Peru Lycaenidae Butterfly Guide 09may2022 Butterflycatalogs

“Este listado está dedicado a todas las personas que nos han ayudado a construirlo y a
todos aquellos que están trabajando por el conocimiento y la conservación de la

naturaleza.”
“This list is dedicated to all the people who have helped us to build it and to all those who
are working for the knowledge and conservation of nature.”
Para citar este documento utilice: To cite this document use:
GARWOOD K., JARAMILLO J.G. Guía de mariposas Lycaenidae de Perú – Butterfly

guide to the Lycaenidae of Peru [en línea], BioButterfly Database. Disponible en:<
http://www.butterflycatalogs.com/ 130 pp. 05 de mayo de 2022
Diseño y diagramación Juan Guillermo Jaramillo V.
Annamaria ganimedes Symbiopsis perulera

Foto cubierta (Front cover): Foto cubierta inglés (English cover):
David Geale David Geale
Introducción Introduction
Con el propósito fundamental de hacer educación With the fundamental purpose of educating and
y crear conciencia sobre la importancia de las raising awareness about the importance of
mariposas en el ecosistema y su necesidad de butterflies in the ecosystem and the need to
conservarlas hemos recopilado información en la conserve them, we have compiled information in
base de datos BioButterfly. the BioButterfly database.
Con esta base de datos elaboramos catálogos y With this database we create butterfly catalogs
guías de mariposas, tanto para todo el Neotrópico and guides, both for the entire Neotropics and for
como para países y localidades desde México countries and localities from Mexico to Argentina.
hasta Argentina.
The guides contain photographs that have been
Las guías contienen fotografías que han compiled and taken by the authors and 366 other
recopilado y tomado los autores y otros 366 photographers who selflessly have given them up
fotógrafos quienes desinteresadamente nos las for our work. That is why at the end of this
han cedido para nuestro trabajo. Por ello al final document we have a complete list of all those
de este documento tenemos un listado completo who have contributed their photos. These
de todos aquellos que han contribuido con sus photographs represent thousands of hours of
fotos. Estas fotografías representan miles de fieldwork that would otherwise have been nearly
horas de trabajo de campo que de otra manera impossible to collect.
hubiera sido casi imposible de recopilar.
This information is constantly evolving and
Esta información está en constante evolución y changing, which is why we believe that the
cambio por eso creemos que las versiones electronic versions are the best option, because
electrónicas son la mejor opción, pues al poder by being able to generate lists and guides without
generar listados y guías sin costo de impresión printing costs whenever you want, it can be the
cada que se quiera puede ser el mecanismo más most practical mechanism for collecting and
práctico para recopilar y compartir la información sharing information about neotropical butterflies.
sobre las mariposas neotropicales.
For the presentation and layout of the document
Para la presentación y diagramación del we chose a format between scientific and didactic
documento escogimos un formato entre científico because, although the information is very
y didáctico pues, aunque la información es muy scientific, the use of photos helps to attract the
científica, el uso de fotos ayuda a atraer público public, and this is part of the objective: to
general y este es parte del objetivo: comunicar communicate to educate and conserve.
para educar y conservar.
At the end we add a numbered list with the
Al final agregamos una lista numerada con las species of the family in the country (without
especies de la familia en el país (sin subespecies) subspecies) and the numbered genera. Those
y también los géneros numerados. Quien desee who want more information should refer to the
más información se debe remitir al listado complete list of the country or the list of the
completo del país o a la lista del Neotrópico. Neotropics.
Esperamos que esta información sea de utilidad al We hope that this information will be useful in
aumentar el conocimiento sobre las mariposas y increasing knowledge about butterflies and that it
