Neotropic Lycaenidae Butterfly Guide 02may2022

“Este listado está dedicado a todas las personas que nos han ayudado a construirlo y a
todos aquellos que están trabajando por el conocimiento y la conservación de la

naturaleza.”
“This list is dedicated to all the people who have helped us to build it and to all those who
are working for the knowledge and conservation of nature.”
Para citar este documento utilice: To cite this document use:
GARWOOD K., JARAMILLO J.G. Guía de Mariposas Lycaenidae del Neotrópico –

Butterfly guide to the Lycaenidae of the Neotropic [en línea], BioButterfly Database.
Disponible en:< http://www.butterflycatalogs.com/ 02 de marzo de 2022 145pp.
Diseño y diagramación Juan Guillermo Jaramillo V.
Laothus gibberosa Pseudolycaena marsyas

Foto cubierta (Front cover): Foto cubierta inglés (English cover):
Juan Guillermo Jaramillo V. Juan Guillermo Jaramillo V.
Introducción Introduction
Con el propósito fundamental de hacer educación With the fundamental purpose of educating and
y crear conciencia sobre la importancia de las raising awareness about the importance of
mariposas en el ecosistema y su necesidad de butterflies in the ecosystem and the need to
conservarlas hemos recopilado información en la conserve them, we have compiled information in
base de datos BioButterfly. the BioButterfly database.
Con esta base de datos elaboramos catálogos y With this database we create butterfly catalogs
guías de mariposas, tanto para todo el Neotrópico and guides, both for the entire Neotropics and for
como para países y localidades desde México countries and localities from Mexico to Argentina.
hasta Argentina.
The guides contain photographs that have been
Las guías contienen fotografías que han compiled and taken by the authors and 364 other
recopilado y tomado los autores y otros 364 photographers who selflessly have given them up
fotógrafos quienes desinteresadamente nos las for our work. That is why at the end of this
han cedido para nuestro trabajo. Por ello al final document we have a complete list of all those
de este documento tenemos un listado completo who have contributed their photos. These
de todos aquellos que han contribuido con sus photographs represent thousands of hours of
fotos. Estas fotografías representan miles de fieldwork that would otherwise have been nearly
horas de trabajo de campo que de otra manera impossible to collect.
hubiera sido casi imposible de recopilar.
This information is constantly evolving and
Esta información está en constante evolución y changing, which is why we believe that the
cambio por eso creemos que las versiones electronic versions are the best option, because
electrónicas son la mejor opción, pues al poder by being able to generate lists and guides without
generar listados y guías sin costo de impresión printing costs whenever you want, it can be the
cada que se quiera puede ser el mecanismo más most practical mechanism for collecting and
práctico para recopilar y compartir la información sharing information about neotropical butterflies.
sobre las mariposas neotropicales.
For the presentation and layout of the document
Para la presentación y diagramación del we chose a format between scientific and didactic
documento escogimos un formato entre científico because, although the information is very
y didáctico pues, aunque la información es muy scientific, the use of photos helps to attract the
científica, el uso de fotos ayuda a atraer público general public and this is part of the objective: to
general y este es parte del objetivo: comunicar communicate to educate and conserve.
para educar y conservar.
We hope that this information will be useful in
Esperamos que esta información sea de utilidad al increasing knowledge about butterflies and that it
aumentar el conocimiento sobre las mariposas y will help promote the observation, study,
que ayude a fomentar la observación, estudio, photography and enjoyment of these, and that
fotografía y disfrute de estas y que la información the information can be used by governments,
pueda ser utilizada por gobiernos, instituciones o institutions or individuals to make decisions and
personas para tomar decisiones y crear conciencia raise awareness about the need for its study and
sobre la necesidad de su estudio y conservación conservation
1
Nota importante Note
Estas guías y listados de chequeo son el These guides and checklists are the result of
resultado de más de veinte años de trabajo de more than twenty years of work by the authors.
los autores. Para llegar al resultado que Usted To reach the result you have in your hands, we
tiene en sus manos hemos contado con la have had the generous help of many expert
generosa ayuda de muchos científicos expertos scientists and many photographers.
y de un gran número de fotógrafos. These publications are the product of the
Estas publicaciones son producto de la compilation of bibliography and photographic
recopilación de bibliografía y registros records from many locations in Neotropical
fotográficos de muchas localidades en países del countries. The user must know that identifying
Neotrópico. El usuario debe saber que some species from only photographs has an
identificar algunas especies a partir solamente implicit margin of error, so it is possible that the
de fotografías tiene implícito un margen de guides have some incorrect identifications or
error, por lo que es posible que las guías tengan lack the precision that we would like for all of
algunas identificaciones incorrectas, o carentes them, despite our efforts and the knowledge
de la precisión que desearíamos para todas, a that the consulted experts have.
pesar de nuestro esfuerzo y del conocimiento To prepare the guides we use photos of
que los expertos consultados tienen. butterflies taken in different countries and from
Para elaborar las guías usamos fotos de these we select the photo or photos that best
mariposas tomadas en diferentes países y de represent the species, therefore, we use that
estas seleccionamos la foto o fotos que mejor photo or photos for all the family guides or for
representen a la especie, por lo tanto, esa foto the different countries. It is for this reason that
o fotos las usamos para todas las guías de familia it is possible, for example, to find a butterfly
o para los diferentes países. Es por esta razón photographed in Peru in a guide to Ecuador. We
que es posible, por ejemplo, encontrar en una are certain that this butterfly has been
guía del Ecuador una mariposa fotografiada en registered in Ecuador, but we use the photo that
el Perú. Nosotros tenemos la certeza de que best represents the species, even if it was taken
dicha mariposa ha sido registrada en Ecuador, in another country, because the idea is that the
pero usamos la foto que mejor represente la user has the best possible idea of what the
especie, así haya sido tomada en otro país, butterfly looks like.
porque la idea es que el usuario tenga la mejor The publication, being generated electronically,
idea posible de cómo luce la mariposa. is easy to correct, does not imply waste of paper
La publicación, al ser generada and can be permanently updated. If you find any
electrónicamente, es fácil de corregir, no error or inconsistency, we have a methodology
implica desperdicio de papel y se puede to correct it: please call Juan Guillermo Jaramillo
actualizar permanentemente. Si encuentra V. at +57-310 830 6732 or write to
algún error o inconsistencia, tenemos una [email protected] or Kim Garwood
metodología para corregirla: por favor llame a [email protected].
Juan Guillermo Jaramillo V. al celular +57-310
830 6732 o escriba al correo
[email protected] o a Kim Garwood
[email protected].
BioButterfly Database – Butterflycatalogs.com
2
Convenciones Keys
Para cada foto de una especie se trata de dar la For each photo of a species, it is about giving as
mayor cantidad de información posible en el much information as possible in the smallest
menor espacio, los números en azul significan: space, the numbers in blue mean:
1- Nombre científico 1- Scientific name

2- Nombre anterior 2- Previous name
3- Sexo 3- Sex
4- Fotógrafo 4- Photographer
5- Información de la foto 5- Photo information
6- Rango geográfico 6- Geographic range
7- Secuencia de la foto 7- Photo sequence
3
1 Nombre científico 1 Scientific name
El nombre científico con género, especie y The scientific name with genus, species and
subespecie cuando sea posible. subspecies when possible.
Los nombres científicos con el género y sp The scientific names with the genus and sp
corresponden a especies que tienen una correspond to species that have an unknown or
identificación desconocida o dudosa, ya que doubtful identification, since there may be
puede haber varias especies con dudas será sp1, several species with doubts, it will be sp1, sp2,
sp2, etc., una vez que se identifique etc., once it is positively identified, the sp will be
positivamente, la sp se cambiará al nombre changed to the correct name of the species.
correcto de la especie. In the case of species not described to science,
En el caso de especies no descritas para la which do not yet have a name, we put sp nov.
ciencia, que todavía no tienen nombre, Please, if you can help us to complete or correct,
colocamos sp nov. Por favor, si puede ayudarnos write to the contacts.
a completar o corregir, escríbanos a los In many places there are species without
contactos. positive identification, we prefer to post photos
En muchos lugares hay especies sin of these species so that it is known that this
identificación positiva, preferimos colocar fotos species is in that place and so that in the future
de estas especies para que se sepa que esta someone can describe it. In case we have doubts
especie está en ese lugar y para que en el futuro even with the gender, we put the gender that
alguien pueda describirla. En caso de que we consider closest and two question marks like
tengamos dudas incluso con el género, this: ??.
ponemos el género que consideramos más In some cases of species found in taxonomic
cercano y dos signos de interrogación como studies, we place the names as follows: sp
este: ??. [group]
En algunos casos de especies que se encuentran The taxonomy and sequence of this catalog
en estudios taxonómicos colocamos mientras se follows the authors that are specified in our
definen los nombres así: sp [grupo] publication: Neotropical Butterflies Checklist
La taxonomía y la secuencia de este catálogo South America.
sigue los autores que están especificados en
2 Former name
nuestra publicación: lista Mariposas del
Neotrópico Lista de Chequeo Suramérica. With the taxonomy recently revised with DNA,
several species changed the genus or name, in
2 Nombre anterior those cases we will put the old genus or name
Con la taxonomía revisada recientemente con el with the red text.
ADN, varias especies cambiaron el género o el
3 Sex
nombre, en esos casos colocaremos el antiguo
género o nombre con el texto en rojo. Whenever possible, we put the sex of the
specimen in the photo, either because they
3 Sexo have sexual dimorphism or because in some
Cuando sea posible, ponemos el sexo del cases, we observed the female laying eggs.
espécimen de la foto, ya sea porque tienen
4
dimorfismo sexual o porque en algunos casos 4 Photographer
observamos a la hembra poniendo huevos.
The abbreviation of the name of the
4 Fotógrafo photographer who took the photo
La abreviación del nombre del fotógrafo que acknowledges their collaboration, please inform
tomó la foto reconoce su colaboración, por us of any errors for correction.
favor nos informan de cualquier error para su 5 Photo information
corrección.
Whenever possible, the photo information will
5 Información de la foto contain the abbreviation of the country, the
Cuando sea posible, la información de la foto locality, the altitude, and the month/day the
tendrá la abreviación del país, la localidad, la photo was taken.
altitud y el mes/día en que se tomó la foto. Most of the photos were taken of live butterflies
La mayoría de las fotos fueron tomadas de in their natural environment. The country where
mariposas vivas en su entorno natural. El país it was taken is indicated by a two-letter
donde fue tomado se indica con una abreviatura abbreviation. This abbreviation corresponds to
de dos letras. Esta abreviatura corresponde al the international code used as the Internet
código internacional utilizado como la termination of the code of each country, for
terminación de Internet del código de cada país, example. Brazil BR, Ecuador EC, Peru PE etc.
por ejemplo. Brasil BR, Ecuador EC, Perú PE etc. After the country code, we put an abbreviation
Después del código del país, ponemos una of the locality (see the table of equivalent codes
abreviatura de la localidad (consulte la tabla de for localities), then the altitude where the photo
códigos equivalentes para las localidades), luego was taken in meters above sea level and the
la altitud donde se tomó la foto en metros sobre month and day, to give an idea of the date they
el nivel del mar y el mes y día, para dar una idea were flying at this location.
de la fecha que estuvo volando en este lugar. From each species or subspecies, we choose the
De cada especie o subespecie escogemos las best photos that represent it both ventral-dorsal
mejores fotos que la representen tanto ventral- and male-female and these photos are used in
dorsal como macho-hembra y estas fotos se the guides for all countries. There are many
usan en las guías para todos los países. Hay species that are common to almost all countries
muchas especies que son comunes a casi todos and that do not differ significantly in their shape
los países y que no difieren significativamente or color between countries, because trying to
en su forma o color entre los países, porque make the guide of a country or locality using
tratar de que la guía de un país o localidad solo only photos taken in that country or locality
tenga fotos tomadas en ese país o localidad se becomes a titanic project knowing that when we
convierte en un proyecto titánico, sabiendo que have more than 14,000 taxa for the Neotropics.
tenemos más de 14.000 taxones para el 6 Range
Neotrópico.
The distribution range of the species with an
6 Rango abbreviation of the name of the countries from
El rango de distribución de las especies con una north to south, in some cases with species that
abreviatura del nombre de los países de norte a inhabit the United States and Mexico, we use
sur, en algunos casos con especies que habitan the code of the southernmost states where the
en los Estados Unidos y México, utilizamos el species are Tx Texas, Chi Chiapas. In big
5
código de los estados más australes donde están countries like Brazil, the upper part of the
las especies Tx Texas, Chi Chiapas. En países Amazon, the lower part, etc. Colombia and
grandes como Brasil, la parte superior del other countries may have a position relative to
Amazonas, la parte inferior, etc. Colombia y the Andes Mountain range and this information
otros países pueden tener una posición relativa is placed in parentheses after the country
a la cordillera de los Andes y esta información se abbreviation.
coloca entre paréntesis después de la
7 Photo Sequence
abreviación del país.
In the upper right corner, there is a consecutive
7 Secuencia de la foto number that helps to quickly locate or refer to
En la esquina superior derecha, hay un número the position of a species in the guide.
consecutivo que ayuda a localizar o hacer
referencia rápidamente a la posición de una
Specimen photos are used to show some
especie en la guía.
features that are not possible to photograph in
the wild or to show species for which we do not
Las fotos de especímenes se usan para mostrar yet have live specimens. In addition, in some
algunas características que no son posibles de cases dorsal photos of captured species are
fotografiar en la naturaleza o para mostrar taken to better identify them, these butterflies
especies de las cuales aún no tenemos are released immediately after the photo.
especímenes vivos. Además, en algunos casos se Photos of specimens or photos of low quality
toman fotos dorsales de especies capturadas will be replaced once better photos of these
para identificarlas mejor, estas mariposas se species are taken, any collaboration that allows
liberan inmediatamente después de la foto. Las to complete the guide will be welcome and the
fotos de especímenes o fotos de baja calidad photographer will have the credits.
serán reemplazadas una vez que se tomen
mejores fotos de estas especies, cualquier
colaboración que permita completar la guía será
bienvenida y el fotógrafo tendrá los créditos.
6
Clasificación familia Lycaenidae
En este listado colocamos todos los géneros en el Neotrópico de la familia agrupados por subfamilia, de tal
manera que se pueda saber la ubicacion de un genero. Todos los generos no necesariamente estan en un pais
o localidad.
We have listed below all the Neotropic genera in the family within each subfamily, to help with locating a
particular genus. All genus are not necessarily in one country or locality.
Lycaenidae
11 Podanotum
12 Rhamma
Subfamilia - Subfamily
13 Salazaria
Lycaeninae Subtribu - Subtribe
Tribu - Tribe
Timaetina
Lycaenini 14 Busbiina
15 Marachina
Subtribu - Subtribe
Sin Subtribu
16 Nesiostrymon
17 Paraspiculatus
1 Iophanus
18 Penaincisalia
19 Phothecla
Subfamilia - Subfamily 20 Timaeta
Theclinae Subtribu - Subtribe
Atlidina
Tribu - Tribe
21 Arcas
Eumaeini
22 Atlides
Subtribu - Subtribe
23 Brangas
Eumaeina
24 Denivia
2 Brevianta
25 Pseudolycaena
3 Eumaeus
26 Theritas
4 Micandra Subtribu - Subtribe
5 Mithras Evenina
6 Theorema 27 Evenus
7 Thestius Subtribu - Subtribe
Subtribu - Subtribe Jantheclina

Rhammina 28 Allosmaitia
8 Balintus 29 Aveexcrenota
9 Johnsonita 30 Contrafacia
10 Lathecla
BioButterfly Database - Butterflycatalogs
7
Lycaenidae
31 Enos 56 Camissecla
32 Janthecla 57 Electrostrymon
33 Laothus 58 Gigantorubra
Subtribu - Subtribe 59 Kisutam
Paiwarriina 60 Lamprospilus
34 Annamaria 61 Mercedes
35 Fasslantonius 62 Pendantus
36 Kolana 63 Rubroserrata
37 Paiwarria 64 Serratofalca
Subtribu - Subtribe
65 Ziegleria
Cupatheclina
Subtribu - Subtribe
38 Cupathecla Strymonina
39 Dabreras 66 Arawacus
Subtribu - Subtribe
67 Heterosmaitia
Parrhasiina
68 Hypostrymon
40 Apuecla
69 Panthiades
41 Beatheclus
70 Rekoa
42 Caerofethra
71 Strymon
43 Dicya
72 Thereus
44 Ignata
Subtribu - Subtribe
45 Michaelus Strephonotina
46 Oenomaus 73 Aubergina
47 Olynthus 74 Celmia
48 Parrhasius 75 Crimsinota
49 Symbiopsis 76 Decussata
50 Thepytus 77 Gargina
Subtribu - Subtribe
78 Gossenia
Ipideclina
79 Iaspis
51 Ipidecla
Subtribu - Subtribe
80 Ministrymon
Calycopidina 81 Nicolaea
52 Argentostriatus 82 Ostrinotes
53 Arzecla 83 Rindgea
54 Badecla 84 Siderus
55 Calycopis 85 Strephonota
8
Lycaenidae
86 Terenthina 108 Echinargus
87 Theclopsis 109 Eldoradina
88 Tmolus 110 Elkalyce
Subtribu - Subtribe 111 Hemiargus
Trichonidina 112 Itylos
89 Bistonina 113 Leptotes
90 Manticia 114 Nabokovia
91 Megathecla 115 Paralycaeides
92 Trichonis 116 Pseudochrysops
Subtribu - Subtribe
117 Pseudolucia
Callophryidina
118 Zizula
93 Callophrys
94 Cyanophrys
95 Chalybs
96 Chlorostrymon
97 Erora
98 Magnastigma
99 Ocaria
100 Radissima
101 Satyrium
102 Semonina
103 Thaeides
Subfamilia - Subfamily
Polyommatinae
Tribu - Tribe
Polyommatini
Subtribu - Subtribe
Sin Subtribu
104 Brephidium
105 Celastrina
106 Cupido
107 Cyclargus
9
10
Lycaenidae del Neotrópico
Subfamilia Lycaeninae
Tribu - Tribe
Lycaenini
1 2
Subtribu - Subtribe
Sin Subtribu
Iophanus pyrrhias ♂ Iophanus pyrrhias ♂

MR MR
MX-, Jul14 MX(Chi),GT,HN MX-, Jul14 MX(Chi),GT,HN
3 4 5
Iophanus pyrrhias ♂ Iophanus pyrrhias ♂ Iophanus pyrrhias ♂

JFG JMz MR
GT-, Oct14 MX(Chi),GT,HN GT-, Jul29 MX(Chi),GT,HN MX-, Jul14 MX(Chi),GT,HN
6 7 8
Iophanus pyrrhias ♂ Iophanus pyrrhias ♂ Iophanus pyrrhias ♀

JMz LMz JMz
GT-, Nov11 MX(Chi),GT,HN GT-, Jul29 MX(Chi),GT,HN GT-, May05 MX(Chi),GT,HN
BioButterfly DataBase - Butterflycatalogs.com Lycaeninae

11
Tribu - Tribe
Lycaenini
9
Iophanus pyrrhias
RGd
HN-, Mar23 MX(Chi),GT,HN
AA infNEOTROPICO SUBFAMILIA Photo Guide
Esta versión del catálogo se generó el lunes, 2 de mayo de 2022 Date of report
Busque versión nueva de este catálogo en: Butterflycatalogs.com

Look for a newer version of this catalog in: Neotropicalbutterflies.com
BioButterfly DataBase - Butterflycatalogs.com Lycaeninae Página 2 de 2

12
13
Subfamilia Theclinae
Tribu - Tribe
Eumaeini
1 2
Subtribu - Subtribe
Eumaeina
Brevianta bathoryon Brevianta busa

DvG RGd
EC-Cal, W CO-EC HO-, MX-EC
3 4 5
Brevianta celelata Brevianta ematheon Brevianta ematheon

DvG KK DvG
AR-ChM, Dec28 BR-AR EC-Atrk, VZ-PE,BR,SU PE-ARCC, VZ-PE,BR,SU
6 7 8
Brevianta hyas Brevianta undulata Eumaeus childrenae

DW DvG KP
PN-Bqt, Apr24 MX-CR PE-Qtz, VZ-PE MX-Oax, May16 MX-HN
BioButterfly DataBase - Butterflycatalogs.com Theclinae

