks established it as a species of ''Tragelaphus''.<ref name=malcolm>{{cite journal|last1=Malcolm|first1=J.|last2=Evangelista|first2=P.H.|title=The range and status of the mountain nyala|date=2005|pages=1–43|url=http://w.ewca.gov.et/sites/default/files/MalcolmEvangelistaMountainNyalaReport_0.pdf|url-status=dead|archive-url=https://web.archive.org/web/20160128041346/http://w.ewca.gov.et/sites/default/files/MalcolmEvangelistaMountainNyalaReport_0.pdf|archive-date=2016-01-28}}</ref> Lydekker believed that the species was related more closely to the [[nyala]] (''T. angasii'') than to the two kudu species (then placed under the genus ''Strepsiceros''). In fact, mountain nyala's similarity to both ''Tragelaphus'' and ''Strepsiceros'' led to the merger of the latter into the former genus.<ref name=lydekker>{{cite journal|last1=Lydekker|first1=R.|title=On the Mountain Nyala, ''Tragelaphus buxtoni''*|journal=Proceedings of the Zoological Society of London|date=June 1911|volume=81|issue=2|pages=348–53|doi=10.1111/j.1096-3642.1911.tb01934.x|url=https://www.biodiversitylibrary.org/part/72349 }}</ref><ref name=evangelista2>{{cite journal|last1=Evangelista|first1=P.|last2=Waltermire|first2=R.|last3=Swartzinski|first3=P.|title=A profile of the mountain nyala (''Tragelaphus buxtoni'')|journal=African Indaba|date=2007|pages=1–47}}</ref>
-In 2005, Sandi Willows-Munro of the [[University of KwaZulu-Natal]] ([[Durban]]) carried out a [[mitochondria]]l analysis of the nine ''Tragelaphus'' species. [[mtDNA]] and [[nDNA]] data were compared. The results showed that the mountain nyala is sister [[taxon]] to [[kéwel]] (''T. scriptus''), [[sitatunga]] (''T. spekii'') and [[Bongo (antelope)|bongo]] (''T. eurycerus'') in the mitochondrial tree (phylogenetic relationships obtained using mitochondrial data). However, in the nuclear tree, kéwel, mountain nyala and sitatunga plus the bongo are three equal branches. Hence the mountain nyala forms a [[monophyletic]] [[clade]] with these three species.<ref name=cladogram>{{cite journal|last1=Willows-Munro|first1=S.|last2=Robinson|first2=T. J.|last3=Matthee|first3=C. A.|title=Utility of nuclear DNA intron markers at lower taxonomic levels: Phylogenetic resolution among nine ''Tragelaphus'' spp.|journal=Molecular Phylogenetics and Evolution|date=June 2005|volume=35|issue=3|pages=624–36|doi=10.1016/j.ympev.2005.01.018|pmid=15878131}}</ref><ref name=groves>{{cite book|last1=Groves|first1=C.|last2=Grubb|first2=P.|title=Ungulate Taxonomy|date=2011|publisher=Johns Hopkins University Press|location=Baltimore, Maryland|isbn=978-1-4214-0093-8|pages=126, 137}}</ref> The greater kudu clade split from the clade formed by mountain nyala, bongo, sitatunga and kéwel approximately 8.6 million years ago.<ref name="ropiquet">{{cite journal|last1=Ropiquet|first1=A.|title=Etude des radiations adaptatives au sein des Antilopinae (Mammalia, Bovidae)|journal=Ph.D. Thesis, Université Paris|date=2006|volume=6|issue=1–247}}</ref><ref name="hassanin">{{cite journal|last1=Hassanin|first1=A.|last2=Delsuc|first2=F.|last3=Ropiquet|first3=A.|last4=Hammer|first4=C.|last5=Jansen van Vuuren|first5=B.|last6=Matthee|first6=C.|last7=Ruiz-Garcia|first7=M.|last8=Catzeflis|first8=F.|last9=Areskoug|first9=V.|last10=Nguyen|first10=T.T.|last11=Couloux|first11=A.|title=Pattern and timing of diversification of Cetartiodactyla (Mammalia, Laurasiatheria), as revealed by a comprehensive analysis of mitochondrial genomes|journal=Comptes Rendus Biologies|date=2012|volume=335|issue=1|pages=32–50|doi=10.1016/j.crvi.2011.11.002|pmid=22226162}}</ref>