que ayude a fomentar la observación, estudio, will help promote the observation, study,
fotografía y disfrute de estas y que la información photography, and enjoyment of these, and that
pueda ser utilizada por gobiernos, instituciones o the information can be used by governments,
personas para tomar decisiones y crear conciencia institutions, or individuals to make decisions and
sobre la necesidad de su estudio y conservación. raise awareness about the need for its study and
conservation
1
Nota importante Note
Estas guías y listados de chequeo son el These guides and checklists are the result of
resultado de más de veinte años de trabajo de more than twenty years of work by the authors.
los autores. Para llegar al resultado que Usted To reach the result you have in your hands, we
tiene en sus manos hemos contado con la have had the generous help of many expert
generosa ayuda de muchos científicos expertos scientists and many photographers.
y de un gran número de fotógrafos. These publications are the product of the
Estas publicaciones son producto de la compilation of bibliography and photographic
recopilación de bibliografía y registros records from many locations in Neotropical
fotográficos de muchas localidades en países del countries. The user must know that identifying
Neotrópico. El usuario debe saber que some species from only photographs has an
identificar algunas especies a partir solamente implicit margin of error, so it is possible that the
de fotografías tiene implícito un margen de guides have some incorrect identifications or
error, por lo que es posible que las guías tengan lack the precision that we would like for all of
algunas identificaciones incorrectas, o carentes them, despite our efforts and the knowledge
de la precisión que desearíamos para todas, a that the consulted experts have.
pesar de nuestro esfuerzo y del conocimiento To prepare the guides we use photos of
que los expertos consultados tienen. butterflies taken in different countries and from
Para elaborar las guías usamos fotos de these we select the photo or photos that best
mariposas tomadas en diferentes países y de represent the species, therefore, we use that
estas seleccionamos la foto o fotos que mejor photo or photos for all the family guides or for
representen a la especie, por lo tanto, esa foto the different countries. It is for this reason that
o fotos las usamos para todas las guías de familia it is possible, for example, to find a butterfly
o para los diferentes países. Es por esta razón photographed in Peru in a guide to Ecuador. We
que es posible, por ejemplo, encontrar en una are certain that this butterfly has been
guía del Ecuador una mariposa fotografiada en registered in Ecuador, but we use the photo that
el Perú. Nosotros tenemos la certeza de que best represents the species, even if it was taken
dicha mariposa ha sido registrada en Ecuador, in another country, because the idea is that the
pero usamos la foto que mejor represente la user has the best possible idea of what the
especie, así haya sido tomada en otro país, butterfly looks like.
porque la idea es que el usuario tenga la mejor The publication, being generated electronically,
idea posible de cómo luce la mariposa. is easy to correct, does not imply waste of paper
La publicación, al ser generada and can be permanently updated. If you find any
electrónicamente, es fácil de corregir, no error or inconsistency, we have a methodology
implica desperdicio de papel y se puede to correct it: please call Juan Guillermo Jaramillo
actualizar permanentemente. Si encuentra V. at +57-310 830 6732 or write to
algún error o inconsistencia, tenemos una [email protected] or Kim Garwood
metodología para corregirla: por favor llame a [email protected].
Juan Guillermo Jaramillo V. al celular +57-310
830 6732 o escriba al correo
[email protected] o a Kim Garwood
[email protected].
BioButterfly Database – Butterflycatalogs.com
2
Convenciones Keys
Para cada foto de una especie se trata de dar la For each photo of a species, it is about giving as
mayor cantidad de información posible en el much information as possible in the smallest
menor espacio, los números en azul significan: space, the numbers in blue mean:
1- Nombre científico 1- Scientific name