14
Tribu - Tribe
Eumaeini
9 10 11
Eumaeus godartii Eumaeus godartii Eumaeus toxana

JMD WC AN
PA-, NI-EC PA-, NI-EC EC-Atrk, VZ-BO,BR
12 13 14
Eumaeus toxana Eumaeus toxea Micandra aegides ♂

DW BBt KG
PE-, VZ-BO,BR MX-Plq, May13 US(Tx)-NI EC-Cal, VZ-PE
15 16 17
Micandra aegides ♀ Micandra comae ♂ Micandra comae ♀

DvG DvG DvG
EC-Bañ, VZ-PE EC-Cal, CO-PE EC-Orq, CO-PE
18 19 20
Micandra comae ♀ Micandra comae ♀ Micandra comae ♀

JAB DvG DvG
CO-Oqy, 1850m CO-PE EC-Cal, CO-PE EC-Caj, CO-PE
BioButterfly DataBase - Butterflycatalogs.com Theclinae Página 2 de 114

15
Tribu - Tribe
Eumaeini
21 22 23
Micandra comae ♀ Micandra comae Micandra dignota

KK DvG DvG
CO-RlT, CO-PE CO-Mzm, 2600m CO-PE PE-Wyq, CO-BO
24 25 26
Micandra ion Micandra platyptera ♂ Micandra platyptera ♂

DvG KG DvG
PE-Rctl, CO-PE CO-SM, CR-BO,GF PE-PNYC, CR-BO,GF
27 28 29
Micandra platyptera ♀ Micandra platyptera ♀ Micandra platyptera ♀

BBt AN
EC-, CR-BO,GF EC-WS, CR-BO,GF CO-QEV, 1975m CR-BO,GF
30 31 32
Micandra platyptera ♀ Micandra platyptera ♀ Micandra platyptera ♀
CO-QEV, 1975m CR-BO,GF CO-PSJ, 1850m Oct17 CR-BO,GF CO-PSJ, 1800m Nov21 CR-BO,GF

16
Tribu - Tribe
Eumaeini
33 34 35
Micandra platyptera ♀ Micandra sp1 Mithras nautes ♂

DW DvG
CO-QEV, 1975m CR-BO,GF CO-, - PE-ARCC, VZ-PE,BR
36 37 38
Mithras nautes ♀ Mithras nautes Theorema dysmenia

DvG RH FM
PE-ARCC, VZ-PE,BR BR-CJL, VZ-PE,BR CO-, CO-PE
39 40 41
Thestius epopea Thestius epopea Thestius epopea

DvG DvG DvG
CO-Mzm, 1350m NI-PE CO-Mzm, 1350m NI-PE CO-Mzm, 1350m NI-PE
42 43 44
Thestius meridionalis ♂ Thestius meridionalis ♂ Thestius meridionalis ♂

KNc DvG DvG
PE-PNM, PN-BO,BR EC-, 550m PN-BO,BR CO-SM, 750m PN-BO,BR

17
Tribu - Tribe
Eumaeini
45 46 47
Thestius meridionalis ♂ Thestius meridionalis ♂ Thestius meridionalis ♀

FM DvG DvG
CO-RP, 560m Ago23 PN-BO,BR CO-MMt, 200m PN-BO,BR EC-AJL, 400m PN-BO,BR
48
Thestius meridionalis ♀
SS
CO-MMt, 200m Nov22 PN-BO,BR
49 50
Subtribu - Subtribe
Rhammina
Balintus tityrus Balintus tityrus

DvG DvG
EC-WS, CO-BO EC-WS, CO-BO
51 52 53
Balintus tityrus Johnsonita auda ♂ Johnsonita auda
CO-RlR, 1850m - CO-BO CO-, - PN-PE,VZ CO-, PN-PE,VZ

18
Tribu - Tribe
Eumaeini
54 55 56
Johnsonita auda Johnsonita auda Johnsonita auda

DvG KG
PE-Rctl, PN-PE,VZ CO-SFx, 2600m May18 PN-PE,VZ PE-Mrd, PN-PE,VZ
57 58 59
Johnsonita auda Johnsonita auda Johnsonita chlamydem ♂

SN DvG
CO-Oqy, 1850m Sep26 PN-PE,VZ CO-QEV, 1940m PN-PE,VZ PE-Salb, May31 PE-BO
60 61 62
Johnsonita chlamydem ♂ Johnsonita chlamydem ♀ Johnsonita chlamydem ♀

DvG DvG DvG
PE-Salb, May31 PE-BO PE-Pro, Jun03 PE-BO PE-PMp, Sep28 PE-BO
63 64 65
Johnsonita chlamydem Johnsonita chlamydem Johnsonita pardoa

THn KK THn
PE-, PE-BO PE-Pro, Jun05 PE-BO CO-Jard, VZ-BO

19
Tribu - Tribe
Eumaeini
66 67 68
Johnsonita pardoa Johnsonita turquisca Lathecla latagus

AN THn DvG
PE-, VZ-BO EC-, Nov24 CO-EC CO-Mzm, MX-BO,GF,BR?
69 70 71
Lathecla latagus Lathecla mimula Lathecla mimula

DvG DvG AN
PE-Mrd, MX-BO,GF,BR? CO-Mzm, 1300m CO?EC-PE,BR PE-, CO?EC-PE,BR
72 73 74
Lathecla mimula Lathecla mimula Rhamma adunca

DvG DvG AN
PE-MWC, CO?EC-PE,BR CO-Mzm, 1300m CO?EC-PE,BR EC-, CO-EC
75 76 77
Rhamma arria ♂ Rhamma arria Rhamma aurugo

FM AH DvG
CO-RPF, 3000m VZ-BO,AR CO-Mzm, VZ-BO,AR PE-Cus, PE

20
Tribu - Tribe
Eumaeini
78 79 80
Rhamma bilix Rhamma bilix Rhamma commodus

Leo Leo
CO-Mzm, CO-EC CO-Mzm, CO-EC CO-PSJ, 1800m Dic19 VZ-BO
81 82 83
Rhamma commodus Rhamma commodus Rhamma comstocki ♀

FM
CO-PSJ, 1800m Dic19 VZ-BO CO-PSJ, 1800m Dic19 VZ-BO CO-RPF, 3000m CO-PE
84 85 86
Rhamma familiaris Rhamma familiaris Rhamma hybla

Paralustrus DvG Paralustrus KG Shapiroana DvG
EC-Caj, CO-BO EC-Sidr, CO-BO EC-Caj, CO-PE
87 88 89
Rhamma mishma Rhamma oxida Rhamma sp1

JSn DvG
CO-Jard, CO(SNSM) CO-, 2600m Feb03 CO-EC CO-SFx, 2800m May16 -

21
Tribu - Tribe
Eumaeini
90 91 92
Rhamma sp2 Rhamma sp2 Salazaria sala

DvG DvG DvG
PE-, PE-, CO-Mzm, May28 CO-PE
93 94 95
Salazaria sala Salazaria sala Salazaria sp1

PB THn GM
CO-Oqy, 1850m Jun04 CO-PE EC-, Nov12 CO-PE CO-Mzm, Ago30 -
96 97
Subtribu - Subtribe
Timaetina
Busbiina bosora Marachina maraches

AN DvG
EC-Orq, CO-EC,BR,GF PE-Rctl, CO-BO
98 99 100
Marachina maraches Marachina sp1 Nesiostrymon calchinia

DvG DvG KG
CO-Mzm, 1700m CO-BO CO-Mzm, 1350m Oct04 - CO-RlT, 1700m Sep22 MX-AR

22
Tribu - Tribe
Eumaeini
101 102 103
Nesiostrymon calchinia Nesiostrymon celona Nesiostrymon celona

DvG DvG DvG
CO-Mzm, 1700m MX-AR PE-PNYC, MX-BR PE-Rmp-Mq, MX-BR
104 105 106
Nesiostrymon celona Nesiostrymon hyccara Nesiostrymon hyccara

OE DvG DvG
CO-, MX-BR BR-, BR BR-, BR
107 108 109
Nesiostrymon tristis Paraspiculatus azul Paraspiculatus colombiensis

PB THn Mithras DvG
CO-, 2600m HN-CO,BR(MT) EC-, E EC-PE CO-Mzm, 1800m CO-BO,BR,GF
110 111 112
Paraspiculatus elis Paraspiculatus elis Paraspiculatus oroanna

Mithras AN Mithras DvG DvG
EC-Ten, SE CO,E EC-PE,BR,SU EC-PmpL, SE CO,E EC-PE,BR,SU PE-VCm, E EC-PE

23
Tribu - Tribe
Eumaeini
113 114 115
Paraspiculatus orobia Paraspiculatus orobia Paraspiculatus orobia

Mithras DvG Mithras DvG Mithras DvG
PE-Plm, CO-BO,BR,AR PE-MWC, CO-BO,BR,AR EC-Orq, CO-BO,BR,AR
116 117 118
Paraspiculatus orobia Paraspiculatus orobia Paraspiculatus orocana

Mithras TH Mithras AN Mithras DJF
EC-Atrk, CO-BO,BR,AR PE-, CO-BO,BR,AR CO-, 1300m Jul16 CO-PE
119 120 121
Paraspiculatus orocana Penaincisalia amatista Penaincisalia atymna

Mithras DJF Abloxurina DvG DvG
CO-, 1300m Jul16 CO-PE PE-Wyq, EC-PE EC-Loj, EC-BO
122 123 124
Penaincisalia culminicola Penaincisalia culminicola Penaincisalia ianusi

KG DvG Ianusanta Mmi
EC-Pulu, PE-BO PE-Cus, PE-BO EC-Baz, EC-PE

24
Tribu - Tribe
Eumaeini
125 126 127
Penaincisalia loxurina ♂ Penaincisalia loxurina Penaincisalia loxurina

Thecloxurina Thecloxurina Thecloxurina
CO-SSb, 2500m CO-BO,VZ,AR CO-SFx, 2800m May16 CO-BO,VZ,AR CO-SFx, 2800m May16 CO-BO,VZ,AR
128 129 130
Penaincisalia loxurina Penaincisalia loxurina Penaincisalia loxurina

Thecloxurina Thecloxurina DvG Thecloxurina
CO-SFx, 2800m May16 CO-BO,VZ,AR PE-Wyq, CO-BO,VZ,AR CO-SFx, 2800m May16 CO-BO,VZ,AR
131 132 133
Penaincisalia loxurina Penaincisalia loxurina Penaincisalia loxurina

Thecloxurina Thecloxurina DvG Thecloxurina BBt
CO-SFx, 2600m May18 CO-BO,VZ,AR CO-Mzm, 2600m CO-BO,VZ,AR CO-Jard, CO-BO,VZ,AR
134 135 136
Penaincisalia penai Penaincisalia purpurea Penaincisalia purpurea

AN Pons KK Pons DvG
EC-PaP, CO-EC PE-Wyq, EC-PE PE-Wyq, EC-PE

25
Tribu - Tribe
Eumaeini
137 138 139
Penaincisalia santamarta Penaincisalia swarthea Phothecla photismos

Thecloxurina DvG Thecloxurina DvG DvG
CO-SLz, 2600m May18 CO EC-Loj, CO-EC PE-Mrd, VZ-PE
140 141 142
Timaeta gabriela ♀ Timaeta sp n Timaeta tema

DvG Temecla DvG Temecla KK
EC-WS, EC-BO PE-Yab, Oct26 PE BO-Vtu, CO-BO,BR
143 144 145
Timaeta tema Timaeta tema Timaeta timaeus ♂

Temecla DvG Temecla DvG
PE-PntL, CO-BO,BR PE-ARCC, CO-BO,BR CO-, - CO-BO
146 147 148
Timaeta timaeus ♂ Timaeta timaeus ♂ Timaeta timaeus ♂

DvG
CO-, - CO-BO PE-Rctl, CO-BO CO-PSJ, 1800m CO-BO

26
Tribu - Tribe
Eumaeini
149 150 151
Timaeta timaeus ♂ Timaeta timaeus ♀ Timaeta timaeus ♀
CO-, CO-BO CO-RlR, 1850m - CO-BO CO-, - CO-BO
152 153 154
Timaeta timaeus ♀ Timaeta trochus ♀ Timaeta werneri ♀

DvG KK DvG
PE-CoR, CO-BO PE-CoR, CO-PE EC-, 1400m CO-EC
155
Timaeta werneri ♀
DvG
EC-, 1400m CO-EC
156 157
Subtribu - Subtribe
Atlidina
Arcas cypria Arcas ducalis

RSb LBd
MX-Ctm, Aug20 MX-VZ AR-ChM, Mar09 S BR-AR,UR

27
Tribu - Tribe
Eumaeini
158 159 160
Arcas gozmanyi Arcas gozmanyi Arcas imperialis ♂

KP DvG
CO-PSJ, 1800m Ago06 CR-EC PA-, CR-EC PE-ARCC, MX-AR
161 162 163
Arcas imperialis ♂ Arcas imperialis ♂ Arcas imperialis ♀

DvG DvG
PE-ARCC, MX-AR PE-Qtz, MX-AR CO-, - MX-AR
164 165 166
Arcas splendor Arcas tuneta Arcas tuneta tuneta

Lep THn
CO-Mzm, CR-EC EC-, CO-BO,BR,Guianas CO-Rmo, 1830m Feb26 CO-BO,BR
167 168 169
Atlides atys ♂ Atlides atys ♀ Atlides carpasia

DvG FM BBt
PE-ARCC, PN-BO,BR CO-, PN-BO,BR MX-Teo, May29 MX-PN CO?

28
Tribu - Tribe
Eumaeini
170 171 172
Atlides dahnersi Atlides dahnersi Atlides gaumeri

DvG DvG BBt
CO-Mzm, 2450m Oct19 W CO CO-Mzm, 2450m Oct19 W CO MX-Teo, May29 MX-PN CO?
173 174 175
Atlides gaumeri Atlides gaumeri Atlides halesus ♂

BBt KD KG
MX-Teo, May29 MX-PN CO? MX-HTn, Nov23 MX-PN CO? TX-RGv, Oct13 US-MX(Ox)
176 177 178
Atlides halesus Atlides havila Atlides inachus

RL GS AN
TX-RGv, Dec20 US-MX(Ox) BO-Cor, CO-BO PE-, Abr19 MX-BR
179 180 181
Atlides inachus Atlides inachus Atlides polybe ♂

Leo AN DW
CO-Mzm, MX-BR PE-PNM, MX-BR PN-Bqt, May03 MX-AR

29
Tribu - Tribe
Eumaeini
182 183 184
Atlides polybe ♂ Atlides polybe ♂ Atlides rustan ♂

KD DvG FM
MX-HTn, Nov23 MX-AR CO-Mzm, Sep26 MX-AR CO-, May31 MX-BO,BR,AR
185 186 187
Atlides rustan ♀ Brangas carthaea Brangas coccineifrons

DvG DW DvG
PE-Jgu, MX-BO,BR,AR PN-Bqt, MX-EC PE-ARCC, MX-PE
188 189 190
Brangas felderi Brangas getus Brangas getus

CRm DvG DvG
CO-, 1750m Ago01 CO-PE PE-ARCC, MX-BO,BR,AR PE-ARCC, MX-BO,BR,AR
191 192 193
Brangas getus Brangas insolitus Brangas neora ♀

DvG DvG
CO-SM, 750m MX-BO,BR,AR PE-Rctl, EC-PE CO-GMq, 5m MX-BR

30
Tribu - Tribe
Eumaeini
194 195 196
Brangas neora ♀ Brangas polonus Brangas rambutorum

GS JOG
CO-GMq, 5m MX-BR BO-Crv, PE-BO CO-Glp, CO-VZ
197 198 199
Denivia arene Denivia augustinula Denivia danaus

DvG Theritas SS Theritas/Lucilda DvG
PE-Mrd, EC-PE HN-, MX-N CO CO-Mzm, 1400m VZ-PE
200 201 202
Denivia danaus Denivia hemon ♂ Denivia hemon ♂

Theritas/Lucilda DvG Theritas DvG Theritas BBt
CO-Mzm, 1400m VZ-PE PE-ARCC, MX-BO,BR,AR CO-CRC, 320m Jun20 MX-BO,BR,AR
203 204 205
Denivia hemon ♀ Denivia hemon ♀ Denivia hemon ♀

Theritas GC Theritas BBt Theritas
PA-, MX-BO,BR,AR CO-CRC, 320m Jun21 MX-BO,BR,AR CO-HNp, 200m Ene29 MX-BO,BR,AR

31
Tribu - Tribe
Eumaeini
206 207 208
Denivia lisus ♂ Denivia lisus ♂ Denivia lisus ♀

Theritas DvG Theritas KG Theritas DvG
PE-ARCC, MX-BO,BR,AR PE-CoR, MX-BO,BR,AR PE-Qtz, MX-BO,BR,AR
209 210 211
Denivia lisus ♀ Denivia monica Denivia monica

Theritas DvG DvG KG
PE-Qtz, MX-BO,BR,AR PE-Qtz, VZ-BO PE-Qtz, VZ-BO
212 213 214
Denivia monica Denivia phegeus ♂ Denivia phegeus ♀

DvG Theritas AN Theritas DvG
EC-Qtz, VZ-BO PE-, CO-BO,VZ,BR PE-ARCC, CO-BO,VZ,BR
215 216 217
Denivia phegeus ♀ Denivia phegeus Denivia phegeus

Theritas AN Theritas DvG Theritas DvG
PE-, CO-BO,VZ,BR CO-Bvt, 200m Ene31 CO-BO,VZ,BR PE-Qtz, CO-BO,VZ,BR

32
Tribu - Tribe
Eumaeini
218 219 220
Denivia theocritus ♂ Denivia theocritus ♂ Denivia theocritus ♂

Theritas WC Theritas Theritas
MX-Mue, MX-EC,VZ CO-, - MX-EC,VZ CO-, MX-EC,VZ
221 222 223
Denivia viresco ♂ Denivia viresco ♂ Denivia viresco ♀

Theritas DvG Theritas DvG Theritas DvG
PE-ARCC, PE-BO,BR(Ma),GF PE-ARCC, PE-BO,BR(Ma),GF PE-ARCC, PE-BO,BR(Ma),GF
224 225 226
Pseudolycaena damo ♂ Pseudolycaena damo ♂ Pseudolycaena damo ♀

KW KP
EC-, MX-EC,VZ PN-Vant, MX-EC,VZ CO-, - MX-EC,VZ
227 228 229
Pseudolycaena marsyas ♂ Pseudolycaena marsyas ♂ Pseudolycaena marsyas ♂

SS
EC-Ten, NI?-AR CO-Tft, 574m NI?-AR CO-, - NI?-AR

33
Tribu - Tribe
Eumaeini
230 231 232
Pseudolycaena marsyas ♂ Pseudolycaena marsyas ♀ Pseudolycaena marsyas ♀

KK FM
CO-MMt, 200m Oct20 NI?-AR CO-RP, 560m Nov07 NI?-AR CO-SNc, 474m NI?-AR
233 234 235
Pseudolycaena marsyas Pseudolycaena marsyas Pseudolycaena marsyas

KK KK
PE-Jgu, NI?-AR PE-Jgu, NI?-AR CO-Sin, 780m Jun14 NI?-AR
236 237 238
Pseudolycaena marsyas Pseudolycaena marsyas Theritas drucei

DvG MCC
BO-Volc, NI?-AR CO-Sin, 700m Jul05 NI?-AR BR-, BR(RJ,SC)
239 240 241
Theritas harrietta Theritas harrietta Theritas mavors ♂

DvG DvG FM
PE-Rctl, PE-BO PE-Qtz, PE-BO CO-RP, 560m Jul25 MX-BO,BR,GY

34
Tribu - Tribe
Eumaeini
242 243 244
Theritas mavors ♂ Theritas mavors ♂ Theritas mavors ♀

FM FM
CO-, May31 MX-BO,BR,GY CO-RP, 560m Feb09 MX-BO,BR,GY CO-, MX-BO,BR,GY
245 246 247
Theritas paupera Theritas paupera Theritas paupera

DvG
CO-Mzm, 1800m CO-PE,VZ CO-SJn, 800m Sep12 CO-PE,VZ CO-PSJ, 1800m Abr26 CO-PE,VZ
248 249
Subtribu - Subtribe
Evenina
Evenus batesii Evenus batesii

FM DvG
CO-RP, 560m Ago23 MX-BR,AR EC-PmpL, MX-BR,AR
250 251 252
Evenus coronata Evenus felix Evenus floralia