+In 2005, Sandi Willows-Munro of the [[University of KwaZulu-Natal]] ([[Durban]]) carried out a [[mitochondria]]l analysis of the nine ''Tragelaphus'' species. [[mtDNA]] and [[nDNA]] data were compared. The results showed that the mountain nyala is sister [[taxon]] to [[bushbuck]] (''T. scriptus''), [[sitatunga]] (''T. spekii'') and [[Bongo (antelope)|bongo]] (''T. eurycerus'') in the mitochondrial tree (phylogenetic relationships obtained using mitochondrial data). However, in the nuclear tree, bushbuck, mountain nyala and sitatunga plus the bongo are three equal branches. Hence the mountain nyala forms a [[monophyletic]] [[clade]] with these three species.<ref name=cladogram>{{cite journal|last1=Willows-Munro|first1=S.|last2=Robinson|first2=T. J.|last3=Matthee|first3=C. A.|title=Utility of nuclear DNA intron markers at lower taxonomic levels: Phylogenetic resolution among nine ''Tragelaphus'' spp.|journal=Molecular Phylogenetics and Evolution|date=June 2005|volume=35|issue=3|pages=624–36|doi=10.1016/j.ympev.2005.01.018|pmid=15878131|bibcode=2005MolPE..35..624W }}</ref><ref name=groves>{{cite book|last1=Groves|first1=C.|last2=Grubb|first2=P.|title=Ungulate Taxonomy|date=2011|publisher=Johns Hopkins University Press|location=Baltimore, Maryland|isbn=978-1-4214-0093-8|pages=126, 137}}</ref> The greater kudu clade split from the clade formed by mountain nyala, bongo, sitatunga and bushbuck approximately 8.6 million years ago.<ref name="ropiquet">{{cite journal|last1=Ropiquet|first1=A.|title=Etude des radiations adaptatives au sein des Antilopinae (Mammalia, Bovidae)|journal=Ph.D. Thesis, Université Paris|date=2006|volume=6|issue=1–247}}</ref><ref name="hassanin">{{cite journal|last1=Hassanin|first1=A.|last2=Delsuc|first2=F.|last3=Ropiquet|first3=A.|last4=Hammer|first4=C.|last5=Jansen van Vuuren|first5=B.|last6=Matthee|first6=C.|last7=Ruiz-Garcia|first7=M.|last8=Catzeflis|first8=F.|last9=Areskoug|first9=V.|last10=Nguyen|first10=T.T.|last11=Couloux|first11=A.|title=Pattern and timing of diversification of Cetartiodactyla (Mammalia, Laurasiatheria), as revealed by a comprehensive analysis of mitochondrial genomes|journal=Comptes Rendus Biologies|date=2012|volume=335|issue=1|pages=32–50|doi=10.1016/j.crvi.2011.11.002|pmid=22226162|url=https://comptes-rendus.academie-sciences.fr/biologies/articles/10.1016/j.crvi.2011.11.002/ }}</ref>
 ==Description==
-The mountain nyala is a large [[sexually dimorphic]] bovid. The head-and-body length is approximately {{convert|240|-|260|cm|in|abbr=on}} in males and {{convert|190|-|200|cm|in|abbr=on}} in females. The males are typically {{convert|120|-|135|cm|in|abbr=on}} tall while females stand {{convert|90|-|100|cm|in|abbr=on}} at the shoulder. Males weigh {{convert|180|-|300|kg|lb|abbr=on}} and females weigh {{convert|150|-|200|kg|lb|abbr=on}}. The bushy tail reaches the heel, and is {{convert|20|-|25|cm|in|abbr=on}} long.<ref name=kingdon/> The sensitive ears are large and lined with white hair.<ref name=malcolm/>
+The mountain nyala is a large [[sexually dimorphic]] bovid. The head-and-body length is approximately {{cvt|240|-|260|cm|in}} in males and {{cvt|190|-|200|cm|in}} in females. The males are typically {{cvt|120|-|135|cm|in}} tall while females stand {{cvt|90|-|100|cm|in}} at the shoulder. Males weigh {{cvt|180|-|300|kg|lb}} and females weigh {{cvt|150|-|200|kg|lb}}. The bushy tail reaches the heel, and is {{cvt|20|-|25|cm|in}} long.<ref name=kingdon/> The sensitive ears are large and lined with white hair.<ref name=malcolm/>
-The coat is grey to brown, marked with two to five poorly defined white strips extending from the back to the underside, and a row of six to ten white spots. White markings are present on the face, throat and legs as well. The white [[Chevron (anatomy)|chevron]] between the eyes and the white patch on the throat are among the most conspicuous markings. The chest and the rump are white. The lower part of the legs are pale on the inside and clear white spots are present just above the hooves. The coat is fawn brown in male juveniles, and grows darker with age, eventually becoming charcoal in old bulls. Males have a short dark erect crest, about {{convert|10|cm|in|abbr=on}} high, running along the middle of the back, from the neck to the tail.<ref name=malcolm/> Coat texture may vary from smooth to rough, probably by season. Females resemble the [[red deer]] hind in size and proportions. Female juveniles are a bright rufous, and old females are as grey as young males. Females have two pairs of [[Inguinal lymph node|inguinal]] [[Mammary gland|nipple]]s.<ref name=kingdon/>