2- Nombre anterior 2- Previous name
3- Sexo 3- Sex
4- Fotógrafo 4- Photographer
5- Información de la foto 5- Photo information
6- Rango geográfico 6- Geographic range
7- Secuencia de la foto 7- Photo sequence
3
1 Nombre científico 1 Scientific name
El nombre científico con género, especie y The scientific name with genus, species and
subespecie cuando sea posible. subspecies when possible.
Los nombres científicos con el género y sp The scientific names with the genus and sp
corresponden a especies que tienen una correspond to species that have an unknown or
identificación desconocida o dudosa, ya que doubtful identification, since there may be
puede haber varias especies con dudas será sp1, several species with doubts, it will be sp1, sp2,
sp2, etc., una vez que se identifique etc., once it is positively identified, the sp will be
positivamente, la sp se cambiará al nombre changed to the correct name of the species.
correcto de la especie. In the case of species not described to science,
En el caso de especies no descritas para la which do not yet have a name, we put sp nov.
ciencia, que todavía no tienen nombre, Please, if you can help us to complete or correct,
colocamos sp nov. Por favor, si puede ayudarnos write to the contacts.
a completar o corregir, escríbanos a los In many places there are species without
contactos. positive identification, we prefer to post photos
En muchos lugares hay especies sin of these species so that it is known that this
identificación positiva, preferimos colocar fotos species is in that place and so that in the future
de estas especies para que se sepa que esta someone can describe it. In case we have doubts
especie está en ese lugar y para que en el futuro even with the gender, we put the gender that
alguien pueda describirla. En caso de que we consider closest and two question marks like
tengamos dudas incluso con el género, this: ??.
ponemos el género que consideramos más In some cases of species found in taxonomic
cercano y dos signos de interrogación como studies, we place the names as follows: sp
este: ??. [group]
En algunos casos de especies que se encuentran The taxonomy and sequence of this catalog
en estudios taxonómicos colocamos mientras se follows the authors that are specified in our
definen los nombres así: sp [grupo] publication: Neotropical Butterflies Checklist
La taxonomía y la secuencia de este catálogo South America.
sigue los autores que están especificados en
2 Former name
nuestra publicación: lista Mariposas del
Neotrópico Lista de Chequeo Suramérica. With the taxonomy recently revised with DNA,
several species changed the genus or name, in
2 Nombre anterior those cases we will put the old genus or name
Con la taxonomía revisada recientemente con el with the red text.
ADN, varias especies cambiaron el género o el
3 Sex
nombre, en esos casos colocaremos el antiguo
género o nombre con el texto en rojo. Whenever possible, we put the sex of the
specimen in the photo, either because they
3 Sexo have sexual dimorphism or because in some
Cuando sea posible, ponemos el sexo del cases, we observed the female laying eggs.
espécimen de la foto, ya sea porque tienen
4
dimorfismo sexual o porque en algunos casos 4 Photographer
observamos a la hembra poniendo huevos.
The abbreviation of the name of the
4 Fotógrafo photographer who took the photo
La abreviación del nombre del fotógrafo que acknowledges their collaboration, please inform
tomó la foto reconoce su colaboración, por us of any errors for correction.
favor nos informan de cualquier error para su 5 Photo information
corrección.
Whenever possible, the photo information will
5 Información de la foto contain the abbreviation of the country, the
Cuando sea posible, la información de la foto locality, the altitude, and the month/day the
tendrá la abreviación del país, la localidad, la photo was taken.
altitud y el mes/día en que se tomó la foto. Most of the photos were taken of live butterflies
La mayoría de las fotos fueron tomadas de in their natural environment. The country where
mariposas vivas en su entorno natural. El país it was taken is indicated by a two-letter
donde fue tomado se indica con una abreviatura abbreviation. This abbreviation corresponds to
de dos letras. Esta abreviatura corresponde al the international code used as the Internet
código internacional utilizado como la termination of the code of each country, for
terminación de Internet del código de cada país, example. Brazil BR, Ecuador EC, Peru PE etc.
por ejemplo. Brasil BR, Ecuador EC, Perú PE etc. After the country code, we put an abbreviation
Después del código del país, ponemos una of the locality (see the table of equivalent codes
abreviatura de la localidad (consulte la tabla de for localities), then the altitude where the photo
códigos equivalentes para las localidades), luego was taken in meters above sea level and the