CRm AN AN
CO-, 2300m Oct02 MX-PE EC-Ten, VZ-PE EC-Atrk, EC-BO,BR,GF

35
Tribu - Tribe
Eumaeini
253 254 255
Evenus gabriela Evenus gabriela Evenus latreillii ♂

DvG DvG DvG
EC-Atrk, CO-PE,BR,GF EC-Atrk, CO-PE,BR,GF AR-ChM, Jan20 BR-AR,PY
256 257 258
Evenus latreillii ♂ Evenus latreillii ♀ Evenus regalis

DvG DvG MCC
AR-ChM, Jan20 BR-AR,PY AR-ChM, Jan20 BR-AR,PY BR-, Abr02 MX-BR,AR
259 260 261
Evenus regalis Evenus regalis Evenus regalis

KK JJ JJ
EC-Atrk, MX-BR,AR CO-, May06 MX-BR,AR CO-, May06 MX-BR,AR
262 263 264
Evenus satyroides ♂ Evenus satyroides ♂ Evenus satyroides ♂

GC KK DvG
BR-CJL, CO-BO,VZ,BR EC-Atrk, CO-BO,VZ,BR PE-, CO-BO,VZ,BR

36
Tribu - Tribe
Eumaeini
265 266 267
Evenus satyroides ♀ Evenus sumptuosa Evenus sumptuosa

DvG AN KK
PE-, CO-BO,VZ,BR EC-Atrk, EC-S BR EC-Atrk, EC-S BR
268 269 270
Evenus sumptuosa Evenus tagyra Evenus tagyra

THn WC KK
EC-, EC-S BR BR-CJL, PE-C BR PE-ARCC, PE-C BR
271 272
Subtribu - Subtribe
Jantheclina
Allosmaitia myrtusa Allosmaitia myrtusa

DvG DvG
PE-Jgu, PN-BO EC-Orq, PN-BO
273 274 275
Allosmaitia strophius Allosmaitia strophius Allosmaitia strophius

DvG AN
PE-Qtz, US(Tx)-BO,BR,AR PE-VCm, US(Tx)-BO,BR,AR CO-Mco, US(Tx)-BO,BR,AR

37
Tribu - Tribe
Eumaeini
276 277 278
Allosmaitia strophius Allosmaitia strophius Allosmaitia strophius

DvG DvG
CO-Mco, US(Tx)-BO,BR,AR EC-Atrk, US(Tx)-BO,BR,AR PE-ARCC, US(Tx)-BO,BR,AR
279 280 281
Allosmaitia strophius Allosmaitia strophius Aveexcrenota anna ♂

DvG DvG Theritas DPt
PE-ARCC, US(Tx)-BO,BR,AR EC-WS, US(Tx)-BO,BR,AR CO-PYm, 2100m Nov04 CO-PE
282 283 284
Contrafacia ahola Contrafacia ahola Contrafacia bassania

WC KG MS
MX-HCcd, Nov23 MX-VZ MX-Oax, Jul08 MX-VZ MX-Ord, Nov20 MX-CO
285 286 287
Contrafacia bassania Contrafacia imma Contrafacia imma

KG
MX-Ord, Nov20 MX-CO CO-VCC, 1650m Mar18 MX-BO,BR,AR CO-VCC, 1650m Mar18 MX-BO,BR,AR

38
Tribu - Tribe
Eumaeini
288 289 290
Contrafacia imma Contrafacia imma Contrafacia imma
CO-VCC, 1650m Mar18 MX-BO,BR,AR CO-VPN, 1450m MX-BO,BR,AR CO-VPN, 1450m MX-BO,BR,AR
291 292 293
Contrafacia marmoris Enos falerina Enos falerina

DvG JM RnM
EC-Mnd, Dec02 CO-EC GT-, MX-BO,BR VZ-, MX-BO,BR
294 295 296
Enos maculata Enos myrtea Enos polka

WC FM DvG
BR-CJL, CO?EC-BR CO-RNM, 150m Jun21 CO-PE,BR PE-Qtz, EC-BO,BR
297 298 299
Enos thara Janthecla cydonia Janthecla cydonia

BBn
PN-Vant, Jul07 MX-AR CO-FCy, - CR-CO CO-FCy, - CR-CO

39
Tribu - Tribe
Eumaeini
300 301 302
Janthecla janthina ♂ Janthecla leea Janthecla leea

DvG AN
CO-RSm, 400m Mar05 MX-VZ BR-RG, CO-BO,BR,GF EC-Atrk, CO-BO,BR,GF
303 304 305
Janthecla malvina Janthecla rocena Janthecla rocena

DW KK DvG
PE-Jgu, PE-BO,BR(RJ) PE-ARCC, MX-BO,BR,AR PE-ARCC, MX-BO,BR,AR
306 307 308
Janthecla rocena Janthecla sista Janthecla sista

DvG DvG DvG
PE-ARCC, MX-BO,BR,AR PE-Qtz, CO-BO,BR,VZ PE-Qtz, CO-BO,BR,VZ
309 310 311
Janthecla sista Janthecla sp1 Janthecla sp2

AN DvG DvG
PE-, CO-BO,BR,VZ CO-MMt, 200m - CO-SM, 800m -

40
Tribu - Tribe
Eumaeini
312 313 314
Laothus gibberosa ♂ Laothus gibberosa ♂ Laothus gibberosa ♂

Gibbosa FM Gibbosa Gibbosa DvG
CO-RP, 560m CO-BO CO-, - CO-BO EC-Rho, CO-BO
315 316 317
Laothus gibberosa ♀ Laothus gibberosa ♀ Laothus gibberosa ♀

Gibbosa KG Gibbosa DvG Gibbosa
CO-PSJ, 1800m CO-BO PE-Qtz, CO-BO CO-PSJ, 1800m CO-BO
318 319 320
Laothus gibberosa ♀ Laothus gibberosa Laothus oceia

Gibbosa DvG Gibbosa DvG DvG
EC-Orq, CO-BO PE-Qtz, CO-BO CO-Mzm, 1700m MX-CO
321 322 323
Laothus oceia Laothus phydela Laothus viridicans

DW MZ DvG
CO-, MX-CO BR-, BR(RJ)-AR EC-Cal, VZ-BO

41
Tribu - Tribe
Eumaeini
324 325 326
Laothus viridicans Laothus viridicans Laothus viridicans

AN DvG
EC-Loj, VZ-BO PE-Qtz, VZ-BO CO-, - VZ-BO
327 328
Subtribu - Subtribe
Paiwarriina
Annamaria columbia ♂ Annamaria columbia ♂

Airamanna Leo Airamanna DvG
CO-Mzm, CO CO-Mzm, Sep23 CO
329 330 331
Annamaria columbia ♀ Annamaria columbia ♀ Annamaria columbia ♀

Airamanna DvG Airamanna DvG Airamanna DvG
CO-Mzm, 1700m Jun28 CO CO-Mzm, 1700m Jun28 CO CO-Mzm, 1150m Ene07 CO
332 333 334
Annamaria draudti Annamaria draudti Annamaria ganimedes

Lamasina JK Lamasina Awr Lamasina DvG
MX-Oax, Aug30 GT-EC EC-Mnd, Nov03 GT-EC PE-PMp, Sep28 Guianas-PE

42
Tribu - Tribe
Eumaeini
335 336 337
Annamaria ganimedes Annamaria rhaptissima ♂ Annamaria rhaptissima ♀

Lamasina DvG Airamanna DvG Airamanna DvG
PE-Qtz, Nov15 Guianas-PE PE-Mrd, CO-BO PE-Qtz, CO-BO
338 339 340
Annamaria rhaptissima ♀ Fasslantonius episcopalis ♂ Fasslantonius episcopalis ♂

Airamanna AN Paiwarria Leo Paiwarria Leo
PE-PNM, CO-BO CO-Mzm, CO-PE CO-Mzm, CO-PE
341 342 343
Fasslantonius episcopalis ♀ Kolana ergina Kolana ergina

Paiwarria TGz AN KK
CO-, Oct02 CO-PE PE-Qtz, CO-PE,BR(Am),GF BO-Vtu, CO-PE,BR(Am),GF
344 345 346
Kolana ergina Kolana ergina Kolana ligurina

FM DvG KK
CO-RP, 560m Nov07 CO-PE,BR(Am),GF PE-Qtz, CO-PE,BR(Am),GF EC-Atrk, MX-BO,BR,AR

43
Tribu - Tribe
Eumaeini
347 348 349
Kolana ligurina Kolana lyde Paiwarria telemus ♂

AN KK GC
PE-PNM, MX-BO,BR,AR EC-Atrk, MX-PE,VZ BR-CJL, CO-BO,BR
350 351 352
Paiwarria telemus ♀ Paiwarria telemus ♀ Paiwarria telemus ♀

DvG WC DvG
EC-Atrk, CO-BO,BR BR-CJL, CO-BO,BR PE-Qtz, CO-BO,BR
353 354 355
Paiwarria venulius Paiwarria venulius Paiwarria venulius

MCC AL DvG
BR-, Nov19 CO-BO,BR,AR EC-, CO-BO,BR,AR PE-ARCC, CO-BO,BR,AR
356
Paiwarria venulius
TC
EC-AJL, CO-BO,BR,AR

44
Tribu - Tribe
Eumaeini
357 358
Subtribu - Subtribe
Cupatheclina
Cupathecla cupentus Cupathecla cupentus

Megathecla THn Megathecla DvG
EC-, MX-BO,GY,BR PE-ARCC, Sep28 MX-BO,GY,BR
359
Cupathecla cupentus
Megathecla DvG
CO-Mzm, 1400m Jun23 MX-BO,GY,BR
360 361
Subtribu - Subtribe
Parrhasiina
Apuecla n sp Apuecla picus

DvG DvG
BO-VTn, Nov27 BO PE-MWC, Oct20 PE-BR,GY,GF
362 363 364
Apuecla upupa Dicya carnica Dicya carnica

KK KG DvG
PE-, 900m May26 MX-VZ,EC CO-Rpau, MX-AR AR-ChM, MX-AR

45
Tribu - Tribe
Eumaeini
365 366 367
Dicya carnica Dicya dicaea Dicya dicaea

DvG DvG DvG
EC-Cpl, MX-AR AR-ChM, MX-AR AR-ChM, Dec23 MX-AR
368 369 370
Dicya iambe Dicya iambe Dicya iambe

FM
CO-, CR-BO,GF CO-PSJ, 1800m CR-BO,GF CO-PSJ, 1800m CR-BO,GF
371 372 373
Dicya iambe Ignata caldas Ignata caldas

DvG AN AN
PE-Qtz, CR-BO,GF PE-, MX-PE PE-, MX-PE
374 375 376
Ignata caldas Ignata levis Ignata levis

DvG DvG AN
EC-Podc, MX-PE PE-PntL, PE-BO,BR(Pa) PE-, PE-BO,BR(Pa)

46
Tribu - Tribe
Eumaeini
377 378 379
Ignata mulsus Ignata norax Michaelus ira

DvG DvG DvG
PE-ARCC, PE-BR(Pa),GF PE-ARCC, MX-AR PE-Jgu, MX-AR
380 381 382
Michaelus ira Michaelus jebus Michaelus jebus

DvG AN BBt
PE-ARCC, MX-AR PE-, MX-AR EC-Podc, MX-AR
383 384 385
Michaelus jebus Michaelus jebus Michaelus jebus

DvG DvG
PE-Qtz, MX-AR PE-VCm, MX-AR CO-VPN, 1450m MX-AR
386 387 388
Michaelus jebus Michaelus joseph Michaelus joseph

DvG DvG
CO-VPN, 1450m MX-AR PE-ARCC, CR-PE,BR PE-ARCC, CR-PE,BR

47
Tribu - Tribe
Eumaeini
389 390 391
Michaelus joseph Michaelus phoenissa Michaelus phoenissa

DvG DvG
PE-ARCC, CR-PE,BR EC-Podc, MX-PE,BR,GF CO-Tft, 574m MX-PE,BR,GF
392 393 394
Michaelus phoenissa Michaelus phoenissa Michaelus phoenissa

FM DvG
CO-CRC, 320m Ene10 MX-PE,BR,GF CO-RP, 560m Nov07 MX-PE,BR,GF PE-PntL, MX-PE,BR,GF
395 396 397
Michaelus phoenissa Michaelus thordesa Michaelus thordesa

DvG DvG
CO-Tft, 574m MX-PE,BR,GF PE-Qtz, MX-BO,AR PE-ARCC, MX-BO,AR
398 399 400
Michaelus thordesa Oenomaus andi Oenomaus andi

DvG DvG AN
PE-ARCC, MX-BO,AR PE-Qtz, EC-PE PE-VCm, EC-PE

48
Tribu - Tribe
Eumaeini
401 402 403
Oenomaus atena Oenomaus atena Oenomaus atesa

DvG DvG DvG
PE-ARCC, CR-PE,VZ,BR PE-ARCC, CR-PE,VZ,BR EC-BBR, MX-PE,BR,GF
404 405 406
Oenomaus atesa Oenomaus cyanovenata Oenomaus cyanovenata

DvG
EC-BBR, MX-PE,BR,GF CO-QLH, 700m Jun06 PN-BR,GY,GF CO-QLH, 700m Jun06 PN-BR,GY,GF
407 408 409
Oenomaus floreus Oenomaus gemma Oenomaus gemma

MCC Porthecla KG Porthecla DvG
BR-, VZ-BR(Pa) CO-CCri, 280m Ago13 CO-PE,BR PE-CJL, CO-PE,BR
410 411 412
Oenomaus gemma Oenomaus jauffreti Oenomaus lea

Porthecla KG DvG DvG
CO-CCri, 280m Ago13 CO-PE,BR PE-Qtz, PE-BR(Pa),GF PE-ARCC, Oct09 EC-PE

49
Tribu - Tribe
Eumaeini
413 414 415
Oenomaus minyia Oenomaus minyia Oenomaus minyia

Porthecla DvG Porthecla SDM Porthecla WC
EC-Atrk, VZ-PE,BR(Am),GF BR-CJL, VZ-PE,BR(Am),GF BR-CJL, VZ-PE,BR(Am),GF
416 417 418
Oenomaus moseri Oenomaus ortygnus Oenomaus ortygnus

MCC DvG DW
BR-, Nov15 BR(SC) PE-ARCC, MX-BO,BR PN-Bqt, MX-BO,BR
419 420 421
Oenomaus ortygnus Oenomaus ortygnus Oenomaus porthura

DGz Porthecla PL
CO-RlT, MX-BO,BR CO-, - MX-BO,BR CO-, HN-W EC
422 423 424
Oenomaus ravus Oenomaus ravus Olynthus essus

Porthecla MCC Porthecla MCC RC
BR-, BR-PY,GF BR-, BR-PY,GF BR-CJL, PE-BR,SU,GF

50
Tribu - Tribe
Eumaeini
425 426 427
Olynthus lorea Olynthus narbal Olynthus narbal

DvG BB AN
PE-MWC, PE-BR,SU,GF CO-Mor, PN-PE,VZ,BR,GF EC-Atrk, PN-PE,VZ,BR,GF
428 429 430
Olynthus narbal Olynthus narbal Olynthus ophelia

TS
PN-CTwr, PN-PE,VZ,BR,GF CO-Mor, 80m May09 PN-PE,VZ,BR,GF CO-Mco, CO-PE,BR,AR,GF
431 432 433
Olynthus ophelia Olynthus ophelia Parrhasius m-album

DvG
PE-ARCC, CO-PE,BR,AR,GF CO-Mco, CO-PE,BR,AR,GF CO-, - CN-US,MX?
434 435 436
Parrhasius orgia Parrhasius orgia Parrhasius orgia

DvG DvG
CO-Pña, MX-AR CO-MMt, 200m MX-AR PE-ARCC, MX-AR

51
Tribu - Tribe
Eumaeini
437 438 439
Parrhasius orgia Parrhasius orgia Parrhasius polibetes

DvG DvG
PE-ARCC, MX-AR PE-Qtz, MX-AR CO-Gurc, 690m - MX-AR
440 441 442
Parrhasius polibetes Parrhasius polibetes Parrhasius polibetes

DvG
PE-Qtz, MX-AR CO-PSJ, 1800m Feb07 MX-AR CO-PSJ, 1800m Feb07 MX-AR
443 444 445
Parrhasius polibetes Parrhasius polibetes Parrhasius polibetes

DvG
CO-Jard, MX-AR PE-Jul, MX-AR CO-HLB, 524m - MX-AR
446 447 448
Parrhasius polibetes Parrhasius selika Parrhasius selika

FM
CO-, May31 MX-AR CO-Jard, CO-BO,BR CO-QEV, 1940m CO-BO,BR

52
Tribu - Tribe
Eumaeini
449 450 451
Parrhasius selika Parrhasius selika Parrhasius selika

AN PB
PE-Qtz, CO-BO,BR CO-QEV, 1940m CO-BO,BR CO-, 2600m Oct13 CO-BO,BR
452 453 454
Parrhasius selika Parrhasius selika Symbiopsis aprica

DvG
CO-PSJ, 1800m Ago06 CO-BO,BR CO-PSJ, 1800m Ago06 CO-BO,BR PE-Qtz, Nov11 CO?EC-PE,BR,SU
455 456 457
Symbiopsis aprica Symbiopsis aprica Symbiopsis lenitas

THn DvG MCC
PE-, CO?EC-PE,BR,SU PE-ARCC, Sep25 CO?EC-PE,BR,SU BR-, May05 BR-AR
458 459 460
Symbiopsis pentas Symbiopsis perulera Symbiopsis perulera

KK DvG DvG
PE-Rmpt, Oct01 PE?-BO,BR PE-Rmpt, Sep27 PE-BR PE-Rmpt, Sep30 PE-BR

53
Tribu - Tribe
Eumaeini
461 462 463
Symbiopsis perulera Symbiopsis pupilla Symbiopsis pupilla

DvG
PE-ARCC, Oct07 PE-BR CO-RSm, 400m Jul19 CO-BO,BR,GY,GF CO-RSm, 400m Jul19 CO-BO,BR,GY,GF
464 465 466
Symbiopsis rickmani Symbiopsis strenua Symbiopsis strenua

DvG DvG DvG
PN-PplRd, Apr20 NI-CO,GF AR-Yab, Jan27 BR-AR,PY AR-Yab, Jan20 BR-AR,PY
467 468 469
Symbiopsis strenua Symbiopsis tanais ♂ Symbiopsis tanais ♂

DvG
AR-Yab, Jan29 BR-AR,PY CO-PSJ, 1800m CR-W EC CO-PSJ, 1800m CR-W EC
470 471 472
Symbiopsis tanais Symbiopsis tanais Symbiopsis tanais

MM BB
CO-, CR-W EC CO-, CR-W EC CO-SJn, 800m Sep12 CR-W EC

54
Tribu - Tribe
Eumaeini
473 474
Thepytus echelta Thepytus thyrea

KG JAR
BR-Rgu, MX-BR,GF BR-Blm, EC-PE,BR,AR,GF
475 476
Subtribu - Subtribe
Ipideclina
Ipidecla crepundia Ipidecla crepundia

DvG AN
BO-Volc, PE-BO,BR,GF PE-VCm, PE-BO,BR,GF
477 478 479
Ipidecla crepundia Ipidecla miadora ♀ Ipidecla schausi ♂

AN DvG
PE-, PE-BO,BR,GF PN-Fort, Apr26 W MX CO-Gai, 582m MX-AR
480 481
Ipidecla schausi ♀ Ipidecla schausi ♀

DvG MM
PE-ARCC, MX-AR CO-Tft, 574m MX-AR

55
Tribu - Tribe
Eumaeini
482 483
Subtribu - Subtribe
Calycopidina
Arzecla albolineata Arzecla arza

DvG AN
CO-Mzm, 1600m Oct01 CO-PE PE-, MX-BO,BR,AR
484 485 486
Arzecla arza Arzecla arza Arzecla arza

DvG BBt
CO-PSJ, 1800m Nov14 MX-BO,BR,AR PE-ARCC, MX-BO,BR,AR CO-Sin, 700m May21 MX-BO,BR,AR
487 488 489
Arzecla arza Arzecla calatia Arzecla calatia

KG DvG
CO-PSJ, 1850m Oct17 MX-BO,BR,AR PE-ARCC, MX,NI-PE,BR,GF PE-MWC, MX,NI-PE,BR,GF
490 491 492
Arzecla canacha Arzecla canacha Arzecla canacha