+The coat is grey to brown, marked with two to five poorly defined white strips extending from the back to the underside, and a row of six to ten white spots. White markings are present on the face, throat and legs as well. The white [[Chevron (anatomy)|chevron]] between the eyes and the white patch on the throat are among the most conspicuous markings. The chest and the rump are white. The lower part of the legs are pale on the inside and clear white spots are present just above the hooves. The coat is fawn brown in male juveniles, and grows darker with age, eventually becoming charcoal in old bulls. Males have a short dark erect crest, about {{cvt|10|cm|in}} high, running along the middle of the back, from the neck to the tail.<ref name=malcolm/> Coat texture may vary from smooth to rough, probably by season. Females resemble the [[red deer]] hind in size and proportions. Female juveniles are a bright rufous, and old females are as grey as young males. Females have two pairs of [[Inguinal lymph node|inguinal]] [[Mammary gland|nipple]]s.<ref name=kingdon/>
-Only males possess [[Horn (anatomy)|horns]], whose maximum recorded length is {{convert|188|cm|in|abbr=on}}. Horns appear as cream-coloured nubs at around six months and start growing in a spiral pattern, reaching full growth by two years.<ref name=malcolm/> The horns have only one or two spirals, though a few males have been reported to have two-and-a-half turns. The final form might vary in different males – the horns could form well-defined spirals, or could diverge in a structure similar to a lyre, resembling the horns of an [[impala]], but with the final spiral incomplete. Growth rings are visible on the horn sheaths, but the annual patterns might be difficult to comprehend. Though the horns might wear out with age, the cream colour of the tips persists.<ref name=malcolm/><ref name=adw>{{cite web|last1=Aleman|first1=M.|title=''Tragelaphus buxtoni'' Mountain nyala|url=http://animaldiversity.org/accounts/Tragelaphus_buxtoni/|website=Animal Diversity Web|publisher=University of Michigan Museum of Zoology|access-date=21 January 2016}}</ref>
+Only males possess [[Horn (anatomy)|horns]], whose maximum recorded length is {{cvt|188|cm|in}}. Horns appear as cream-coloured nubs at around six months and start growing in a spiral pattern, reaching full growth by two years.<ref name=malcolm/> The horns have only one or two spirals, though a few males have been reported to have two-and-a-half turns. The final form might vary in different males – the horns could form well-defined spirals, or could diverge in a structure similar to a lyre, resembling the horns of an [[impala]], but with the final spiral incomplete. Growth rings are visible on the horn sheaths, but the annual patterns might be difficult to comprehend. Though the horns might wear out with age, the cream colour of the tips persists.<ref name=malcolm/><ref name=adw>{{cite web|last1=Aleman|first1=M.|title=''Tragelaphus buxtoni'' Mountain nyala|url=http://animaldiversity.org/accounts/Tragelaphus_buxtoni/|website=Animal Diversity Web|publisher=University of Michigan Museum of Zoology|access-date=21 January 2016}}</ref>
 The mountain nyala resembles the [[greater kudu]] in that both have an array of white spots along their flanks and possess spiral horns. However, the greater kudu can be told apart from the mountain nyala due to the former's greater height and paler colour. Moreover, the horns in greater kudu have two to three spirals, and the tips are farther apart. Another species similar to the mountain nyala is the nyala, but the latter can be easily distinguished from the former due to its smaller size and a fringe of long hair along its throat and neck. The horns of nyala, though very similar, are slender and narrower.<ref name=kingdon/>