la altitud donde se tomó la foto en metros sobre month and day, to give an idea of the date they
el nivel del mar y el mes y día, para dar una idea were flying at this location.
de la fecha que estuvo volando en este lugar. From each species or subspecies, we choose the
De cada especie o subespecie escogemos las best photos that represent it both ventral-dorsal
mejores fotos que la representen tanto ventral- and male-female and these photos are used in
dorsal como macho-hembra y estas fotos se the guides for all countries. There are many
usan en las guías para todos los países. Hay species that are common to almost all countries
muchas especies que son comunes a casi todos and that do not differ significantly in their shape
los países y que no difieren significativamente or color between countries, because trying to
en su forma o color entre los países, porque make the guide of a country or locality using
tratar de que la guía de un país o localidad solo only photos taken in that country or locality
tenga fotos tomadas en ese país o localidad se becomes a titanic project knowing that when we
convierte en un proyecto titánico, sabiendo que have more than 14,000 taxa for the Neotropics.
tenemos más de 14.000 taxones para el 6 Range
Neotrópico.
The distribution range of the species with an
6 Rango abbreviation of the name of the countries from
El rango de distribución de las especies con una north to south, in some cases with species that
abreviatura del nombre de los países de norte a inhabit the United States and Mexico, we use
sur, en algunos casos con especies que habitan the code of the southernmost states where the
en los Estados Unidos y México, utilizamos el species are Tx Texas, Chi Chiapas. In big
5
código de los estados más australes donde están countries like Brazil, the upper part of the
las especies Tx Texas, Chi Chiapas. En países Amazon, the lower part, etc. Colombia and
grandes como Brasil, la parte superior del other countries may have a position relative to
Amazonas, la parte inferior, etc. Colombia y the Andes Mountain range and this information
otros países pueden tener una posición relativa is placed in parentheses after the country
a la cordillera de los Andes y esta información se abbreviation.
coloca entre paréntesis después de la
7 Photo Sequence
abreviación del país.
In the upper right corner, there is a consecutive
7 Secuencia de la foto number that helps to quickly locate or refer to
En la esquina superior derecha, hay un número the position of a species in the guide.
consecutivo que ayuda a localizar o hacer
referencia rápidamente a la posición de una
Specimen photos are used to show some
especie en la guía.
features that are not possible to photograph in
the wild or to show species for which we do not
Las fotos de especímenes se usan para mostrar yet have live specimens. In addition, in some
algunas características que no son posibles de cases dorsal photos of captured species are
fotografiar en la naturaleza o para mostrar taken to better identify them, these butterflies
especies de las cuales aún no tenemos are released immediately after the photo.
especímenes vivos. Además, en algunos casos se Photos of specimens or photos of low quality
toman fotos dorsales de especies capturadas will be replaced once better photos of these
para identificarlas mejor, estas mariposas se species are taken, any collaboration that allows
liberan inmediatamente después de la foto. Las to complete the guide will be welcome and the
fotos de especímenes o fotos de baja calidad photographer will have the credits.
serán reemplazadas una vez que se tomen
mejores fotos de estas especies, cualquier
colaboración que permita completar la guía será
bienvenida y el fotógrafo tendrá los créditos.
6
Clasificación familia Lycaenidae
En este listado colocamos todos los géneros en el Neotrópico de la familia agrupados por subfamilia, de tal
manera que se pueda saber la ubicacion de un genero. Todos los generos no necesariamente estan en un pais
o localidad.
We have listed below all the Neotropic genera in the family within each subfamily, to help with locating a
particular genus. All genus are not necessarily in one country or locality.
Lycaenidae
11 Podanotum
12 Rhamma
Subfamilia - Subfamily
13 Salazaria
Lycaeninae Subtribu - Subtribe
Tribu - Tribe
Timaetina
Lycaenini 14 Busbiina
15 Marachina
Subtribu - Subtribe
Sin Subtribu
16 Nesiostrymon
17 Paraspiculatus
1 Iophanus
18 Penaincisalia
19 Phothecla
Subfamilia - Subfamily 20 Timaeta
Theclinae Subtribu - Subtribe
Atlidina
Tribu - Tribe
21 Arcas
Eumaeini
22 Atlides
Subtribu - Subtribe
23 Brangas
Eumaeina
24 Denivia
2 Brevianta
25 Pseudolycaena
3 Eumaeus
26 Theritas
4 Micandra Subtribu - Subtribe
5 Mithras Evenina
6 Theorema 27 Evenus
7 Thestius Subtribu - Subtribe
Subtribu - Subtribe Jantheclina