KK
CO-Jard, VZ-PE CO-Pña, VZ-PE BO-Volc, VZ-PE

56
Tribu - Tribe
Eumaeini
493 494 495
Arzecla canacha Arzecla canacha Arzecla canacha

RW DvG DvG
EC-, VZ-PE CO-, 1200m Ene12 VZ-PE EC-Loj, VZ-PE
496 497 498
Arzecla morpheus Arzecla morpheus Arzecla paralus

DvG DvG
EC-Mnd, Dec06 EC EC-Mnd, Dec06 EC CO-PSJ, 1800m GT-PE
499 500 501
Arzecla paralus Arzecla sethon Arzecla taminella

DvG FM DvG
PE-CoR, GT-PE CO-RPF, 3000m MX-VZ PE-ARCC, MX-BO,BR,GF,AR
502 503 504
Arzecla taminella Arzecla taminella Arzecla taminella

KK DvG DvG
EC-Ten, MX-BO,BR,GF,AR PE-ARCC, MX-BO,BR,GF,AR CO-Mzm, 1400m MX-BO,BR,GF,AR

57
Tribu - Tribe
Eumaeini
505 506 507
Arzecla tucumanensis Arzecla tucumanensis Arzecla tucumanensis

DvG MCC KK
PE-TngM, PE-BO,BR,AR BR-, PE-BO,BR,AR PE-Qtz, PE-BO,BR,AR
508 509 510
Arzecla tucumanensis Badecla badaca ♀ Badecla badaca

DvG Lamprospilus DvG Lamprospilus DvG
PE-TngM, PE-BO,BR,AR PE-PntL, VZ-BO,BR,AR BO-Cor, VZ-BO,BR,AR
511 512 513
Badecla badaca Badecla badaca Badecla badaca

Lamprospilus DvG Lamprospilus MIW Lamprospilus AH
PE-ARCC, VZ-BO,BR,AR BR-, VZ-BO,BR,AR CO-, Feb01 VZ-BO,BR,AR
514 515 516
Badecla lanckena Badecla lanckena Badecla sp

DvG DvG DvG
PE-ARCC, PE-BO PE-CoR, PE-BO PE-ARCC, PE

58
Tribu - Tribe
Eumaeini
517 518 519
Calycopis anastasia ♀ Calycopis anastasia Calycopis anthora

DvG DvG FM
BO-LHt, PE-BO BO-Pcy, PE-BO CO-RP, 560m CO-BO,BR
520 521 522
Calycopis anthora Calycopis atnius Calycopis atnius

JZ DvG DvG
EC-, CO-BO,BR PE-PntL, CR-PE,BR,GF PE-PntL, CR-PE,BR,GF
523 524 525
Calycopis atnius Calycopis atnius Calycopis atnius

DvG GC
CO-, - CR-PE,BR,GF PE-ARCC, CR-PE,BR,GF BR-CJL, CR-PE,BR,GF
526 527 528
Calycopis bactra Calycopis bactra Calycopis bactra

AH XD DvG
CO-SM, NI-PE,VZ,GF VZ-, NI-PE,VZ,GF PN-PplRd, NI-PE,VZ,GF

59
Tribu - Tribe
Eumaeini
529 530 531
Calycopis bellera ♀ Calycopis bellera Calycopis bellera

KG FM AN
PE-ARCC, PN-BO,BR,GF,AR CO-RP, 560m Feb09 PN-BO,BR,GF,AR EC-Atrk, PN-BO,BR,GF,AR
532 533 534
Calycopis bellera Calycopis bellera Calycopis bellera

DvG DvG
PE-Pro, PN-BO,BR,GF,AR EC-Orq, PN-BO,BR,GF,AR CO-Pña, PN-BO,BR,GF,AR
535 536 537
Calycopis buphonia Calycopis buphonia Calycopis buphonia

DvG DvG
PE-ARCC, CR-PE,BR,GF CO-CRC, 320m Ene12 CR-PE,BR,GF EC-PmpL, CR-PE,BR,GF
538 539 540
Calycopis buphonia Calycopis caesaries Calycopis caesaries

AN KG DvG
PE-, CR-PE,BR,GF CO-CRC, 320m Ago31 BZ-PE,BR PE-ARCC, BZ-PE,BR

60
Tribu - Tribe
Eumaeini
541 542 543
Calycopis calus Calycopis calus Calycopis calus

DvG
CO-PSJ, 1800m Abr26 MX-BO,AR CO-, MX-BO,AR EC-Podc, MX-BO,AR
544 545 546
Calycopis calus Calycopis calus Calycopis caulonia

DvG
CO-, Dic31 MX-BO,AR CO-, Dic31 MX-BO,AR AR-ChM, PE-BO,BR,AR
547 548 549
Calycopis caulonia Calycopis centoripa ♂ Calycopis centoripa ♂

DvG DvG AN
AR-ChM, PE-BO,BR,AR EC-AJL, 400m CO-BO,BR PE-, CO-BO,BR
550 551 552
Calycopis centoripa ♀ Calycopis centoripa ♀ Calycopis cerata

DvG GC
PE-Aco, CO-BO,BR BR-CJL, CO-BO,BR CO-HFr, 1100m Mar11 HN-AR

61
Tribu - Tribe
Eumaeini
553 554 555
Calycopis cerata Calycopis cerata Calycopis cerata

DvG
CO-RSm, 400m Jul19 HN-AR CO-HFr, 1100m Mar11 HN-AR CO-CRC, 550m HN-AR
556 557 558
Calycopis cerata Calycopis cerata Calycopis cicero

DvG DvG DvG
PE-ARCC, HN-AR PE-ARCC, HN-AR PE-PNYC, EC-PE
559 560 561
Calycopis cicero Calycopis clarina Calycopis clarina

DvG BBt KG
PE-Jgu, EC-PE MX-Mue, Aug11 MX MX-Teo, Jul06 MX
562 563 564
Calycopis clarina Calycopis demonassa Calycopis demonassa

KG
MX-Mue, Aug10 MX CO-, - MX-BO,BR,GF CO-VPN, 1450m MX-BO,BR,GF

62
Tribu - Tribe
Eumaeini
565 566 567
Calycopis demonassa Calycopis demonassa Calycopis demonassa
CO-CRC, 350m Ago23 MX-BO,BR,GF CO-Sin, 700m Nov01 MX-BO,BR,GF CO-VPN, 1450m Sep21 MX-BO,BR,GF
568 569 570
Calycopis demonassa Calycopis demonassa Calycopis gentilla ♂

DvG
CO-VPN, 1450m Sep21 MX-BO,BR,GF CO-Sin, 700m Nov01 MX-BO,BR,GF AR-ChM, PE,BR(RJ)-AR
571 572 573
Calycopis gentilla Calycopis isobeon Calycopis isobeon

DvG
BO-Chl, PE,BR(RJ)-AR CO-HLB, 524m Jul31 US(Tx)-EC,BR CO-Sin, 780m Ago23 US(Tx)-EC,BR
574 575 576
Calycopis isobeon Calycopis isobeon Calycopis isobeon
CO-RRm, 320m US(Tx)-EC,BR CO-HLB, 524m - US(Tx)-EC,BR CO-, - US(Tx)-EC,BR

63
Tribu - Tribe
Eumaeini
577 578 579
Calycopis isobeon Calycopis isobeon Calycopis janeirica

MCC
CO-, US(Tx)-EC,BR CO-, US(Tx)-EC,BR BR-, BR(RJ)-AR
580 581 582
Calycopis johnsoni Calycopis johnsoni Calycopis lerbela

RRt DvG DvG
EC-Cpl, CO-BO PE-Qtz, CO-BO GY-, EC-BR(Pa),GF
583 584 585
Calycopis malta Calycopis mimas Calycopis mimas

DvG DvG DvG
PE-PH, CO?EC-BO,BR EC-Podc, PN-PE,BR,GF EC-Podc, PN-PE,BR,GF
586 587 588
Calycopis nicolayi Calycopis orcilla Calycopis orcillula ♀

DvG DvG DvG
BR-SBr, PE-BO,BR(Pb) PE-Qtz, CO?EC-BO PN-PplRd, HN-PE,BR,VZ

64
Tribu - Tribe
Eumaeini
589 590 591
Calycopis orcillula Calycopis orcillula Calycopis orcillula

DvG AN DvG
PN-PplRd, HN-PE,BR,VZ EC-Atrk, HN-PE,BR,VZ PN-PplRd, HN-PE,BR,VZ
592 593 594
Calycopis origo Calycopis origo Calycopis partunda

DvG DvG DvG
PE-, CR-BO,BR,GF PN-PplRd, CR-BO,BR,GF PE-PNYC, PE-BR
595 596 597
Calycopis petaurister Calycopis pisis Calycopis pisis

DvG DvG
CO-VCC, 1640m Oct11 VZ-PE,GY PN-PplRd, PN-PE,BR PE-ARCC, PN-PE,BR
598 599 600
Calycopis sp1 Calycopis sp3 Calycopis sp4

LEM DvG THn
CO-HLB, 524m - - CO-Mzm, 1450m - CO-CRC, 320m -

65
Tribu - Tribe
Eumaeini
601 602 603
CO-RSm, 400m Ago02 - CO-RSm, 400m Ago02 - CO-FCy, -
604 605 606
CO-FCy, - CO-FCy, - CO-Sin, 700m May24 -
607 608 609
Calycopis sp9 Calycopis suda Calycopis suda

KK KK
CO-Sin, 700m Jun01 - PE-BSht, PE-BO PE-BSht, PE-BO
610 611 612
Calycopis sullivani Calycopis sullivani Calycopis talama

DvG DvG MCC
CO-Mzm, 1400m CR-PN, Atlantic slope CO-Mzm, 1400m CR-PN, Atlantic slope BR-, BR(RJ)

66
Tribu - Tribe
Eumaeini
613 614 615
Calycopis tamos Calycopis tamos Calycopis thama

DvG DvG DvG
CO-Mzm, 1400m CR-PE CO-Mzm, 1400m CR-PE PN-PplRd, May03 CR-CO
616 617 618
Calycopis thama Calycopis thama Calycopis trebula

DvG DvG KG
PN-PplRd, May03 CR-CO PN-PplRd, May03 CR-CO HN-, MX-PE,BR
619 620 621
Calycopis trebula Calycopis trebula Calycopis trebula

DvG DvG DvG
PE-ARCC, MX-PE,BR PE-ARCC, MX-PE,BR PE-VCm, MX-PE,BR
622 623 624
Calycopis wolfii Calycopis wolfii Calycopis xeneta ♂

DvG DvG DvG
PE-ARCC, CO-PE,BR,GF PE-ARCC, CO-PE,BR,GF PE-ARCC, NI-PE,VZ,GF

67
Tribu - Tribe
Eumaeini
625 626 627
Calycopis xeneta ♀ Calycopis xeneta Calycopis xeneta

KK DvG DvG
PE-Pro, NI-PE,VZ,GF PE-PNYC, NI-PE,VZ,GF PE-ARCC, NI-PE,VZ,GF
628 629 630
Camissecla camissa Camissecla camissa Camissecla cleocha

DvG RC
PE-VCm, CO? EC-PE PE-TRC, CO? EC-PE CO-PSJ, 1800m Nov21 CO-PE
631 632 633
Camissecla cleocha Camissecla cleocha Camissecla cleocha

KG DvG KG
PE-Qtz, 1150m CO-PE PE-Qtz, 1150m CO-PE CO-Afl, CO-PE
634 635 636
Camissecla charichlorus Camissecla melma Camissecla melma

SS DvG DvG
PE-PntL, HN-EC CO-Mzm, 1350m CR-PN,CO? CO-Mzm, 1350m CR-PN,CO?

68
Tribu - Tribe
Eumaeini
637 638 639
Camissecla pactya Camissecla pactya Camissecla pactya

DvG JHg MC
PE-, CO-BO EC-WS, CO-BO CO-, 2090m CO-BO
640 641 642
Camissecla sp1 Camissecla sp2 Camissecla sp2

FM FM
CO-PSJ, 1800m - CO-RP, 560m Feb09 - CO-RP, 560m Feb09 -
643 644 645
Camissecla verbenaca Camissecla verbenaca Camissecla vesper

DvG DvG DvG
CO-Mzm, VZ-PE,BR CO-Mzm, VZ-PE,BR PE-VCm, CO-BO,BR,GF
646 647 648
Camissecla vesper Camissecla vesper Camissecla vesper

DvG DvG DvG
PE-ARCC, CO-BO,BR,GF CO-Mzm, 1400m CO-BO,BR,GF CO-Mzm, 1400m CO-BO,BR,GF

69
Tribu - Tribe
Eumaeini
649 650 651
Electrostrymon denarius Electrostrymon denarius Electrostrymon endymion

DW DW RH
PN-Bqt, Jun03 MX-PN CO? PN-Bqt, Jun03 MX-PN CO? BR-CJL, PE-BO,BR,AR
652 653 654
Electrostrymon hugon ♂ Electrostrymon hugon ♂ Electrostrymon hugon ♂
CO-Sin, 780m Jun14 US(Tx)-VZ CO-Sin, 780m Jun14 US(Tx)-VZ CO-Sin, 750m Jun14 US(Tx)-VZ
655 656 657
Electrostrymon hugon ♂ Electrostrymon hugon ♀ Electrostrymon hugon
CO-Sin, 750m Jun14 US(Tx)-VZ CO-Sin, 700m Jul19 US(Tx)-VZ CO-VCC, 1650m Ene24 US(Tx)-VZ
658 659 660
Electrostrymon hugon Electrostrymon joya Electrostrymon joya

DvG DvG DvG
PN-PplRd, May03 US(Tx)-VZ EC-Vilc, Nov30 US(Tx)-PE PE-Cus, Aug22 US(Tx)-PE

70
Tribu - Tribe
Eumaeini
661 662 663
Electrostrymon perisus Electrostrymon picoloro Electrostrymon thurman

DvG DvG DvG
CO-, 2600m CO-VZ,EC CO-Mzm, 2500m W CO PN-Tot, Apr27 CR-W PN
664 665 666
Electrostrymon thurman Gigantorubra aruma ♀ Gigantorubra aruma

DvG Arumecla DvG Arumecla DvG
PN-Tot, Apr27 CR-W PN PE-CoR, CO-PE,BR,GF PE-ARCC, CO-PE,BR,GF
667 668 669
Gigantorubra aruma Gigantorubra collucia Gigantorubra collucia

Arumecla AH Lamprospilus Lamprospilus BBt
CO-SM, Feb05 CO-PE,BR,GF CO-CRC, 350m Feb04 MX-EC,VZ CO-Sin, 700m May21 MX-EC,VZ
670 671 672
Gigantorubra collucia Gigantorubra galliena Gigantorubra netesca ♂♀

Lamprospilus DvG Arumecla DW Arumecla RnM
CO-Mzm, Sep30 MX-EC,VZ PN-Bqt, MX-EC,VZ VZ-, PE-BO,GY

71
Tribu - Tribe
Eumaeini
673 674 675
Gigantorubra orcidia ♂ Gigantorubra orcidia ♂ Gigantorubra orcidia ♀

Lamprospilus DvG Lamprospilus DvG Lamprospilus DvG
PE-ARCC, MX-BO,BR,PY,GF PE-ARCC, MX-BO,BR,PY,GF PE-ARCC, MX-BO,BR,PY,GF
676 677 678
Gigantorubra orcidia Gigantorubra orcidia Kisutam syllis

Lamprospilus Lamprospilus THn FM
CO-HLB, 524m - MX-BO,BR,PY,GF PE-, MX-BO,BR,PY,GF CO-, May31 US(Tx)-PE,BR,GF
679 680 681
Kisutam syllis Kisutam syllis Kisutam syllis

DvG DvG
CO-, US(Tx)-PE,BR,GF EC-Podc, US(Tx)-PE,BR,GF PE-MWC, US(Tx)-PE,BR,GF
682 683 684
Kisutam syllis Lamprospilus coelicolor ♂ Lamprospilus coelicolor ♀

KK AN KK
CO-MMt, US(Tx)-PE,BR,GF PE-, MX-BO,BR,GF PE-Qtz, MX-BO,BR,GF

72
Tribu - Tribe
Eumaeini
685 686 687
Lamprospilus coelicolor ♀ Lamprospilus coelicolor ♀ Lamprospilus decorata ♀

DvG AN DvG
CO-Mzm, 1300m Sep30 MX-BO,BR,GF PE-Qtz, MX-BO,BR,GF PE-, CO-PE
688 689 690
Lamprospilus decorata Lamprospilus draudti ♂ Lamprospilus draudti ♂

DvG AN Leo
PE-VCm, CO-PE CO-QEV, 1850m Feb27 CO-PE CO-Mzm, CO-PE
691 692 693
Lamprospilus draudti ♀ Lamprospilus draudti ♀ Lamprospilus genius ♂

DvG DvG
CO-PSJ, 1800m Feb07 CO-PE CO-Mzm, 2100m CO-PE BR-SBr, EC-PE,BR,GF
694 695 696
Lamprospilus genius ♀ Lamprospilus genius Lamprospilus nicetus

KK DBn THn
BR-, EC-PE,BR,GF PE-ARCC, EC-PE,BR,GF EC-, VZ-BO

73
Tribu - Tribe
Eumaeini
697 698 699
Lamprospilus nicetus Lamprospilus occidentalis Lamprospilus occidentalis

DvG DvG LEM
PE-ARCC, VZ-BO CO-Mzm, 1350m CO CO-, 2000m CO
700 701 702
Rubroserrata ecbatana ♂ Rubroserrata ecbatana ♀ Rubroserrata ecbatana

DvG DvG
PE-ARCC, VZ-PE,BR,GF PE-TngM, VZ-PE,BR,GF CO-CRC, 320m Ene26 VZ-PE,BR,GF
703 704 705
Rubroserrata ecbatana Rubroserrata ecbatana Rubroserrata ecbatana
CO-VCC, 1650m Mar18 VZ-PE,BR,GF CO-Pña, VZ-PE,BR,GF CO-SNc, 474m Nov23 VZ-PE,BR,GF
706 707 708
Rubroserrata mathewi ♂ Ziegleria ceromia Ziegleria ceromia

DvG
PN-PplRd, May03 MX-PN,CO?BR? CO-Sin, 780m May24 MX-BO,BR,UR,GF CO-Sin, 780m May24 MX-BO,BR,UR,GF

74
Tribu - Tribe
Eumaeini
709 710 711
Ziegleria ceromia Ziegleria hesperitis Ziegleria hesperitis

DvG
CO-Sin, 780m May21 MX-BO,BR,UR,GF CO-VCC, 1650m Mar18 MX-BO,BR,AR,GF CO-VCC, 1650m Mar18 MX-BO,BR,AR,GF
712 713
Ziegleria hesperitis Ziegleria hesperitis

LC
EC-Atrk, Oct31 MX-BO,BR,AR,GF CO-, MX-BO,BR,AR,GF
714 715
Subtribu - Subtribe
Strymonina
Arawacus dolylas Arawacus dolylas
CO-VPN, 1450m PN-BR,AR CO-VPN, 1450m PN-BR,AR
716 717 718
Arawacus dolylas Arawacus dumenilii Arawacus dumenilii

FM LG
CO-, - PN-BR,AR CO-, HN-BR,GY PA-CTwr, HN-BR,GY

75
Tribu - Tribe
Eumaeini
719 720 721
Arawacus dumenilii Arawacus dumenilii Arawacus dumenilii

DvG DvG
CO-QLH, 750m Oct24 HN-BR,GY CO-RAD, May19 HN-BR,GY CO-RAD, May19 HN-BR,GY
722 723 724
Arawacus ellida Arawacus ellida Arawacus ellida

TH KG KG
BO-Corc, Nov16 VZ-BO,BR,AR PE-Rmp-Mq, Nov14 VZ-BO,BR,AR PE-Rmp-Mq, Nov14 VZ-BO,BR,AR
725 726 727
Arawacus hypocrita Arawacus jada Arawacus jada

WC KG JK
MX-CNv, Apr25 MX-CR MX-Teo, Oct07 US(Az)-CR MX-Oax, Aug11 US(Az)-CR
728 729 730
Arawacus jada Arawacus leucogyna ♂ Arawacus leucogyna ♂