@@ Line 67: / Line 72: @@
 Four to five individuals may congregate for short intervals of time to form small herds. However, groups containing as many as 100 individuals have been reported from the [[Bale Zone]], composed of several family units moving in and out periodically. Size and longevity of such large herds depends upon the season, habitat type and the time of the day. Female and juvenile groups have adult females accompanied by a calf of her previous year and another of the current year. These groups are led by adult males, depending on the presence of [[oestrus]]. Bachelor herds are formed by non-dominant adult bulls and young males, consisting of up to 13 individuals. Mixed sex groups may also be formed. Old bulls tend to lead a solitary life, though they may occasionally visit female herds looking for females in oestrus. [[Dominance hierarchies]] are observed in both sexes.<ref name=hillman2>{{cite journal|last1=Hillman|first1=J.C.|last2=Hillman|first2=S.M.|title=The mountain nyala ''Tragelaphus buxtoni'' and the Simien fox ''Canis simensis'' in the Bale Mountains National Park|journal=Walia|date=1987|volume=10|pages=3–6}}</ref>
-Males are not [[Territory (animal)|territorial]], and have [[home range]]s covering {{convert|15|-|20|sqkm|sqmi|abbr=on}} in the wet season. Females and juveniles, on the other hand, occupy much smaller ranges of about {{convert|5|sqkm|sqmi|abbr=on}} in the wet season. Dry season ranges are much larger than the wet season ones for both sexes.<ref name=hillman>{{cite journal|last1=Hillman|first1=J. C.|title=Conservation in Bale Mountains National Park, Ethiopia|journal=Oryx|date=24 April 2009|volume=20|issue=2|pages=89–94|doi=10.1017/S0030605300026314}}</ref> Males often wrestle using their horns.<ref name=adw/> Though usually silent, the mountain nyala may "cough" noticing a potential threat, or utter a low bark if the threat is more serious.<ref name=brown/> The [[leopard]] is the main predator of the mountain nyala.<ref name=adw/>
+Males are not [[Territory (animal)|territorial]], and have [[home range]]s covering {{cvt|15|-|20|sqkm|sqmi|abbr=on}} in the wet season. Females and juveniles, on the other hand, occupy much smaller ranges of about {{cvt|5|sqkm|sqmi|abbr=on}} in the wet season. Dry season ranges are much larger than the wet season ones for both sexes.<ref name=hillman>{{cite journal|last1=Hillman|first1=J. C.|title=Conservation in Bale Mountains National Park, Ethiopia|journal=Oryx|date=24 April 2009|volume=20|issue=2|pages=89–94|doi=10.1017/S0030605300026314|doi-access=free}}</ref> Males often wrestle using their horns.<ref name=adw/> Though usually silent, the mountain nyala may "cough" noticing a potential threat, or utter a low bark if the threat is more serious.<ref name=brown/> The [[leopard]] is the main predator of the mountain nyala.<ref name=adw/>
 ===Diet===
-Primarily a browser, the mountain nyala may switch to grazing occasionally. It feeds on low-height herbs, bushes, shrubs and general foliage. They might even eat [[lichen]]s, [[fern]]s and aquatic plants. Grasses are specially preferred during the early wet season. They pick up fallen leaves and use their horns to reach higher branches. Favoured species include ''[[Artemsia afra]]'' (sage brush), ''[[Hypericum revolutum]]'', ''[[Kniphofia foliosa]]'', ''[[Solanum sessilistellatum]]'' and ''[[Hagenia abyssinica]]'' leaves. They prefer ''[[Alchemilla rotti]]'', ''[[Helichrysum splendidum]]'' and lower foliage of ''[[Lobelia rynchopetalum]]''.<ref name=gagnon/><ref name=hillman/>
+Primarily a browser, the mountain nyala may switch to grazing occasionally. It feeds on low-height herbs, bushes, shrubs and general foliage. They might even eat [[lichen]]s, [[fern]]s and aquatic plants. Grasses are specially preferred during the early wet season. They pick up fallen leaves and use their horns to reach higher branches. Favoured species include ''[[Artemisia afra]]'' (African wormwood), ''[[Hypericum revolutum]]'', ''[[Kniphofia foliosa]]'', ''[[Solanum sessilistellatum]]'' and ''[[Hagenia abyssinica]]'' leaves. They prefer ''[[Alchemilla rotti]]'', ''[[Helichrysum splendidum]]'' and lower foliage of ''[[Lobelia rhynchopetalum]]''.<ref name=gagnon/><ref name=hillman/>