JK BBt
MX-Oax, Jul20 US(Az)-CR CO-Afl, 1400m BZ-EC,VZ CO-QEV, 1920m BZ-EC,VZ

76
Tribu - Tribe
Eumaeini
731 732 733
Arawacus leucogyna ♂ Arawacus leucogyna ♀ Arawacus leucogyna

Leo
CO-QEV, 1920m BZ-EC,VZ CO-Mzm, BZ-EC,VZ CO-, - BZ-EC,VZ
734 735 736
Arawacus leucogyna Arawacus lincoides Arawacus lincoides
CO-PSJ, 1800m BZ-EC,VZ CO-RSm, 400m Ago02 CR-EC,VZ,GF CO-VPN, 1450m CR-EC,VZ,GF
737 738 739
Arawacus lincoides Arawacus lincoides Arawacus separata

GM KWs KK
CO-CRC, 320m Ago22 CR-EC,VZ,GF CO-, CR-EC,VZ,GF CO-MMt, 200m Oct17 CO-AR
740 741 742
Arawacus separata Arawacus sito Arawacus sito

FM KD KD
CO-RP, 560m Ago01 CO-AR MX-GF, Nov26 MX-CR MX-Ord, Nov20 MX-CR

77
Tribu - Tribe
Eumaeini
743 744 745
Arawacus sito Arawacus sp1 Arawacus tarania

KG DvG MCC
MX-Mue, Jun05 MX-CR CO-Bvt, 200m - BR-, Abr17 BO-BR(MG),AR
746 747 748
Arawacus togarna Arawacus togarna Hypostrymon aepeona

KG
CO-, - MX-Amz CO-LPC, 800m MX-Amz PE-BSht, PE
749 750 751
Hypostrymon asa Hypostrymon asa Hypostrymon asa

DvG DvG DvG
EC-Podc, GT-PE,BR,GF,AR CO-MMt, 200m GT-PE,BR,GF,AR EC-Atrk, GT-PE,BR,GF,AR
752 753 754
Hypostrymon asa Hypostrymon renidens Hypostrymon renidens

DvG DvG DvG
PE-ARCC, GT-PE,BR,GF,AR PE-CoR, CO?EC-PE CO-RAA, CO?EC-PE

78
Tribu - Tribe
Eumaeini
755 756 757
Panthiades aeolus Panthiades aeolus Panthiades aeolus

FM DvG DvG
CO-RNM, 150m Jun21 PN-BO,BR PE-Pro, PN-BO,BR CO-MMt, 200m Mar06 PN-BO,BR
758 759 760
Panthiades aeolus Panthiades bathildis Panthiades bathildis

DvG
PE-Rmp-Mq, PN-BO,BR CO-, MX-EC,BR,GF CO-Tkk, 5m Jun17 MX-EC,BR,GF
761 762 763
Panthiades bathildis Panthiades bitias ♂ Panthiades bitias ♂

RC
PE-TRC, MX-EC,BR,GF CO-HLB, 524m - MX-BO,BR,GF CO-RSm, 400m Ago02 MX-BO,BR,GF
764 765 766
Panthiades bitias ♂ Panthiades bitias ♂ Panthiades bitias ♀
CO-RSm, 400m Jul19 MX-BO,BR,GF CO-QLH, 750m Jul25 MX-BO,BR,GF CO-Rch, 1000m Feb06 MX-BO,BR,GF

79
Tribu - Tribe
Eumaeini
767 768 769
Panthiades bitias ♀ Panthiades bitias ♀ Panthiades bitias ♀

SS
CO-MMt, 200m MX-BO,BR,GF CO-HLB, 524m - MX-BO,BR,GF CO-, - MX-BO,BR,GF
770 771 772
Panthiades bitias ♀ Panthiades boreas Panthiades boreas

GC GC
CO-HLB, 524m Ago18 MX-BO,BR,GF BR-CJL, CO-BR,GF BR-CJL, CO-BR,GF
773 774 775
Panthiades hebraeus Panthiades hebraeus Panthiades hebraeus

MCC AH MCC
BR-, BR(Ba)-AR BR-Rgu, BR(Ba)-AR BR-, BR(Ba)-AR
776 777 778
Panthiades ochus Panthiades paphlagon Panthiades paphlagon

PT KG KK
PN-, MX-EC PE-, CO-PE,VZ BO-Volc, 1400m CO-PE,VZ

80
Tribu - Tribe
Eumaeini
779 780 781
Panthiades paphlagon Panthiades paphlagon Panthiades paphlagon

LC DvG
CO-PSJ, 1800m CO-PE,VZ CO-Mzm, CO-PE,VZ PE-PNYC, CO-PE,VZ
782 783 784
Panthiades phaleros Panthiades phaleros Panthiades phaleros

KG DvG KG
PA-, MX-AR PE-ARCC, MX-AR PE-ARCC, MX-AR
785 786 787
Panthiades phaleros Panthiades phaleros Rekoa marius

DvG KG DvG
PE-ARCC, MX-AR PA-, MX-AR PE-PH, US(Tx)-AR
788 789 790
Rekoa marius Rekoa marius Rekoa marius

KK PS
PE-Pro, US(Tx)-AR CO-, US(Tx)-AR PE-PNYC, US(Tx)-AR

81
Tribu - Tribe
Eumaeini
791 792 793
Rekoa meton Rekoa meton Rekoa meton

DvG
EC-Podc, MX-AR CO-HLB, 524m - MX-AR CO-Sin, 780m Nov15 MX-AR
794 795 796
Rekoa meton Rekoa palegon Rekoa palegon

DvG DvG
PE-VCm, MX-AR CO-Mzm, 1350m US(Tx)-AR CO-, - US(Tx)-AR
797 798 799
Rekoa palegon Rekoa palegon Rekoa palegon

DvG DvG
CO-, US(Tx)-AR CO-Mzm, 1350m US(Tx)-AR EC-Pro, US(Tx)-AR
800 801 802
Rekoa palegon Rekoa palegon Rekoa palegon

DvG KK
CO-Jard, 1700m May30 US(Tx)-AR PE-Qtz, US(Tx)-AR EC-Atrk, US(Tx)-AR

82
Tribu - Tribe
Eumaeini
803 804 805
Rekoa stagira ♀ Rekoa stagira Strymon albata

DvG
CO-, - MX-AR PE-AMZ, Oct26 MX-AR CO-, - US(Tx)-VZ
806 807 808
Strymon albata Strymon albata Strymon albata
CO-HLB, 524m - US(Tx)-VZ CO-RSD, US(Tx)-VZ CO-Tft, 574m US(Tx)-VZ
809 810 811
Strymon alea ♂ Strymon alea ♂ Strymon alea

JK JK BBc
MX-Oax, Sep19 US(Tx)-VZ MX-Oax, Sep19 US(Tx)-VZ US-, US(Tx)-VZ
812 813 814
Strymon alea Strymon alea Strymon astiocha

WC BBc KG
MX-CNv, Nov05 US(Tx)-VZ US-, US(Tx)-VZ BO-Chl, Nov15 MX-AR

83
Tribu - Tribe
Eumaeini
815 816 817
Strymon astiocha Strymon astiocha Strymon astiocha

DW KG KG
PE-Hual, Nov16 MX-AR PE-Hual, Nov16 MX-AR BO-Chl, Nov15 MX-AR
818 819 820
Strymon astiocha Strymon bazochii bazochii Strymon bazochii bazochii

DW
PE-Hual, Nov16 MX-AR CO-VPN, 1450m Feb19 US(Tx)-AR CO-, - US(Tx)-AR
821 822 823
Strymon bazochii bazochii Strymon bazochii bazochii Strymon bazochii bazochii
CO-, US(Tx)-AR CO-Gai, 600m US(Tx)-AR CO-Sin, 780m Jun14 US(Tx)-AR
824 825 826
Strymon bazochii bazochii Strymon bazochii bazochii Strymon bebrycia

RL
CO-Sin, 780m Jun14 US(Tx)-AR CO-, - US(Tx)-AR MX-CNv, Nov15 US(Tx)-CR

84
Tribu - Tribe
Eumaeini
827 828 829
Strymon bebrycia Strymon bebrycia Strymon bebrycia

MS KG KD
MX-CNv, Nov18 US(Tx)-CR MX-Mue, May14 US(Tx)-CR MX-CNv, Nov18 US(Tx)-CR
830 831 832
Strymon bubastus bubastus ♂ Strymon bubastus bubastus ♂ Strymon bubastus bubastus
CO-, VZ-PE,BR,AR CO-, VZ-PE,BR,AR CO-HLB, 524m Oct24 VZ-PE,BR,AR
833 834 835
Strymon bubastus bubastus Strymon bubastus bubastus Strymon bubastus bubastus
CO-HLB, 524m - VZ-PE,BR,AR CO-, - VZ-PE,BR,AR CO-CCri, 300m Ago06 VZ-PE,BR,AR
836 837 838
Strymon bubastus bubastus Strymon bubastus bubastus Strymon bubastus bubastus

DvG
CO-, - VZ-PE,BR,AR CO-Sin, 700m May21 VZ-PE,BR,AR CO-Tft, 574m VZ-PE,BR,AR

85
Tribu - Tribe
Eumaeini
839 840 841
Strymon bubastus bubastus Strymon cestri Strymon cestri
CO-HLB, 524m Oct24 VZ-PE,BR,AR CO-, US(Az,Tx)-BO,BR,AR CO-, US(Az,Tx)-BO,BR,AR
842 843 844
Strymon cestri Strymon colombiana Strymon crambusa

KG KG
CO-, US(Az,Tx)-BO,BR,AR CO-, CO BR-RG, Nov21 PE-BO,BR,AR
845 846 847
Strymon crambusa Strymon crambusa Strymon crambusa

DvG DvG DvG
PE-Rmpt, Sep24 PE-BO,BR,AR PE-Rmpt, Sep24 PE-BO,BR,AR PE-Rmpt, Sep24 PE-BO,BR,AR
848 849 850
Strymon crambusa Strymon crambusa Strymon daraba

KG DvG DvG
BR-RG, Nov21 PE-BO,BR,AR PE-Rmpt, Sep24 PE-BO,BR,AR PE-, EC-PE

86
Tribu - Tribe
Eumaeini
851 852 853
Strymon daraba Strymon daraba Strymon daraba

KK KK DvG
PE-, EC-PE PE-, EC-PE PE-, EC-PE
854 855 856
Strymon daraba Strymon davara Strymon eurytulus ♀

DvG AN DvG
PE-, EC-PE PE-, EC-BO,BR AR-ChM, Jan11 BO-AR,CH,PY
857 858 859
Strymon gabatha ♀ Strymon gabatha ♀ Strymon gabatha

DvG DvG
CO-Mzm, MX-PE,VZ CO-Mzm, MX-PE,VZ CO-CRC, 320m MX-PE,VZ
860 861 862
Strymon gabatha Strymon istapa istapa Strymon istapa istapa

DvG KG KG
CO-Mzm, 1350m Sep25 MX-PE,VZ MX-Cat, MX-AR MX-Plq, MX-AR

87
Tribu - Tribe
Eumaeini
863 864 865
Strymon istapa istapa Strymon istapa istapa Strymon megarus

DvG DvG
BR-, MX-AR PE-, MX-AR CO-CRC, 320m Ene12 MX-BO,BR,AR
866 867 868
Strymon megarus Strymon megarus Strymon melinus ♂

FM BBn
CO-RP, 560m Ago01 MX-BO,BR,AR CO-CRC, 320m Ene12 MX-BO,BR,AR MX-Ctm, Jun03 US-EC,VZ
869 870 871
Strymon melinus ♂ Strymon melinus Strymon melinus

BBn KG KG
MX-Ctm, Jun03 US-EC,VZ TX-RGv, Dec30 US-EC,VZ MX-Ctm, Jun03 US-EC,VZ
872 873 874
Strymon melinus caldasensis Strymon melinus caldasensis Strymon melinus caldasensis

DvG DvG DvG
CO-Mzm, CO CO-, 1200m Ene12 CO CO-Mzm, CO

88
Tribu - Tribe
Eumaeini
875 876 877
Strymon melinus caldasensis Strymon mulucha Strymon mulucha
CO-Sin, 780m Oct10 CO CO-VCC, 1650m Mar18 MX-BO,BR,AR CO-VCC, 1650m Mar18 MX-BO,BR,AR
878 879 880
Strymon mulucha Strymon mulucha Strymon mulucha
CO-Tft, 574m MX-BO,BR,AR CO-RSm, 400m Ago02 MX-BO,BR,AR CO-RSD, MX-BO,BR,AR
881 882 883
CO-PSJ, 1800m Dic05 MX-BO,BR,AR CO-PSJ, 1800m MX-BO,BR,AR CO-, MX-BO,BR,AR
884 885 886
CO-, MX-BO,BR,AR CO-, - MX-BO,BR,AR CO-SNc, 474m Dic07 MX-BO,BR,AR

89
Tribu - Tribe
Eumaeini
887 888 889
Strymon oreala ♀ Strymon rufofusca Strymon rufofusca

DvG
BR-SBc, Jan30 BR(Ba,SP)-AR CO-SNc, 474m Dic14 US(Tx)-AR CO-SNc, 474m Nov16 US(Tx)-AR
890 891 892
Strymon rufofusca Strymon rufofusca Strymon rufofusca
CO-Sin, 780m Ago23 US(Tx)-AR CO-Gai, 582m Ago24 US(Tx)-AR CO-, US(Tx)-AR
893 894 895
Strymon rufofusca Strymon rufofusca Strymon rufofusca

LEM
CO-, US(Tx)-AR CO-Gai, 582m Ago24 US(Tx)-AR CO-HLB, 524m - US(Tx)-AR
896 897 898
Strymon rufofusca Strymon rufofusca Strymon sp1
CO-Gai, 582m Ago24 US(Tx)-AR CO-Sin, 780m Ago23 US(Tx)-AR CO-HLB, 524m - -

90
Tribu - Tribe
Eumaeini
899 900 901
Strymon sp7 Strymon yojoa Strymon yojoa

DvG
CO-, - - CO-Mnc, 600m US(Tx,Az)-PE,BR,GF CO-SNc, 474m Nov23 US(Tx,Az)-PE,BR,GF
902 903 904
Strymon yojoa Strymon yojoa Strymon yojoa

DvG
CO-SNc, 474m Nov23 US(Tx,Az)-PE,BR,GF CO-SNc, 474m Nov23 US(Tx,Az)-PE,BR,GF CO-Mnc, 600m US(Tx,Az)-PE,BR,GF
905 906 907
Strymon ziba Strymon ziba Strymon ziba
CO-Sin, 700m Sep20 MX-BO,AR,GF CO-Sin, 780m Ago23 MX-BO,AR,GF CO-Sin, 780m Ago23 MX-BO,AR,GF
908 909 910
Strymon ziba Strymon ziba Strymon ziba
CO-, - MX-BO,AR,GF CO-, MX-BO,AR,GF CO-Mor, 80m May01 MX-BO,AR,GF

91
Tribu - Tribe
Eumaeini
911 912 913
Thereus cithonius Thereus cithonius Thereus cithonius

Noreena FM Noreena FM Noreena FM
CO-RP, 560m Nov07 MX-AR CO-, May31 MX-AR CO-RP, 560m Ago01 MX-AR
914 915 916
Thereus endera Thereus endera Thereus lausus

DvG RW JK
CO-MMt, CO-PE,BR(Am) PE-, CO-PE,BR(Am) MX-Oax, Aug30 MX-BR,GY,Amz
917 918 919
Thereus ortalus Thereus ortalus Thereus pedusa

BBn KG Pedusa JJ
MX-Ctm, MX-BO,BR,PY MX-Ctm, MX-BO,BR,PY CO-, 250m May12 PN-PE,BR
920 921
Thereus pedusa Thereus timoclea

Pedusa AN GS
PE-, PN-PE,BR BO-, PN-BO,GF

92
Tribu - Tribe
Eumaeini
922 923
Subtribu - Subtribe
Strephonotina
Aubergina alda Aubergina hesychia

AN DvG
PE-, CO-BO,BR(Pa),GF CO-Mzm, 1400m Jun09 CR-PE,BR
924 925 926
Aubergina hesychia Aubergina paetus Aubergina paetus

AN SS
PE-, CR-PE,BR CO-CRC, 320m Ago30 MX-CO CO-FCy, MX-CO
927 928 929
Aubergina paetus Aubergina vanessoides Aubergina vanessoides
CO-FCy, MX-CO CO-LPC, 800m CO-BO,BR,AR CO-LPC, 800m CO-BO,BR,AR
930 931 932
Aubergina vanessoides Celmia anastomosis Celmia celmus

THn
CO-, CO-BO,BR,AR PE-, PE-BR,Guianas CO-VPN, 1450m Sep21 MX-AR

93
Tribu - Tribe
Eumaeini
933 934 935
Celmia celmus Celmia celmus Celmia celmus

KK
CO-MMt, 200m Oct16 MX-AR CO-VPN, 1450m Sep21 MX-AR CO-Mor, 80m May01 MX-AR
936 937 938
Celmia color Celmia conoveria Celmia conoveria

DvG DvG AN
PE-, PE-BO,BR(Am),GY BR-, MX-PE,BR,GF EC-Atrk, MX-PE,BR,GF
939 940 941
Celmia mecrida Celmia sp Celmia uzza

JB DvG DvG
BR-CJL, VZ-PE,BR,Guianas PE-ARCC, BR-SBc, SE BR
942 943 944
Gargina caninius ♂ Gargina caninius ♂ Gargina caninius ♂

DvG DvG DvG
PE-PntL, MX-BO,BR,GF PE-, MX-BO,BR,GF PE-PntL, MX-BO,BR,GF

94
Tribu - Tribe
Eumaeini
945 946 947
Gargina caninius Gargina caninius Gargina emessa ♀

KK
CO-VPN, 1450m MX-BO,BR,GF EC-Ten, MX-BO,BR,GF CO-RSm, 400m Jul19 NI-PE,BR,GF
948 949 950
Gargina emessa ♀ Gargina emessa ♀ Gargina emessa

GM
CO-RSm, 400m Ago02 NI-PE,BR,GF CO-RSm, 400m Jul19 NI-PE,BR,GF CO-CRC, 320m Ago21 NI-PE,BR,GF
951 952 953
Gargina gargophia Gargina gargophia Gargina gargophia

DvG
CO-VCC, 1650m Mar18 MX-BO,BR,GF CO-Mzm, 1400m MX-BO,BR,GF CO-PSJ, 1800m May15 MX-BO,BR,GF
954 955 956
Gargina gargophia Gargina gnosia ♀ Gargina gnosia

FM DW DvG
CO-RP, 560m Feb09 MX-BO,BR,GF BO-VTn, Nov23 MX-BO,AR,GF BO-Chl, Nov08 MX-BO,AR,GF

95
Tribu - Tribe
Eumaeini
957 958 959
Gargina panchaea Gargina panchaea Gargina panchaea

DvG DvG DvG
PE-PMp, PE-BR,AR PE-Jgu, PE-BR,AR PE-Jul, PE-BR,AR
960 961 962
Gargina thyesta ♀ Gargina thyesta ♀ Gossenia gossei

DvG DvG OE
CO-PGt, CR-BR CO-PGt, CR-BR CO-, HN-CO
963 964 965
Gossenia lycabas Gossenia lycabas Gossenia lycabas

Thestius DvG Thestius DvG Thestius DvG
PE-VCm, Oct09 CO-PE,BR PE-AMZ, Oct17 CO-PE,BR PE-AMZ, CO-PE,BR
966 967 968
Gossenia lycabas Iaspis andersoni Iaspis andersoni

Thestius DvG
PE-ARCC, CO-PE,BR CO-VPN, 1450m Sep21 MX-CO CO-VPN, 1450m MX-CO

96
Tribu - Tribe
Eumaeini
969 970 971
Iaspis andersoni Iaspis castitas Iaspis exiguus

DW GC
CO-VPN, 1450m MX-CO BR-RG, PE-BR(Pa,ES),GF BR-CJL, SU-BR
972 973 974
Iaspis talayra Iaspis temesa Iaspis temesa

JAR DvG DvG
VZ-Mdb, Jun15 SE BR-PY PE-ARCC, MX-PE,BR,GY GY-, MX-PE,BR,GY
975 976 977
Iaspis thabena Iaspis thabena Iaspis verania

FM PB DvG
CO-RP, 560m Nov07 CO-BR CR-Lsel, May09 CO-BR EC-Orq, Nov23 EC-BR,GF
978 979 980
Iaspis verania Ministrymon arola Ministrymon arola

DvG
EC-Orq, Nov23 EC-BR,GF CO-Sin, 780m Ago30 MX-CR,CO? CO-Sin, 780m Oct10 MX-CR,CO?

97
Tribu - Tribe
Eumaeini
981 982 983
Ministrymon arola Ministrymon arthuri Ministrymon arthuri

DvG DvG
CO-Sin, 700m Jul26 MX-CR,CO? PE-VCm, EC-PE,BR(RJ) PE-ARCC, EC-PE,BR(RJ)
984 985 986
Ministrymon arthuri Ministrymon azia Ministrymon azia

DvG KG
PE-PntL, EC-PE,BR(RJ) MX-, US(Az)-BO,BR,AR CO-PSJ, 1800m US(Az)-BO,BR,AR
987 988 989
Ministrymon azia Ministrymon azia Ministrymon azia
CO-PSJ, 1800m US(Az)-BO,BR,AR CO-PSJ, 1800m Feb07 US(Az)-BO,BR,AR CO-PSJ, 1800m Abr26 US(Az)-BO,BR,AR
990 991 992
Ministrymon azia Ministrymon azia Ministrymon azia

DvG DvG DvG
PE-ARCC, US(Az)-BO,BR,AR EC-Vilc, US(Az)-BO,BR,AR EC-Vilc, US(Az)-BO,BR,AR

98
Tribu - Tribe
Eumaeini
993 994 995
Ministrymon azia Ministrymon azia Ministrymon cleon

WC DvG
MX-, US(Az)-BO,BR,AR CO-PSJ, 1800m Dic05 US(Az)-BO,BR,AR PE-Qtz, CR-BO,BR,AR
996 997 998
Ministrymon cleon Ministrymon cleon Ministrymon clytie

DvG
CO-PSJ, 1800m CR-BO,BR,AR EC-Orq, CR-BO,BR,AR CO-, US(Tx)-CR
999 1000 1001
Ministrymon coronta Ministrymon coronta Ministrymon cruenta

KG BBn DvG
MX-Ctm, Jun06 MX-Guianas MX-Ctm, Jun06 MX-Guianas EC-Vilc, CO-BO,AR,GF
1002 1003 1004
Ministrymon cruenta Ministrymon inoa Ministrymon inoa

DvG
EC-Vilc, CO-BO,AR,GF CO-PSJ, 1800m Nov21 MX-VZ CO-PSJ, 1800m Nov21 MX-VZ

99
Tribu - Tribe
Eumaeini
1005 1006 1007
Ministrymon inoa Ministrymon inoa Ministrymon inoa
CO-PSJ, 1800m Nov21 MX-VZ CO-Sin, 700m Jul19 MX-VZ CO-Sin, 700m Jul19 MX-VZ
1008 1009 1010
Ministrymon inoa Ministrymon leda Ministrymon leda

KG KG
CO-Sin, 700m Jul19 MX-VZ MX-Teo, Jul06 SW US-MX MX-Teo, Jul06 SW US-MX
1011 1012 1013
Ministrymon megacles Ministrymon megacles Ministrymon phrutus

DvG DvG
PE-ARCC, VZ-BO,BR,SU,GF PE-ARCC, VZ-BO,BR,SU,GF CO-Sin, 780m Oct10 MX-BO,BR
1014 1015 1016
Ministrymon phrutus Ministrymon phrutus Ministrymon phrutus

LEM
CO-Sin, 780m Ago23 MX-BO,BR CO-Gai, 582m MX-BO,BR CO-HLB, 524m - MX-BO,BR

100
Tribu - Tribe
Eumaeini
1017 1018 1019
Ministrymon phrutus Ministrymon una scopas Ministrymon una scopas

DvG DvG
CO-, - MX-BO,BR PN-Boc, Apr23 MX-EC PN-Boc, Apr27 MX-EC
1020 1021 1022
Ministrymon una una Ministrymon una una Ministrymon una una

KK
CO-, - CO-BO,BR,AR CO-Rmo, 1830m CO-BO,BR,AR CO-SM, Oct11 CO-BO,BR,AR
1023 1024 1025
Ministrymon una una Ministrymon una una Ministrymon una una
CO-Mor, 80m May01 CO-BO,BR,AR CO-Mor, 80m May01 CO-BO,BR,AR CO-, - CO-BO,BR,AR
1026 1027 1028
Ministrymon una una Ministrymon zilda Ministrymon zilda

DvG DvG
CO-, - CO-BO,BR,AR PE-ARCC, MX-BO,BR,GF PE-ARCC, MX-BO,BR,GF

101
Tribu - Tribe
Eumaeini
1029 1030 1031
Ministrymon zilda Nicolaea bagrada Nicolaea besidia

MCC DvG
CO-Mco, MX-BO,BR,GF BR-, VZ-BR(Am) PE-, VZ-PE,BR,GF
1032 1033 1034
Nicolaea besidia Nicolaea cupa ♀ Nicolaea cupa

JL KG DvG
VZ-, VZ-PE,BR,GF BR-, BR(RS,SC)-PY AR-ChM, BR(RS,SC)-PY
1035 1036 1037
Nicolaea dolium Nicolaea lemuria Nicolaea lemuria

AH DvG DvG
CO-SM, Feb15 MX-BO,BR CO-Mzm, 1400m Jun09 CO-BR,VZ,GF PE-ARCC, CO-BR,VZ,GF
1038 1039 1040
Nicolaea lemuria Nicolaea lemuria Nicolaea lemuria

DvG DvG DvG
CO-Mzm, 1400m Jun09 CO-BR,VZ,GF PE-PNYC, CO-BR,VZ,GF CO-Mzm, 1400m Jun09 CO-BR,VZ,GF

102
Tribu - Tribe
Eumaeini
1041 1042 1043
Nicolaea lemuria Nicolaea lemuria Nicolaea munditia

DvG
CO-RSm, 400m Jul19 CO-BR,VZ,GF CO-RSm, 400m Jul19 CO-BR,VZ,GF PE-VCm, EC-BO,Guianas
1044 1045 1046
Nicolaea munditia Nicolaea munditia Nicolaea pyxis

DvG DvG DvG
PE-VCm, EC-BO,Guianas PE-VCm, EC-BO,Guianas PE-PNYC, CO-PE,BR,GF
1047 1048 1049
Nicolaea pyxis Nicolaea schausa Nicolaea velina

DvG MCC Bsu
PE-Qtz, CO-PE,BR,GF BR-, BR(Pr) MX-, MX-BR,GF
1050 1051 1052
Ostrinotes empusa Ostrinotes empusa Ostrinotes empusa

KK DvG DvG
PE-Jgu, PN-AR PE-, 650m PN-AR PE-, 650m PN-AR

103
Tribu - Tribe
Eumaeini
1053 1054 1055
Ostrinotes empusa Ostrinotes gentiana Ostrinotes halciones

DvG KK DvG
CO-Mzm, 1400m Sep23 PN-AR PE-Rmp-Mq, CO-BO,BR CO-Mzm, 1400m Sep23 MX-PE,BR
1056 1057 1058
Ostrinotes halciones Ostrinotes keila Ostrinotes keila

KK
PE-Rmp-Mq, MX-PE,BR CO-PSJ, 1800m Dic05 MX-CO CO-VCC, 1650m Ene24 MX-CO
1059 1060 1061
Ostrinotes keila Ostrinotes keila Ostrinotes keila
CO-VCC, 1650m Mar23 MX-CO CO-, - MX-CO CO-PSJ, 1800m Ago06 MX-CO
1062 1063 1064
Ostrinotes keila Ostrinotes keila Ostrinotes sophocles

HB
CO-PSJ, 1800m Dic05 MX-CO CO-VCC, 1650m Mar23 MX-CO BR-, Ene24 BR-AR

104
Tribu - Tribe
Eumaeini
1065 1066 1067
Ostrinotes sophocles Ostrinotes sospes Ostrinotes tarena

BBt DvG KK
BR-, Ene25 BR-AR PE-ARCC, CO-PE PE-Qtz, VZ-PE,BR,GF
1068 1069 1070
Ostrinotes tympania Ostrinotes tympania Ostrinotes tympania

DvG DvG KK
PE-ARCC, EC-PE,BR(Pa),GF EC-Podc, EC-PE,BR(Pa),GF PE-Jgu, EC-PE,BR(Pa),GF
1071 1072 1073
Ostrinotes tympania Ostrinotes tympania Ostrinotes tympania

DvG BB
CO-Mco, EC-PE,BR(Pa),GF PE-PMp, EC-PE,BR(Pa),GF CO-, EC-PE,BR(Pa),GF
1074 1075 1076
Siderus athymbra Siderus athymbra Siderus bouvieri

DvG DvG DBn
PE-ARCC, CO-PE,BR,VZ,GF GY-, CO-PE,BR,VZ,GF PE-ARCC, EC-PE

105
Tribu - Tribe
Eumaeini
1077 1078 1079
Siderus bouvieri Siderus giapor Siderus giapor

DvG DvG DvG
PE-, EC-PE AR-ChM, Dec24 BR(SP)-PY,AR AR-ChM, Dec24 BR(SP)-PY,AR
1080 1081 1082
Siderus leucophaeus ♀ Siderus leucophaeus ♀ Strephonota acameda

DvG DvG DvG
PE-ARCC, Nov11 GT-PE,BR,GF PE-ARCC, Nov11 GT-PE,BR,GF PE-VCm, PE-BO,BR(Pa)
1083 1084 1085
Strephonota adela Strephonota agrippa Strephonota ambrax

GC AN Jpe
BR-CJL, PE-BR(Rd),GF EC-Atrk, Oct24 VZ-BO,BR(Pa) CO-Sin, 700m Jul05 MX-PE,BR,AR
1086 1087 1088
Strephonota ambrax Strephonota ambrax Strephonota azurinus

DvG DvG DvG
CO-Sin, 700m May21 MX-PE,BR,AR PE-ARCC, MX-PE,BR,AR PE-Qtz, CR-PE

106
Tribu - Tribe
Eumaeini
1089 1090 1091
Strephonota carteia ♂ Strephonota carteia ♀ Strephonota carteia ♀

DvG AN DvG
PE-ARCC, CO-PE,BR,GY EC-PmpL, CO-PE,BR,GY PE-ARCC, CO-PE,BR,GY
1092 1093 1094
Strephonota carteia Strephonota cyllarissus Strephonota cyllarissus

DvG AN KK
PE-ARCC, CO-PE,BR,GY PE-VCm, CO-PE,BR,SU PE-Qtz, 1050m Nov22 CO-PE,BR,SU
1095 1096 1097
Strephonota cyllarissus Strephonota cyllarissus Strephonota elika

KK GC DvG
CO-MMt, 200m Oct15 CO-PE,BR,SU BR-CJL, CO-PE,BR,SU BR-PNI, Feb03 PE-BR,AR
1098 1099 1100
Strephonota foyi Strephonota foyi Strephonota foyi

KK DvG DvG
PE-Rmp-Mq, PE-BO,BR PE-, PE-BO,BR PE-ARCC, PE-BO,BR

107
Tribu - Tribe
Eumaeini
1101 1102 1103
Strephonota foyi Strephonota jactator Strephonota metanira

DvG JAR Exorbaetta DvG
PE-Jgu, PE-BO,BR BR-, PE-BO,BR,PY,AR PE-Qtz, CO-BO,BR(Am)
1104 1105 1106
Strephonota metanira Strephonota parvipuncta Strephonota parvipuncta

Exorbaetta DvG Robustana DvG Robustana DvG
PE-Qtz, CO-BO,BR(Am) PE-VCm, CO-PE,BR,GF PE-ARCC, CO-PE,BR,GF
1107 1108 1109
Strephonota porphyritis ♀ Strephonota porphyritis Strephonota porphyritis

Serratonotes DvG Serratonotes DvG Serratonotes SDM
PE-ARCC, PE-BR(Pa),GY GY-, PE-BR(Pa),GY BR-, PE-BR(Pa),GY
1110 1111 1112
Strephonota prina Strephonota sphinx Strephonota sphinx

DvG FM DvG
PE-ARCC, CO-PE CO-, May31 PN-AR PE-Qtz, Nov26 PN-AR

108
Tribu - Tribe
Eumaeini
1113 1114 1115
Strephonota sphinx Strephonota sphinx Strephonota strephon

DvG KP DvG
AR-ChM, Jan20 PN-AR PN-Vant, Mar07 PN-AR PE-ARCC, CO-PE,BR,Guianas
1116 1117 1118
Strephonota strephon Strephonota strephon Strephonota strephon

DvG DvG DvG
PE-MWC, CO-PE,BR,Guianas PE-ARCC, CO-PE,BR,Guianas PE-ARCC, CO-PE,BR,Guianas
1119 1120 1121
Strephonota syedra Strephonota syedra Strephonota syedra

Syedranota DvG Syedranota DvG Syedranota GC
GY-, MX-PE,BR,GF PE-, MX-PE,BR,GF BR-CJL, MX-PE,BR,GF
1122 1123 1124
Strephonota tephraeus Strephonota tephraeus Strephonota tephraeus

DvG DvG AN
EC-Atrk, US(Tx)-PE,BR,AR PE-ARCC, US(Tx)-PE,BR,AR PE-PntL, US(Tx)-PE,BR,AR

109
Tribu - Tribe
Eumaeini
1125 1126 1127
Strephonota tephraeus Strephonota tephraeus Strephonota tephraeus

KK
PE-Qtz, US(Tx)-PE,BR,AR CO-Sin, 780m Ago16 US(Tx)-PE,BR,AR CO-VCC, 1650m Ene24 US(Tx)-PE,BR,AR
1128 1129 1130
Strephonota trebonia Strephonota tyriam Terenthina bradyae

Treboniana LC Diminutina RH
EC-Yach, Apr11 EC-PE BR-CJL, PE-BR(Pa),GF CO-Sin, 780m Oct10 CO-EC
1131 1132 1133
Terenthina bradyae Terenthina terentia ♂ Terenthina terentia ♂

DvG DvG FM
CO-Mzm, 1350m CO-EC EC-Atrk, CR-PE,BR,GF CO-RP, 560m Nov07 CR-PE,BR,GF
1134 1135 1136
Terenthina terentia ♀ Terenthina terentia ♀ Terenthina terentia ♀

DvG
CO-CRC, 320m CR-PE,BR,GF CO-CRC, 320m CR-PE,BR,GF EC-Atrk, CR-PE,BR,GF

110
Tribu - Tribe
Eumaeini
1137 1138 1139
Theclopsis demea ♀ Theclopsis demea ♀ Theclopsis epidius ♀

BBt DvG
CO-CRC, 320m Jun21 NI-EC PE-Qtz, NI-EC CO-QLH, 750m Jun06 PN-CO
1140 1141 1142
Theclopsis epidius ♀ Theclopsis epidius Theclopsis gargara

DvG DvG
CO-QLH, 750m Jun06 PN-CO BO-Corc, Nov19 PN-CO EC-Orq, CO-PE,BR,VZ,GF
1143 1144 1145
Theclopsis gargara Theclopsis gargara Theclopsis gargara

DvG DvG DvG
PE-Pro, CO-PE,BR,VZ,GF PE-Jgu, CO-PE,BR,VZ,GF PE-MWC, CO-PE,BR,VZ,GF
1146 1147 1148
Theclopsis lydus ♂ Theclopsis lydus ♀ Theclopsis lydus ♀

DvG DvG
PE-PntL, VZ-BO,BR,GY,GF EC-Cpl, VZ-BO,BR,GY,GF CO-RSD, VZ-BO,BR,GY,GF

111
Tribu - Tribe
Eumaeini
1149 1150 1151
Theclopsis lydus ♀ Theclopsis lydus Theclopsis lydus

DvG MM FM
CO-Bvt, 200m VZ-BO,BR,GY,GF CO-Gvr, Ene13 VZ-BO,BR,GY,GF CO-RP, 560m Ago01 VZ-BO,BR,GY,GF
1152 1153 1154
Theclopsis lydus Theclopsis lydus Theclopsis lydus

AN AN KNc
EC-Atrk, VZ-BO,BR,GY,GF EC-Atrk, Oct30 VZ-BO,BR,GY,GF PE-PNM, VZ-BO,BR,GY,GF
1155 1156 1157
Theclopsis mycon Theclopsis mycon Theclopsis mycon

MS AH
MX-Mue, MX-VZ,GF CO-, - MX-VZ,GF CO-, MX-VZ,GF
1158 1159 1160
Tmolus crolinus Tmolus crolinus Tmolus crolinus

JK JK DvG
MX-Oax, Aug05 NI-VZ MX-Oax, Jun04 NI-VZ PE-ARCC, Sep22 NI-VZ

112
Tribu - Tribe
Eumaeini
1161 1162 1163
Tmolus cydrara Tmolus cydrara Tmolus cydrara

DvG DvG FM
PE-ARCC, MX-BO,S BR,AR,GF PE-ARCC, MX-BO,S BR,AR,GF CO-RP, 560m Ago01 MX-BO,S BR,AR,GF
1164 1165 1166
Tmolus cydrara Tmolus echion Tmolus echion

KK DvG
CO-MMt, 200m Oct20 MX-BO,S BR,AR,GF CO-PSJ, 1850m Oct17 MX-BO,S BR,AR,GF CO-Mzm, MX-BO,S BR,AR,GF
1167 1168 1169
Tmolus echion Tmolus echion Tmolus echion
CO-GMq, 5m MX-BO,S BR,AR,GF CO-VCC, 1650m Ene25 MX-BO,S BR,AR,GF CO-Sin, 780m Sep06 MX-BO,S BR,AR,GF
1170 1171 1172
CO-Mor, 80m May01 MX-BO,S BR,AR,GF CO-PSJ, 1850m Oct17 MX-BO,S BR,AR,GF CO-PSJ, 1850m Oct17 MX-BO,S BR,AR,GF

113
Tribu - Tribe
Eumaeini
1173 1174 1175
CO-HLB, 524m - MX-BO,S BR,AR,GF CO-GMq, 5m Dic29 MX-BO,S BR,AR,GF CO-PSJ, 1800m Feb22 MX-BO,S BR,AR,GF
1176 1177 1178
Tmolus mutina Tmolus mutina Tmolus mutina

DvG DvG GC
PE-ARCC, CR-PE,N BR,GY,GF PE-ARCC, CR-PE,N BR,GY,GF BR-, CR-PE,N BR,GY,GF
1179 1180
Tmolus ufentina Tmolus ufentina

DvG DvG
PE-Qtz, Nov21 CO-BO,BR(Am),GF PE-Qtz, Nov21 CO-BO,BR(Am),GF
1181
Subtribu - Subtribe
Trichonidina
Trichonis immaculata ♂
GC
BR-CJL, EC-PE,BR,GF

114
Tribu - Tribe
Eumaeini
1182 1183
Subtribu - Subtribe
Callophryidina
Cyanophrys acaste Cyanophrys acaste

DvG KG
AR-Yab, Jan29 BO-BR(RJ),AR CO-Oqy, 1850m Ago28 BO-BR(RJ),AR
1184 1185 1186
Cyanophrys acaste Cyanophrys acaste Cyanophrys agricolor

DvG DvG DW
BR-Rgu, Aug11 BO-BR(RJ),AR AR-Yab, Jan29 BO-BR(RJ),AR PN-Bqt, Jul30 MX-CO
1187 1188 1189
Cyanophrys agricolor Cyanophrys amyntor Cyanophrys amyntor

BBn DvG
MX-Oax, May29 MX-CO CO-Sin, 700m Jun14 MX-PE,BR PE-ARCC, MX-PE,BR
1190 1191 1192
Cyanophrys amyntor Cyanophrys amyntor Cyanophrys amyntor
CO-HLB, 524m - MX-PE,BR CO-HLB, 524m - MX-PE,BR CO-HLB, 524m - MX-PE,BR

115
Tribu - Tribe
Eumaeini
1193 1194 1195
Cyanophrys amyntor Cyanophrys amyntor Cyanophrys amyntor

DvG
CO-SJn, 800m Sep12 MX-PE,BR CO-Jard, MX-PE,BR PE-ARCC, MX-PE,BR
1196 1197 1198
Cyanophrys amyntor Cyanophrys banosensis Cyanophrys bertha

DvG KG BBt
PE-ARCC, MX-PE,BR PE-BSht, EC-PE MX-Mue, Aug10 BR(Pr)-AR
1199 1200 1201
Cyanophrys goodsoni Cyanophrys goodsoni Cyanophrys goodsoni
CO-Sin, 700m Jun14 US(Tx)-CR,CO? CO-, US(Tx)-CR,CO? CO-, US(Tx)-CR,CO?
1202 1203 1204
Cyanophrys herodotus Cyanophrys herodotus Cyanophrys herodotus