 ===Reproduction===
-Females start mating at two years of age, and males too become [[sexually mature]] by that time. Sexual dimorphism, larger home ranges for males, and a female-biased population indicate [[polygyny]] in the species.<ref name=adw/> The mountain nyala breed throughout the year, but the peak occurs in December. Prior to and during the breeding season the adult males tend to get associated with mixed sex herds. Bulls continuously follow receptive females and test their [[vulva]]s. Three to four males may seek a single female, and, if equally ranked in hierarchy, may engage in circling [[Display (zoology)|display]]s. In these displays the males move very slowly and stiffly, with the crest on the back erect and the tail raised. They are less likely to engage in aggressive fights, which, if they happen, last only for a brief period. [[Flehmen response|Flehmen]] follows vulva-testing.<ref name=brown/><ref name=kingdon/>
+Females start mating at two years of age, and males too become [[sexually mature]] by that time. Sexual dimorphism, larger home ranges for males, and a female-biased population indicate [[Polygyny in animals|polygyny]] in the species.<ref name=adw/> The mountain nyala breed throughout the year, but the peak occurs in December. Prior to and during the breeding season the adult males tend to get associated with mixed sex herds. Bulls continuously follow receptive females and test their [[vulva]]s. Three to four males may seek a single female, and, if equally ranked in hierarchy, may engage in circling [[Display (zoology)|display]]s. In these displays the males move very slowly and stiffly, with the crest on the back erect and the tail raised. They are less likely to engage in aggressive fights, which, if they happen, last only for a brief period. [[Flehmen response|Flehmen]] follows vulva-testing.<ref name=brown/><ref name=kingdon/>
 [[Gestation]] lasts for eight to nine months, after which a single calf is born. In the [[Bale Mountains National Park]], births occur throughout the year, but peak from September to November.  Calves remain in cover for the first few weeks after birth. The calves remain close to their mothers for nearly two years. The young females may get pregnant by then.<ref name=brown/> The young males, as they mature by two years, are challenged by other males and driven out of their herds.<ref name=malcolm/> The lifespan of a mountain nyala is around 15 to 20 years.<ref name=adw/>
@@ Line 79: / Line 84: @@
 ==Habitat and distribution==
 [[File:Tragelaphus buxtoni map.svg|thumbnail|The range of mountain nyala is confined within Ethiopia.]]
-The typical habitat for the mountain nyala is composed of montane woodlands at an altitude of {{convert|3000|-|3400|m|ft|abbr=on}}. The vegetation generally includes ''[[Juniper]]'', ''[[Podocarpus]]'' and ''[[Olea]]'' at lower levels and ''[[Haegenia]]'', ''Juniper'' and ''[[Hypericum]]'' in the upper reaches. They often visit the edges of montane grasslands at {{convert|2800|-|3100|m|ft|abbr=on}}, which is covered with ''Artemesia afra'', ''[[Kniphofia]]'' and evergreen ''Hypericum'' species.<ref name=kingdon>{{cite book|last1=Kingdon|first1=J.|title=Mammals of Africa|date=2013|publisher=Bloomsbury|location=London|isbn=978-1-4081-2257-0|pages=159–62}}</ref>
+The typical habitat for the mountain nyala is composed of [[Ethiopian montane grasslands and woodlands|montane woodlands]] at an altitude of {{convert|3000|-|3400|m|ft|abbr=on}}. The vegetation generally includes [[Juniperus procera|African juniper]] (''Juniperus procera''), ''[[Afrocarpus gracilior]]'', and ''[[Olea]]'' at lower levels and ''[[Hagenia abyssinica]]'', juniper, and ''[[Hypericum revolutum]]'' in the upper reaches. They often visit the edges of montane grasslands at {{convert|2800|-|3100|m|ft|abbr=on}}, which is covered with ''[[Artemisia afra]]'', ''[[Kniphofia]]'', and evergreen ''Hypericum'' species.<ref name=kingdon>{{cite book|last1=Kingdon|first1=J.|title=Mammals of Africa|date=2013|publisher=Bloomsbury|location=London|isbn=978-1-4081-2257-0|pages=159–62}}</ref>