THn
CO-CRC, 320m Abr05 MX-AR CO-CRC, 320m MX-AR CO-, MX-AR

116
Tribu - Tribe
Eumaeini
1205 1206 1207
Cyanophrys herodotus Cyanophrys herodotus Cyanophrys longula

DvG
CO-Mco, MX-AR EC-Podc, MX-AR CO-Jard, MX-VZ
1208 1209 1210
Cyanophrys longula Cyanophrys miserabilis Cyanophrys miserabilis

KG KG
CO-Jard, MX-VZ MX-Teo, Jul04 US(Tx)-CR MX-Oax, Sep03 US(Tx)-CR
1211 1212 1213
Cyanophrys pseudolongula Cyanophrys pseudolongula Cyanophrys pseudolongula
CO-PSJ, 1800m CO-BO,VZ,AR CO-PSJ, 1800m CO-BO,VZ,AR CO-PSJ, 1800m Dic05 CO-BO,VZ,AR
1214 1215 1216
Cyanophrys pseudolongula Cyanophrys remus ♀ Cyanophrys remus

DvG DvG
CO-PSJ, 1800m CO-BO,VZ,AR AR-ChM, Dec25 BR-AR,UR AR-ChM, Jan23 BR-AR,UR

117
Tribu - Tribe
Eumaeini
1217 1218 1219
Cyanophrys remus Chalybs chloris Chalybs chloris

DvG DvG DvG
AR-Yab, Jan29 BR-AR,UR AR-ChM, BR-AR BR-PNI, BR-AR
1220 1221 1222
Chalybs hassan Chalybs hassan Chalybs hassan

DvG AN KK
EC-Orq, MX-PE,BR,GF,AR PE-, MX-PE,BR,GF,AR PE-Qtz, 1050m MX-PE,BR,GF,AR
1223 1224 1225
Chalybs hassan Chalybs janias Chalybs janias

DvG DvG FM
PE-ARCC, MX-PE,BR,GF,AR EC-Cpl, MX-PE,BR,GF CO-RP, 560m Ago23 MX-PE,BR,GF
1226 1227 1228
Chalybs janias Chalybs janias Chalybs lineata

GC DvG Smd
BR-CJL, MX-PE,BR,GF CO-Mzm, MX-PE,BR,GF CO-, 1900m Ago06 CO-EC,BR

118
Tribu - Tribe
Eumaeini
1229 1230 1231
Chlorostrymon simaethis Chlorostrymon simaethis Chlorostrymon telea

DvG
CO-Gai, 582m US(Tx)-BR,AR CO-, - US(Tx)-BR,AR PE-ARCC, US(Tx)-BO,BR,AR
1232 1233 1234
Chlorostrymon telea Chlorostrymon telea Erora aura ♀

KK DvG DvG
PE-, US(Tx)-BO,BR,AR PE-ARCC, US(Tx)-BO,BR,AR CO-Mzm, MX-CO
1235 1236 1237
Erora aura Erora badeta Erora badeta

DvG
CO-Mzm, 1400m MX-CO CO-RSm, 400m Ago02 CR-PE,BR,AR CO-RSm, 400m Ago02 CR-PE,BR,AR
1238 1239 1240
Erora badeta Erora badeta Erora badeta

AN KK
PE-, CR-PE,BR,AR CO-QLH, 750m Jul25 CR-PE,BR,AR EC-Atrk, CR-PE,BR,AR

119
Tribu - Tribe
Eumaeini
1241 1242 1243
Erora biblia Erora caespes ♀ Erora caespes

MCC THn AN
BR-, Ago13 VZ-BO,BR(Pa),PY CO-CRC, 320m CO?EC-BO PE-, CO?EC-BO
1244 1245 1246
Erora carla ♀ Erora carla ♀ Erora carla

RRt
CO-PSJ, 1800m MX-BR CO-PSJ, 1800m MX-BR CO-Mzm, MX-BR
1247 1248 1249
Erora carla Erora carla Erora carla
CO-Sin, 780m Ago16 MX-BR CO-PSJ, 1800m Nov21 MX-BR CO-PSJ, 1800m Nov21 MX-BR
1250 1251 1252

RRt DvG
CO-, MX-BR CO-Mzm, MX-BR CO-Mzm, 1350m MX-BR

120
Tribu - Tribe
Eumaeini
1253 1254 1255

DvG
CO-Mzm, 1350m MX-BR CO-PSJ, 1800m Nov21 MX-BR CO-Sin, 780m Ago16 MX-BR
1256 1257 1258
Erora carla Erora gabina Erora gabina

DvG DvG
CO-PSJ, 1800m Ago13 MX-BR PE-Qtz, MX-BO,BR,AR PE-Qtz, MX-BO,BR,AR
1259 1260 1261
Erora melba Erora melba Erora nitetis

MCC MCC BBn
BR-, Ene17 BR BR-, Ene17 BR MX-Oax, MX-PE
1262 1263 1264
Erora opisena ♀ Erora opisena ♀ Erora opisena ♀

KG KG
CO-, 2600m Oct13 MX-VZ CO-, 2600m Oct13 MX-VZ CO-QEV, 1900m MX-VZ

121
Tribu - Tribe
Eumaeini
1265 1266 1267
Erora opisena ♀ Erora phrosine Erora phrosine

KG DvG KG
CO-, 2600m Oct13 MX-VZ PE-Qtz, EC-BO PE-PNYC, EC-BO
1268 1269 1270
Erora sp1 Erora subflorens Magnastigma elsa

DvG DvG DvG
CO-Mzm, 1700m - CO-Mzm, MX-VZ PE-PNYC, MX-EC
1271 1272 1273
Magnastigma elsa Ocaria aholiba Ocaria aholiba

DvG DvG DvG
EC-Orq, MX-EC CO-RlT, CR-BO,VZ PE-CoR, CR-BO,VZ
1274 1275 1276
Ocaria aholiba Ocaria aholiba Ocaria aholiba

DvG AN DvG
PE-Rctl, CR-BO,VZ EC-Cpl, CR-BO,VZ EC-BBR, CR-BO,VZ

122
Tribu - Tribe
Eumaeini
1277 1278 1279
Ocaria aholiba Ocaria aholiba Ocaria arpoxais

KG DvG DvG
CO-RlT, CR-BO,VZ CO-Mzm, 1600m CR-BO,VZ CO-Mzm, MX-BO
1280 1281 1282
Ocaria calesia Ocaria calesia Ocaria calesia

DvG
CO-PSJ, 1800m CO-PE,AR CO-RlR, 1850m - CO-PE,AR CO-Mzm, Jun12 CO-PE,AR
1283 1284 1285
Ocaria calesia Ocaria calesia Ocaria calesia

DvG DvG DvG
CO-Mzm, 2050m CO-PE,AR EC-WS, CO-PE,AR PE-Rctl, CO-PE,AR
1286 1287 1288
Ocaria cinerea Ocaria cinerea Ocaria cinerea

DvG DvG DvG
EC-Atrk, EC,PE-BR(RS) PE-Jgu, EC,PE-BR(RS) PE-Qtz, EC,PE-BR(RS)

123
Tribu - Tribe
Eumaeini
1289 1290 1291
Ocaria clenchi Ocaria clenchi Ocaria clepsydra

DW JK DvG
MX-Ctm, Sep15 MX-SV MX-Oax, Jul31 MX-SV CO-Mzm, 1600m Jun24 CO-PE,VZ
1292 1293 1294
Ocaria elongata Ocaria elongata Ocaria elongata

BBt
CO-Ang, 1900m CO-BO CO-Ang, 1900m CO-BO EC-Baz, CO-BO
1295 1296 1297
Ocaria elongata Ocaria elongata Ocaria ocrisia

Leo KK
CO-Mzm, CO-BO PE-CoR, CO-BO CO-VPN, 1450m MX-AR
1298 1299 1300
Ocaria ocrisia Ocaria ocrisia Ocaria ocrisia

DvG DvG
EC-JSR, MX-AR EC-Cpl, MX-AR CO-Sin, 780m Nov01 MX-AR

124
Tribu - Tribe
Eumaeini
1301 1302 1303

FM
CO-RP, 560m Nov07 MX-AR CO-RlR, 1850m - MX-AR CO-QEV, 1975m MX-AR
1304 1305 1306
CO-Sin, 780m Nov01 MX-AR CO-, - MX-AR CO-QEV, 1975m MX-AR
1307 1308 1309
CO-Ang, 1900m MX-AR CO-, - MX-AR CO-, - MX-AR
1310 1311 1312

DvG
CO-, MX-AR CO-, MX-AR PE-ARCC, MX-AR

125
Tribu - Tribe
Eumaeini
1313 1314 1315
Ocaria ocrisia Ocaria sadei Ocaria sadei

DvG OE KG
PE-ARCC, MX-AR CO-, Feb06 CO-BO PE-Rmp-Mq, CO-BO
1316 1317 1318
Ocaria sadei Ocaria thales Ocaria thales

DvG GC
PE-Rmpt, CO-BO BR-CJL, Ago20 MX-AR CO-QLH, 750m Jul25 MX-AR
1319 1320 1321
Ocaria thales Ocaria thales Ocaria thales

DW KG DvG
PA-PplRd, MX-AR PE-PNYC, 1100m MX-AR PE-PNYC, 1000m MX-AR
1322 1323 1324
Semonina ares Semonina ares Thaeides muela

DvG DvG DvG
PE-Jgu, PN-PE,BR,GF PE-Jgu, PN-PE,BR,GF PE-Wyq, EC-BO,CH

126
Tribu - Tribe
Eumaeini
1325 1326 1327
Thaeides theia ♀ Thaeides theia Thaeides theia

DvG KNc
CO-RAD, May17 MX-BO,BR,AR CO-RlT, Ene06 MX-BO,BR,AR PE-PNM, MX-BO,BR,AR


127
128
Subfamilia Polyommatinae
Tribu - Tribe
Polyommatini
1 2
Subtribu - Subtribe
Sin Subtribu
Cupido comyntas texana ♂ Cupido comyntas texana
CO-Sin, 700m Sep20 US(Tx)-EC CO-GMq, 5m Dic29 US(Tx)-EC
3 4 5
Cupido comyntas texana Cupido comyntas texana Cupido comyntas texana
CO-, - US(Tx)-EC CO-Sin, 780m Oct10 US(Tx)-EC CO-VPN, 1450m US(Tx)-EC
6 7 8
Cupido comyntas texana Hemiargus ceraunus Hemiargus hanno bogotana ♂

FM
CO-, - US(Tx)-EC CO-, - SW US-PN,CO? CO-RPF, 3000m PN-CO
BioButterfly DataBase - Butterflycatalogs.com Polyommatinae

129
Tribu - Tribe
Polyommatini
9 10 11
Hemiargus hanno bogotana ♀ Hemiargus hanno bogotana ♀ Hemiargus hanno hanno ♂

FM FM
CO-RPF, 3000m PN-CO CO-RPF, 3000m PN-CO CO-, - CO-AR
12 13 14
Hemiargus hanno hanno ♀♂ Hemiargus hanno hanno ♀ Hemiargus hanno hanno
CO-VCC, 1650m Oct11 CO-AR CO-VCC, 1650m Oct11 CO-AR CO-VCC, 1650m Oct11 CO-AR
15 16 17
Hemiargus hanno hanno Hemiargus hanno hanno Hemiargus hanno hanno
CO-, CO-AR CO-PSJ, 1800m Feb07 CO-AR CO-VCC, 1650m Oct11 CO-AR
18 19 20
CO-VCC, 1650m Oct11 CO-AR CO-VCC, 1650m Oct11 CO-AR CO-, - CO-AR
BioButterfly DataBase - Butterflycatalogs.com Polyommatinae Página 2 de 6

130
Tribu - Tribe
Polyommatini
21 22 23
CO-, - CO-AR CO-, - CO-AR CO-, CO-AR
24 25 26
CO-PSJ, 1800m Feb07 CO-AR CO-VCC, 1650m Oct11 CO-AR CO-VCC, 1650m Oct11 CO-AR
27 28 29
Hemiargus martha Hemiargus martha Hemiargus ramon

DvG DvG DvG
EC-Vilc, EC EC-Vilc, EC EC-Vilc, EC-PE
30 31 32
Hemiargus ramon Hemiargus ramon Hemiargus ramon

DvG DvG DvG
PE-Lim, EC-PE EC-Vilc, EC-PE EC-Vilc, EC-PE

131
Tribu - Tribe
Polyommatini
33 34 35
Itylos nodo Itylos nodo Itylos nodo

Madeleinea DvG Madeleinea DvG Madeleinea DvG
EC-Pulu, EC EC-Pulu, EC EC-, EC
36 37 38
Itylos pacis Itylos pacis Leptotes andicola

Madeleinea AN Madeleinea AN DvG
PE-, PE-BO PE-, PE-BO EC-, CO-EC
39 40 41
Leptotes bathyllos Leptotes bathyllos Leptotes callanga

KK KK AN
EC-, Oct25 EC-PE EC-Ten, EC-PE PE-, PE-BO
42 43 44
Leptotes callanga Leptotes callanga Leptotes callanga

DvG DvG AN
PE-, PE-BO PE-Chtk, PE-BO EC-PaP, PE-BO

132
Tribu - Tribe
Polyommatini
45 46 47
Leptotes cassius Leptotes cassius Leptotes cassius
CO-VPN, 1450m US-VZ,BO,BR,AR,SU CO-, US-VZ,BO,BR,AR,SU CO-, - US-VZ,BO,BR,AR,SU
48 49 50
Leptotes cassius Leptotes cassius Leptotes cassius

DvG
CO-Mzm, 1450m US-VZ,BO,BR,AR,SU CO-GMq, 5m US-VZ,BO,BR,AR,SU CO-, - US-VZ,BO,BR,AR,SU
51 52 53
Leptotes lamasi Leptotes lamasi Nabokovia cuzquenha

DvG AN DvG
EC-Vilc, EC-PE PE-, EC-PE PE-, 3100m Jul16 PE
54 55 56
Nabokovia cuzquenha Zizula cyna Zizula cyna

DvG
PE-, 3100m Jul16 PE CO-, US-AR CO-, - US-AR

133
Tribu - Tribe
Polyommatini
57 58
Zizula cyna Zizula cyna

Leo
CO-Mzm, Jun05 US-AR CO-Sin, 780m Jun14 US-AR


134
Bibliografía Bibliography
El Neotrópico por su diversidad de mariposas tiene The Neotropics, due to its diversity of butterflies,
mucha información publicada en libros y revistas have a lot of information published in books and
especializadas, esta información nos ha ayudado a specialized magazines. This information has helped
construir el catálogo de BioButterfly Database y a us to build the BioButterfly Database catalog and to
identificar muchas de nuestras fotos. identify many of our photos.
Inicialmente estábamos colocando la bibliografía en Initially we were placing the bibliography in each PDF
cada PDF tratando de asociar los artículos relevantes trying to associate the relevant articles to each
a cada PDF especifico, básicamente si se relaciona specific PDF, basically if it related to a family, a
con una familia, un país o una localidad. Decidimos country or a locality. We decided to make a general
hacer una bibliografía general combinada en formato combined bibliography as a separate PDF with
PDF independiente con citas para todas las citations for all the publications we have been using
publicaciones que hemos estado usando de tal in such a way that it is useful for all our PDFs and at
manera que sea útil para todos nuestros PDF y al the same time it frees up space in all the PDFs.
mismo tiempo libere espacio en todos los PDF. With this link access the PDF with the bibliography:
Con el siguiente vinculo acceda al PDF con la https://www.butterflycatalogs.com/neotropical-
bibliografía.: bibliography.html
https://www.butterflycatalogs.com/neotropical-
bibliography.html If you want to delve into the Neotropical
bibliography, with articles on butterflies from Mexico
Si se quiere profundizar en la bibliografía del to Argentina, we recommend an excellent document
Neotrópico, con artículos de mariposas desde compiled by Dr. Gerardo Lamas 2022 found on the
México hasta Argentina, recomendamos un Butterflies of America BOA page called: An Annotated
excelente documento recopilado por el Dr. Gerardo Bibliography of the Neotropical Butterflies and
Lamas 2022 que se encuentra en la página de Skippers (Lepidoptera: Papilionoidea and
Butterflies of America BOA llamado: An Annotated Hesperioidea) Revised Electronic Edition which can
Bibliography of the Neotropical Butterflies and be consulted at the following link:
Skippers (Lepidoptera: Papilionoidea and
Hesperioidea) Revised Electronic Edition que se http://www.butterfliesofamerica.com/L/Biblio.htm
puede consultar en el siguiente vinculo: We are constantly looking for information to help us
http://www.butterfliesofamerica.com/L/Biblio.htm complete the database, especially changes in the
taxonomy of species, new species and species by
Permanentemente estamos buscando información country or region. We appreciate all the references
que nos ayude a completar la base de datos, that you can send us.
especialmente cambios en la taxonomía de las
especies, nuevas especies y especies por país o
regiones. Agradecemos todas las referencias que nos
puedan enviar.
135
Páginas web recomendadas Recommended web pages
Existen muchísimas fuentes de información sobre There are many sources of information about
mariposas en Internet, sin embargo, presentamos butterflies on the internet, however, we present
acá una selección de aquellas recomendadas que here a selection of those recommended that can
puedan dar información complementaria a la provide complementary information to the
bibliografía, las listas de chequeo o las guías, o bibliography, the checklists, or the guides, or may
pueden tener información útil para la identificación have useful information for the identification or
o el conocimiento de las mariposas. knowledge of butterflies.
Identificación de mariposas - Butterfly Otras - Other

identification https://v3.boldsystems.org (DNA barcodes)
https://www.butterflycatalogs.com (libros de http://tolweb.org/tree/
identificación para la región neotropical) https://www.inaturalist.org/
www.neotropicalbutterflies.com (Mariposas del https://www.nic.funet.fi/
Neotrópico)
https://www.butterfliesofamerica.com Tour de mariposas – Butterfly tours
fotografías de todos los tipos www.butterfliesofcolombia.com
http://www.mariposabutterflytours.com/
Información general de mariposas - General
butterfly information Specific Families or Genera:
https://www.learnaboutbutterflies.com http://parides.genus.free.fr/syst.html
http://www.heliconius.org/2011/neil-rossers-
Nombres - Names maps-preliminary-versions/
https://www.nhm.ac.uk/our- Euptychiina Checklist:
science/data/lepindex/ https://www2.ib.unicamp.br/labor/site/?page_i
https://www.nic.funet.fi/pub/sci/bio/life/insect d=1020
a/lepidoptera/
Plantas hospederas - Host plants

https://www.nhm.ac.uk/our-
science/data/hostplants/search/index.dsml
Inventarios regionales - Regional inventories

http://calydna.com excelente fotografía/E
http://www.sangay.eu/es (Parque
Nacional Sangay, Ecuador)
http://www.cotacachi.eu (Región de Cotacachi,
W Ecuador)
http://www.loja-zamora.eu/en (Ecuador)
https://www.noreste.eu/en/les-papillons (NE
Ecuador)
http://www.butterfliesofecuador.com/
https://papillons-de-guyane.fr/papillons/
136
Localidades de las fotografías de la familia
Photo localities for the family
Abreviacion y Nombre localidad Ubicación Altura Min- Max m
Argentina AR
ChM Chacra Mariposa Obera, Misiones 300 300
Yab Yaboti Biosphere Reserve San Pedro, Misiones 200 200
Bolivia BO
Cor Coroico Nor Yungas, La Paz 1800 2000
Corc Coroico Nor Yungas, La Paz 1800 2000
Crv Caranavi Nor Yungas, La Paz 800 1200
Chl Chulumani Chulumani, La Paz 1200 1800
Volc Refugio Los Volcanes Florida, Santa Cruz 1300 1300
VTn Villa Tunari Chapare, Cochabamba 500 900
Vtu above Villa Tunari Chipiri, Cochabamba 800 800
Brasil BR
Blm Blumenau Blumenau, Santa Catarina
CJL Cristalino Jungle Lodge Alta Floresta, Mato Grosso 200 200
Mdb Massaranduba Massaranduba, Santa Catarina
Plm Reserva Natural Palmari Atalaia do Norte, Amazonas Am 80 80
PNI Parque Nacional do Itatiaia Resende, Rio de Janeiro 9500 3000
RG Rancho Grande Cacaulandia, Rondonia 300 300
Rgu Reserva Guapi Assu Bird Lodge Cachoeiras de Macacu, Rio de Janeir 0 200
SBc Serra do Bocaina Sao Jose do Barreiro, Sao Paulo 1150 1200
SBr Southeast Brazil Ubatuba, Sao Paulo 0
Colombia CO
- -, o
Afl Altos Florencia Florencia, Caquetá
Ang Carretera Caldas - Angelópolis Angelópolis, Antioquia 1800 2000
Bvt Sendero Buena vista Guaviare, Guainía
Cal Caluma Caluma, Bolivar 1200
137
Colombia CO
CCri Caño Cristales La Macarena, Meta 250 260
CdC Cueva del Condor San Luis, Antioquia 400 540
CRC Cañón del Río Claro - Reserva el Refugio San Francisco, Antioquia 350 600
FCy Finca El Caney Puerto Berrío, Antioquia
Gai Piscicola el Gaitero Sopetrán, Antioquia
Glp Municipio de Galapa Galapa, Atlántico
GMq Golfo de Morrosquillo Tolú, Sucre 0 10
Gurc Quebrada La Guaracu San Jerónimo, Antioquia 650 680
Gvr Varias Guaviare San José del Guaviare, Guaviare
HFr Hacienda Farias Concordia, Antioquia 1000 1100
HLB Humedal La Bramadora Sopetrán, Antioquia
HNp Parque tematico Hacienda Napoles Puerto Triunfo, Antioquia 200 250
Jard Municipio Jardín - Fundación Colibrí Jardín, Antioquia 1500 3000
LPC La Piñuela - Reserva la Cristalina Cocorná, Antioquia 800 900
Mco Municipio Mocoa Mocoa, Putumayo
MMt Municipio de Mitú Mitú, Vaupés 180 200
Mnc Minca Santa Marta, Magdalena
Mor Reserva Natural Moru Cumaribo, Vichada 80 100
Mzm Carretera Montezuma - PNN Tatamá Pueblo Rico, Risaralda 1100 2600
Oqy Santuario de F y F Otún Quimbaya Pereira, Risaralda 1800 1800
PGt Santa Cecilia- Piscina Gito Pueblo Rico, Risaralda 400 400
Pña Peñalisa Tarso, Antioquia 600 700
PSJ Pueblito de San Jose Amagá, Antioquia 1800 1900
PYm Parcelacion El Yarumo La Ceja, Antioquia
QEV Quebrada El Viao Cocorná, Antioquia 1870 1980
QLH Quebrada La Habana San Luis, Antioquia 700 750
RAA Reserva natural de las aves Arrierito Antioqueño Anorí, Antioquia 1400 1850
(Piha)
RAD Reserva natural de las aves El Dorado Santa Marta, Magdalena 950 2600
Rch Finca La Rochela Santa Rosa de Osos, Antioquia 1000 1100
RlR Reserva La Romera Sabaneta, Antioquia 1800 2200
RlT Reserva Las Tangaras El Carmen de Atrato, Chocó 1492 1800
Rmo Rincones de Monticello Medellín, Antioquia 1800 1850
138
Colombia CO
RNM Reserva Natural Mururito Puerto Gaitán, Meta 130 180
RP Río Pepino Mocoa, Putumayo 550 580
Rpau El Paujil - Las Quinchas Puerto Boyacá, Boyacá
RPF Reserva Paz y Flora Guasca, Cundinamarca
RRm Reserva Rocamadre Toluviejo, Sucre 92 320
RSD Río Santodomingo Cocorná, Antioquia
RSm Río Samana San Luis, Antioquia 380 420
SFx San Felix - Las Antenas Bello, Antioquia 2500 3100
Sidr San Isidro - Alto de la Ceja La Ceja, Antioquia 2000 2200
Sin Sinifaná - La Holanda Titiribí, Antioquia 600 790
SJn Municipio de San Jeronimo San Jerónimo, Antioquia
SLz San Lorenzo Santa Marta, Magdalena
SM Santa María Santa María, Boyacá
SNc San Nicolas - Parcelación Arreboles Sopetrán, Antioquia
SSb San Sebastian - Alto del Escobero Retiro, Antioquia
Tft Vereda Tafetanes Sopetrán, Antioquia
Tkk Reserva Natural Takarkuna Capurganá - Sapzurro Acandí, Chocó 0 50
VCC Vereda Cristalina Chamuscados Fredonia, Antioquia 1550 1650
VPN Vereda Paso Nivel - Finca Zanzibar Amagá, Antioquia 1450 1500
Costa Rica CR
Lsel La Selva Biological Station Puerto Viejo de Sarapiqui, Heredia
Ecuador EC
AJL Arajuno Jungle Lodge Tena, Napo 350 350
Atrk Apuya Track Tena, Napo 550 600
Bañ Baños Baños, Tungurahua 1500 2800
Baz Baeza Baeza, Orellana 1800 1800
BBR Rio Bigal Biological Reserve Loreto, Orellana 900 1100
Caj Cajanuma Entrance, PN Podocarpus, EC (Refugio Loja, Loja 2800
Cajanuma)
Cpl Copalinga Lodge Zamora, Zamora-Chinchipe 950 950
JSR Jatun Sacha Reserve Misahualli, Napo 400 400
139
Ecuador EC
Loj Loja area, EC (Villonaco Ridge,top of old Loja road ) Loja, Loja 2400 2750
Mnd Mindo San Miguel de Los Bancos, Pichincha 1000 1400
Orq Las Orquideas Yantaza, Zamora-Chinchipe 900 900
PaP Papallacta Pass Papallacta, Pichincha 4100
PmpL Pimpilala Lodge Tena, Napo 600 600
Podc Bombuscaro Ranger Station, Parque Nacional Zamora, Zamora-Chinchipe 1300 1300
Podocarpus
Pulu Pululahua Crater Calicali, Pichincha 2850
Rho Rio Hollin, Reserva Narupa Jondache, Napo 1050 1050
Ten Area de Tena Tena, Napo 510 550
Vilc Vilcabamba Loja, Loja 1500
WS WildSumaco Lodge Loreto, Orellana 1350 1350
Yach Napo lowlands(Yachana, La Selva, Napo Wildlife Coca, Orellana
Center)
Estados Unidos de US
RGv Rio Grande Valley Hidalgo, Texas
México MX
Cat Catemaco Catemaco, Veracruz 0 1100
CNv Cañon Novillo Ciudad Victoria, Tamaulipas 460 460
Ctm Catemaco Catemaco, Veracruz 200 200
GF Reserva de la Biosfera El Cielo Gomez Farias, Tamaulipas 350 1400
HCcd Hotel Cascades Orizaba, Veracruz 1200 1200
HTn Hotel Taninul Ciudad Valles, San Luis Potosí 45 45
LHt La Huerta Cortazar, Guanajuato 1800 1800
Mue Mueller's Xalapa, Veracruz 500
Oax Oaxaca Oaxaca, Oaxaca 1700 1700
Ord Ocampo road Ocampo, San Luis Potosí 1150 1150
Plq Palenque Joaquin Miguel Gutierrez, Chiapas 70 70
Teo Teotitlan de Valle Teotitlan, Oaxaca 1700 1700
Panamá PN
140
Panamá PN
Boc Bocas Bocas, Bocas del Toro
Bqt Boquete Boquete, Chiriquí 1400 1400
CTwr Canopy Tower Gamboa, Panamá 200 200
Fort Fortuna Field Station Hornito, Chiriquí 1000 1400
PplRd Pipeline Road Gamboa, Panamá 200 200
Tot Mt. Totumas Cloud Forest Volcan, Chiriquí 1800 2000
Vant Valle Antón El Valle de Anton, Coclé
Perú PE
Aco Aconabikh Tarapoto, San Martín 750 750
AMZ Amazonia Lodge Atalaya, Madre de Dios 400 400
ARCC Amazon Research and Conservation Center (Lago Las Piedras, Madre de Dios 230 230
Soledad)
BSht Bosque Shollet Oxapampa, Pasco 1800 2250
CoR Cock of the Rock Lodge/Paradise Lodge, Manu Road San Pedro, Cusco 1350 1400
Cus Cusco area, PE (Abra Malaga,etc) Ollantayambo, Cusco 4200
Chtk Chontachaka, Manu Road Pillcopata, Cusco 850 850
Hual Huallaca Valley Tarapoto, Áncash
Jgu Juan Guerra Tarapoto, San Martín 350 350
Jul La Julianita Moyabamba, San Martín 950 950
Lim Lima Lima, Lima 5 50
Mrd San Pedro, Manu Road San Pedro, Cusco 1350 1400
MWC Manu Wildlife Center Rio Madre de Díos, Madre de Dios 270 270
Pcy Picuroyacu Punchana, Loreto 150 150
PH Pampa Hermosa Chanchamayo, Junín 1200 1300
PMp Playa Mariposa Moyabamba, San Martín 1000 1050
PNM Parque Nacional del Manu San Pedro, Cusco 300 4000
PntL Pantiacolla Lodge Rio Madre de Díos, Madre de Dios 400 400
PNYC Parque Nacional Yanachaga-Chemillen Pozuzo, Pasco 1000 1450
Pro La Promisora San Ramon, Junín 1050 1050
Qtz Quitacalzon, Lower Manu Road San Pedro, Cusco 1050 1100
Rctl Rocotal, Manu road Pillahuata, Cusco 2000 2200
Rmp- Rumipata-Mishquiyacu Moyabamba, San Martín 950 1000
141
Perú PE
Rmpt Rumipata-Mishquiyacu Moyobamba, San Martín 950 1000
Salb San Alberto Oxapampa, Pasco 1800 2300
TngM Tingo Maria area Tingo Maria, Huánuco 650 650
TRC Tambopata Research Center Tambopata, Madre de Dios 240 240
VCm Villa Carmen, Manu Road Pillcopata, Cusco 500 550
Wyq Wayquecha Research Station Pillahuata, Cusco 2900 2900
142
Fotógrafos - Photographers
Este documento es el esfuerzo acumulativo de muchas personas. Los siguientes son los fotógrafos
que han contribuido con sus fotos para generar la guía. Muchas gracias a todos.
This list is the result of the cumulative efforts of a large number of people. The following are the
photographers who have contributed their photos to this work, thank you very much.
AWr Adam Winer JOG John Ortega Galvan

AH Adrian Hoskins JAB Johnnier Arango Bermudez
AN Andrew Neild JJ Jorge J Restrepo
Bsu Barry Sullender Jpe Jorge Perez
BBc Bill Beck JFG Jose Francisco Garcia
BBt Bill Berthet JMz Jose Monzon
BBn Bill Bouton BB Juan Guillermo Jaramillo Velasquez
CRm Cristóbal Ríos-Málaver KNc Karen Nichols
DW Dan Wade KW Kay Wade
DJF Daniel Jaramillo Ferrer KK Ken Kertell
DPt Daniel Piedrahita Thiriez KP Kevin Painter
DvG David Geale KD Kim Davis
DGz David Guzman KG Kim Garwood
FM Fredy Montero KWs Kristine Wallstrom
GMr German Muriziasco LG Leif Gabrielsen
GC Gill Carter LBd Leonel Baldoni
GM Glenn Mahler Leo Leopoldina Tapasco
GS Gottfried Siebel LC Les Catchick
HB Hank Brodkin LuL Luc Legal
JB Jim Brock LEM Luis Eduardo Mejia
JHg Jim Hengeveld MCC Marcos Cesar Campis
JSn Jim Snyder MIW Maria Isabel Weyermans
JAR João Amarildo Ranguetti Mmi Maris Midgely
JSl Joe Schelling MZ Maristela Zamoner
JK John Kemner MM Martin Moreno
JMD John MacDonald MR Martin Reed
143
MC Mike Chapman
MS Mike Stangeland
MHN Natural History Museum London
OE Oscar Enciso
PS Pamela Sai
PM Pat Martin
PT Paul Thompson
PL Pingchung Lee
PB Priscilla Brodkin
RnM Renato Mattei
RC Rick Cech
RH Rich Hoyer
RL Richard Lehman
RW Rob Westerduijn
RGd Robert Gallardo
RSb Robert Straub
RRt Roger Rittmaster
RD Rudy Dodero
Smd Santiago Mejía Dugand
SFn Shawneen Finnegan
SN Sherry Nelson
SS Shirley Sekarajasingham
SDM Sidney Dantas de Melo
TGz Terry Gonzales
TS Tinio Sanchez
THn Tom Horton
TH Tony Hoare †
WC Will Carter
144
Agradecimientos Acknowledgments
Estas guías y listados de chequeo son el resultado del These guides and checklist are the result of the effort
esfuerzo de un gran número de colaboradores, of a large number of collaborators, scientists, fans
científicos, aficionados y fotógrafos a quienes damos and photographers to whom we thank, because
las gracias, pues sin ellos, estas publicaciones no without them, these publications would not have
hubieran sido posible de completar. been possible to complete.
En la familia Hesperiidae agradecemos In the Hesperiidae family we especially thank the
especialmente al experto Bernard Hermier por las expert Bernard Hermier for the countless hours and
innumerables horas y páginas de explicaciones y pages of explanations and corrections without which
correcciones sin las cuales habría sido imposible it would have been impossible to make this
hacer este documento. document.
Al Dr. Keith Willmott por sus consejos en la To Dr. Keith Willmott for his advice in preparing the
elaboración de la presentación de este trabajo e presentation of this work and identification of
identificación de fotos. photos.
Agradecemos a los siguientes expertos que We thank the following experts who have generously
generosamente han ayudado con información y en la helped with information and in the identification of
identificación de algunas fotos que han servido de some photos that have served as support for the list
soporte al listado (en orden alfabético): Stéphane (in alphabetical order): Stéphane Attal, Dr. Andrew
Attal, Dr. Andrew Brower, Prof. James Mallet, Brower, Prof. James Mallet, Andrew Neild, Gregory
Andrew Neild, Gregory Nielsen, Dr. Tomasz Pyrcz, Dr. Nielsen, Dr. Tomasz Pyrcz, Dr. Robert Robbins, Julián
Robert Robbins, Julián Salazar y Aaron Soh. Salazar and Aaron Soh.
A Andrés Arenas por sus consejos y ayuda en la To Andrés Arenas for his advice and help in preparing
elaboración de la base de datos. A Andrew Neild por the database. To Andrew Neild for his
sus recomendaciones y sugerencias. A David Geale recommendations and suggestions. To David Geale
por sus innumerables aportes fotográficos. for his countless photographic contributions.
A Gabriel Rodríguez por sus recomendaciones en To Gabriel Rodríguez for his recommendations on
cómo distribuir las zonas biogeográficas y por los how to distribute the biogeographic zones and for
múltiples datos geográficos de su colección de the multiple geographic data of his collection of
mariposas. Al Dr. Juan José Arango, Rodrigo Bernal y butterflies. To Dr. Juan José Arango, Rodrigo Bernal
Daniel Jaramillo F. por sus consejos y colaboración en and Daniel Jaramillo F. for their advice and
la consecución de datos. A Maristela Zamoner por la collaboration in obtaining data. To Maristela
traducción al portugués. Zamoner for the portuguese translation.
Al Museo de Historia Natural de Londres (Natural To the Natural History Museum of London for
History Museum) por permitirnos el acceso a sus allowing us access to their collections, specimens
colecciones, ejemplares y datos. A Blanca Huertas and data. To Blanca Huertas for all her advice and
por todos sus concejos y recomendaciones. recommendations.
Especial agradecimiento a las familias de los autores, Special thanks to the families of the authors, for their
por su apoyo durante la realización de este proyecto. support during the realization of this project.
145
Esta guía es un legado porque varios de los This guide is a legacy because several of the
sitios donde fueron registradas las especies ya sites where the species were recorded have
están destruidos y por lo tanto estas especies already been destroyed and therefore these
ya no existen en esa localidad específica. El species no longer exist in that specific locality.
listado y la guía ilustran la asombrosa The checklists and the guides illustrate the
diversidad de mariposas del Neotrópico para amazing diversity of butterflies of the
motivar su conocimiento y conservación. Neotropics to motivate their knowledge and
conservation.
“Solo podemos conservar “We can only conserve

y proteger lo que and protect what we
conocemos” know" JGJ

Neotropic Lycaenidae Butterfly Guide 02may2022

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Formatos disponibles

“Este listado está dedicado a todas las personas que nos han ayudado a construirlo y a

todos aquellos que están trabajando por el conocimiento y la conservación de la

Para citar este documento utilice: To cite this document use:

GARWOOD K., JARAMILLO J.G. Guía de Mariposas Lycaenidae del Neotrópico –

Laothus gibberosa Pseudolycaena marsyas

BioButterfly Database – Butterflycatalogs.com

1- Nombre científico 1- Scientific name

BioButterfly Database – Butterflycatalogs.com

BioButterfly Database – Butterflycatalogs.com

BioButterfly Database – Butterflycatalogs.com

BioButterfly Database – Butterflycatalogs.com

Subtribu - Subtribe Jantheclina

BioButterfly Database - Butterflycatalogs

BioButterfly Database - Butterflycatalogs

BioButterfly Database - Butterflycatalogs

Iophanus pyrrhias ♂ Iophanus pyrrhias ♂

Iophanus pyrrhias ♂ Iophanus pyrrhias ♂ Iophanus pyrrhias ♂

Iophanus pyrrhias ♂ Iophanus pyrrhias ♂ Iophanus pyrrhias ♀

BioButterfly DataBase - Butterflycatalogs.com Lycaeninae

AA infNEOTROPICO SUBFAMILIA Photo Guide

Busque versión nueva de este catálogo en: Butterflycatalogs.com

BioButterfly DataBase - Butterflycatalogs.com Lycaeninae Página 2 de 2

Brevianta bathoryon Brevianta busa

Brevianta celelata Brevianta ematheon Brevianta ematheon

Brevianta hyas Brevianta undulata Eumaeus childrenae

BioButterfly DataBase - Butterflycatalogs.com Theclinae

Eumaeus godartii Eumaeus godartii Eumaeus toxana

Eumaeus toxana Eumaeus toxea Micandra aegides ♂

Micandra aegides ♀ Micandra comae ♂ Micandra comae ♀

Micandra comae ♀ Micandra comae ♀ Micandra comae ♀

BioButterfly DataBase - Butterflycatalogs.com Theclinae Página 2 de 114

Micandra comae ♀ Micandra comae Micandra dignota

Micandra ion Micandra platyptera ♂ Micandra platyptera ♂

Micandra platyptera ♀ Micandra platyptera ♀ Micandra platyptera ♀

Micandra platyptera ♀ Micandra platyptera ♀ Micandra platyptera ♀

BioButterfly DataBase - Butterflycatalogs.com Theclinae Página 3 de 114

Micandra platyptera ♀ Micandra sp1 Mithras nautes ♂

Mithras nautes ♀ Mithras nautes Theorema dysmenia

Thestius epopea Thestius epopea Thestius epopea

Thestius meridionalis ♂ Thestius meridionalis ♂ Thestius meridionalis ♂

BioButterfly DataBase - Butterflycatalogs.com Theclinae Página 4 de 114

Thestius meridionalis ♂ Thestius meridionalis ♂ Thestius meridionalis ♀

Balintus tityrus Balintus tityrus

Balintus tityrus Johnsonita auda ♂ Johnsonita auda

CO-RlR, 1850m - CO-BO CO-, - PN-PE,VZ CO-, PN-PE,VZ

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Johnsonita auda Johnsonita auda Johnsonita auda

Johnsonita auda Johnsonita auda Johnsonita chlamydem ♂

Johnsonita chlamydem ♂ Johnsonita chlamydem ♀ Johnsonita chlamydem ♀

Johnsonita chlamydem Johnsonita chlamydem Johnsonita pardoa

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Johnsonita pardoa Johnsonita turquisca Lathecla latagus

Lathecla latagus Lathecla mimula Lathecla mimula

Lathecla mimula Lathecla mimula Rhamma adunca

Rhamma arria ♂ Rhamma arria Rhamma aurugo

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Rhamma bilix Rhamma bilix Rhamma commodus

Rhamma commodus Rhamma commodus Rhamma comstocki ♀

Rhamma familiaris Rhamma familiaris Rhamma hybla

Rhamma mishma Rhamma oxida Rhamma sp1

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Rhamma sp2 Rhamma sp2 Salazaria sala

Salazaria sala Salazaria sala Salazaria sp1

Busbiina bosora Marachina maraches