-Nowadays the continuous blocks of woodland habitat have been reduced to a series of pockets, interspersed in large cultivated lands. Human settlement and large livestock population have forced the animal to occupy heath forests at an altitude of above {{convert|3400|m|ft|abbr=on}} rich in ''[[Erica]]'', ''[[Phillippia]]'', ''Hypericum'', ''[[Euphorbia]]'' and ''[[Helichrysum]]'' species. They may even be forced into afromontane grasslands containing ''[[Alchemilla]]'' and ''[[Festuca]]'' species, at an altitude of up to {{convert|4300|m|ft|abbr=on}}. In the eastern borders of its range, the mountain nyala has usually been observed to occupy areas at lower altitudes of about {{convert|1800|-|2400|m|ft|abbr=on}}.<ref name=kingdon/>
+Nowadays the continuous blocks of woodland habitat have been reduced to a series of pockets, interspersed in large cultivated lands. Human settlement and large livestock population have forced the animal to occupy heath forests at an altitude of above {{convert|3400|m|ft|abbr=on}} rich in ''[[Erica arborea]]'', ''[[Erica trimera]]'', ''Hypericum'', ''[[Euphorbia]]'', and ''[[Helichrysum]]'' species. They may even be forced into [[Ethiopian montane moorlands|afroalpine grasslands]] containing ''[[Alchemilla]]'' and ''[[Festuca]]'' species, at an altitude of up to {{convert|4300|m|ft|abbr=on}}. In the eastern borders of its range, the mountain nyala has usually been observed to occupy areas at lower altitudes of about {{convert|1800|-|2400|m|ft|abbr=on}}.<ref name=kingdon/>
-Mountain nyala are endemic to the [[Ethiopian highlands]] east of the [[Main Ethiopian Rift|Rift Valley]], between 6°N and 10°N. Their former range was from [[Mount Gara Muleta]] in the east to [[Shashamene (woreda)|Shashamene]] and the northern Bale Zone to the south. Up to half of the total population of the mountain nyala occurs in the {{convert|200|sqkm|sqmi|abbr=on}} area of Gaysay, in the northern part of the Bale Mountains National Park. Smaller relict populations occur in [[Chercher province|Chercher]], mountains such as [[Mount Chilalo|Chilalo]] in [[Arsi Zone]], and the western Bale.<ref name=east>{{cite book|last1=East|first1=R.|last2=IUCN/SSC Antelope Specialist Group|title=African Antelope Database 1998|date=1999|publisher=The IUCN Species Survival Commission|location=Gland, Switzerland|isbn=978-2-8317-0477-7|pages=130–1}}</ref> A study identified an area of {{convert|39.38|sqkm|sqmi|abbr=on}} on the southern escarpment of the Bale Mountains as the most sustainable habitat for the mountain nyala.<ref name=evangelista>{{cite journal|last1=Evangelista|first1=P. H.|last2=Norman|first2=J.|last3=Berhanu|first3=L.|last4=Kumar|first4=S.|last5=Alley|first5=Nathaniel|title=Predicting habitat suitability for the endemic mountain nyala in Ethiopia|journal=Wildlife Research|date=2008|volume=35|issue=5|pages=409–16|doi=10.1071/WR07173|citeseerx=10.1.1.570.6821}}</ref>
+Mountain nyala are endemic to the [[Ethiopian Highlands]] east of the [[Main Ethiopian Rift|Rift Valley]], between 6°N and 10°N. Their former range was from [[Mount Gara Muleta]] in the east to [[Shashamene (woreda)|Shashamene]] and the northern Bale Zone to the south. Up to half of the total population of the mountain nyala occurs in the {{convert|200|sqkm|sqmi|abbr=on}} area of Gaysay, in the northern part of the Bale Mountains National Park. Smaller relict populations occur in [[Chercher province|Chercher]], mountains such as [[Mount Chilalo|Chilalo]] in [[Arsi Zone]], and the western Bale.<ref name=east>{{cite book|last1=East|first1=R.|last2=IUCN/SSC Antelope Specialist Group|title=African Antelope Database 1998|date=1999|publisher=The IUCN Species Survival Commission|location=Gland, Switzerland|isbn=978-2-8317-0477-7|pages=130–1}}</ref> A study identified an area of {{convert|39.38|sqkm|sqmi|abbr=on}} on the southern escarpment of the Bale Mountains as the most sustainable habitat for the mountain nyala.<ref name=evangelista>{{cite journal|last1=Evangelista|first1=P. H.|last2=Norman|first2=J.|last3=Berhanu|first3=L.|last4=Kumar|first4=S.|last5=Alley|first5=Nathaniel|title=Predicting habitat suitability for the endemic mountain nyala in Ethiopia|journal=Wildlife Research|date=2008|volume=35|issue=5|pages=409–16|doi=10.1071/WR07173|citeseerx=10.1.1.570.6821}}</ref>
 ==Threats and conservation==
+[[File:Mountain Nyala diorama (21982315228).jpg|thumb|Taxidermied specimens at the [[American Museum of Natural History]]]]
-Major threats to the survival of the mountain nyala include illegal hunting, habitat destruction, encroachment by livestock, predation of calves by dogs, expansion of montane cultivation and construction at high altitudes.<ref name=mamo>{{cite journal|last1=Mamo|first1=Y.|last2=Bekele|first2=A.|title=Human and livestock encroachments into the habitat of Mountain Nyala (''Tragelaphus buxtoni'') in the Bale Mountains National Park, Ethiopia|journal=Tropical Ecology|date=2011|volume=52|issue=3|pages=265–73}}</ref><ref name=bussmann>{{cite journal|last1=Bussmann|first1=R.W.|last2=Worede|first2=A.|last3=Swartzinsky|first3=P.|last4=Evangelista|first4=P.|title=Plant use in Odo-Bulu and Demaro, Bale region, Ethiopia|journal=Journal of Ethnobiology and Ethnomedicine|date=2011|volume=7|issue=1|pages=28|doi=10.1186/1746-4269-7-28|pmid=21943288|pmc=3196683}}</ref> The animal is extensively hunted for its horns and meat. The meat is utilised in local medicine and for making nipples for traditional milk bottles. Impact of trophy hunting programs is obscure, and current trophy hunting quotas may be unsustainable in the long term although, if well regulated, trophy hunting could play an important role in the long term management of this species.<ref name = iucn/><ref name=east/>
+Major threats to the survival of the mountain nyala include illegal hunting, habitat destruction, encroachment by livestock, predation of calves by dogs, expansion of montane cultivation and construction at high altitudes.<ref name=mamo>{{cite journal|last1=Mamo|first1=Y.|last2=Bekele|first2=A.|title=Human and livestock encroachments into the habitat of Mountain Nyala (''Tragelaphus buxtoni'') in the Bale Mountains National Park, Ethiopia|journal=Tropical Ecology|date=2011|volume=52|issue=3|pages=265–73}}</ref><ref name=bussmann>{{cite journal|last1=Bussmann|first1=R.W.|last2=Worede|first2=A.|last3=Swartzinsky|first3=P.|last4=Evangelista|first4=P.|title=Plant use in Odo-Bulu and Demaro, Bale region, Ethiopia|journal=Journal of Ethnobiology and Ethnomedicine|date=2011|volume=7|issue=1|pages=28|doi=10.1186/1746-4269-7-28|pmid=21943288|pmc=3196683 |doi-access=free }}</ref> The animal is extensively hunted for its horns and meat. The meat is utilised in local medicine and for making nipples for traditional milk bottles. Impact of trophy hunting programs is obscure, and current trophy hunting quotas may be unsustainable in the long term although, if well regulated, trophy hunting could play an important role in the long term management of this species.<ref name = iucn/><ref name=east/>
-The mountain nyala has been classified under the Endangered category of the [[International Union for Conservation of Nature and Natural Resources]] (IUCN). It has not been listed under the [[Washington Convention]] (CITES). Though legal protection has been fully ensured for the species, the implementation has not been so effective. In 1991, there was widespread unrest in Ethiopia during which several mountain nyala were killed and the population in the Bale Mountains National Park fell to 150.<ref>{{cite journal|last1=Woldegebriel|first1=G.K.|title=The status of mountain nyala (''Tragelaphus buxtoni'') in Bale Mountains National Park 1986–1994|journal=Walia|date=1996|volume=17|pages=27–37}}</ref><ref name=mamo2>{{cite journal|last1=Mamo|first1=Y.|last2=Pinard|first2=M.A.|last3=Bekele|first3=A.|title=Demography and dynamics of mountain nyala ''Tragelaphus buxtoni '' in the Bale Mountains National Park, Ethiopia|journal=Current Zoology|date=2010|volume=56|issue=6|pages=660–9|doi=10.1093/czoolo/56.6.660|doi-access=free}}</ref><ref name=refera>{{cite journal|last1=Refera|first1=B.|last2=Bekele|first2=A.|title=Population status and structure of mountain nyala in the Bale Mountains National Park, Ethiopia|journal=African Journal of Ecology|date=2004|volume=42|issue=1|pages=1–7|doi=10.1111/j.0141-6707.2004.00437.x}}</ref> After the Bale Mountains National Park, the small [[Kuni-Muktar Mountain Nyala Sanctuary]] was established in 1990 was established as a second protected area. But by 1996 the sanctuary had been so severely affected by deforestation, poaching, cultivation and [[gully erosion]], that the populations soon disappeared from there.<ref name=iucn/><ref name=east/>
+The mountain nyala has been classified under the Endangered category of the [[International Union for Conservation of Nature and Natural Resources]] (IUCN). It has not been listed under the [[Washington Convention]] (CITES). Though legal protection has been fully ensured for the species, the implementation has not been so effective. In 1991, there was widespread unrest in Ethiopia during which several mountain nyala were killed and the population in the Bale Mountains National Park fell to 150.<ref>{{cite journal|last1=Woldegebriel|first1=G.K.|title=The status of mountain nyala (''Tragelaphus buxtoni'') in Bale Mountains National Park 1986–1994|journal=Walia|date=1996|volume=17|pages=27–37}}</ref><ref name=mamo2>{{cite journal|last1=Mamo|first1=Y.|last2=Pinard|first2=M.A.|last3=Bekele|first3=A.|title=Demography and dynamics of mountain nyala ''Tragelaphus buxtoni '' in the Bale Mountains National Park, Ethiopia|journal=Current Zoology|date=2010|volume=56|issue=6|pages=660–9|doi=10.1093/czoolo/56.6.660|doi-access=free}}</ref><ref name=refera>{{cite journal|last1=Refera|first1=B.|last2=Bekele|first2=A.|title=Population status and structure of mountain nyala in the Bale Mountains National Park, Ethiopia|journal=African Journal of Ecology|date=2004|volume=42|issue=1|pages=1–7|doi=10.1111/j.0141-6707.2004.00437.x|bibcode=2004AfJEc..42....1R }}</ref> After the Bale Mountains National Park, the small [[Kuni-Muktar Mountain Nyala Sanctuary]] was established in 1990 was established as a second protected area. But by 1996 the sanctuary had been so severely affected by deforestation, poaching, cultivation and [[gully erosion]], that the populations soon disappeared from there.<ref name=iucn/><ref name=east/>
 In the 1960s, the population was estimated at 7,000 to 8,000 (and perhaps up to 12,500). This declined sharply to 2,000 to 4,000 individuals in the 1980s. The population is decreasing even now. The mountain nyala is feared to be extinct in the eastern and southern extremes of its range, though a few probably survive in [[Chiro (town)|Chiro]] (Asebe Teferi) and the border between Bale and [[Sidamo Province|Sidamo]], south of [[Kofele (woreda)|Kofele]]. Apart from Gaysay about 80 to 120 mountain nyala occur in the rest of the Bale Mountains National Park, and a few hundred occur in the areas in the vicinity of the national park.<ref name=iucn/><ref name=east/>
@@ Line 109: / Line 115: @@
 {{Taxonbar|from=Q255576}}
-{{DEFAULTSORT:nyala, mountain}}
 [[Category:Tragelaphus|mountain nyala]]
 [[Category:Mammals of Ethiopia]]
@@ Line 115: / Line 120: @@
 [[Category:Fauna of the Horn of Africa]]
 [[Category:Bale Mountains]]
-[[Category:Ethiopian Highlands]]
+[[Category:Fauna of the Ethiopian Highlands]]
 [[Category:Endangered biota of Africa]]
 [[Category:Mammals described in 1910|mountain nyala]]