Rhammina 28 Allosmaitia
8 Balintus 29 Aveexcrenota
9 Johnsonita 30 Contrafacia
10 Lathecla
BioButterfly Database - Butterflycatalogs
7
Lycaenidae
31 Enos 56 Camissecla
32 Janthecla 57 Electrostrymon
33 Laothus 58 Gigantorubra
Subtribu - Subtribe 59 Kisutam
Paiwarriina 60 Lamprospilus
34 Annamaria 61 Mercedes
35 Fasslantonius 62 Pendantus
36 Kolana 63 Rubroserrata
37 Paiwarria 64 Serratofalca
Subtribu - Subtribe
65 Ziegleria
Cupatheclina
Subtribu - Subtribe
38 Cupathecla Strymonina
39 Dabreras 66 Arawacus
Subtribu - Subtribe
67 Heterosmaitia
Parrhasiina
68 Hypostrymon
40 Apuecla
69 Panthiades
41 Beatheclus
70 Rekoa
42 Caerofethra
71 Strymon
43 Dicya
72 Thereus
44 Ignata
Subtribu - Subtribe
45 Michaelus Strephonotina
46 Oenomaus 73 Aubergina
47 Olynthus 74 Celmia
48 Parrhasius 75 Crimsinota
49 Symbiopsis 76 Decussata
50 Thepytus 77 Gargina
Subtribu - Subtribe
78 Gossenia
Ipideclina
79 Iaspis
51 Ipidecla
Subtribu - Subtribe
80 Ministrymon
Calycopidina 81 Nicolaea
52 Argentostriatus 82 Ostrinotes
53 Arzecla 83 Rindgea
54 Badecla 84 Siderus
55 Calycopis 85 Strephonota
8
Lycaenidae
86 Terenthina 108 Echinargus
87 Theclopsis 109 Eldoradina
88 Tmolus 110 Elkalyce
Subtribu - Subtribe 111 Hemiargus
Trichonidina 112 Itylos
89 Bistonina 113 Leptotes
90 Manticia 114 Nabokovia
91 Megathecla 115 Paralycaeides
92 Trichonis 116 Pseudochrysops
Subtribu - Subtribe
117 Pseudolucia
Callophryidina
118 Zizula
93 Callophrys
94 Cyanophrys
95 Chalybs
96 Chlorostrymon
97 Erora
98 Magnastigma
99 Ocaria
100 Radissima
101 Satyrium
102 Semonina
103 Thaeides
Subfamilia - Subfamily
Polyommatinae
Tribu - Tribe
Polyommatini
Subtribu - Subtribe
Sin Subtribu
104 Brephidium
105 Celastrina
106 Cupido
107 Cyclargus
9
10
Lycaenidae
Subfamilia Theclinae
Tribu - Tribe
Eumaeini
1 2
Subtribu - Subtribe
Eumaeina
Brevianta ematheon Brevianta ematheon

DvG KK
PE-ARCC, VZ-PE,BR,SU EC-Atrk, VZ-PE,BR,SU
3 4 5
Brevianta undulata Eumaeus toxana Eumaeus toxana

DvG DW AN
PE-Qtz, VZ-PE PE-, VZ-BO,BR EC-Atrk, VZ-BO,BR
6 7 8
Micandra aegides ♂ Micandra aegides ♀ Micandra comae ♂

KG DvG DvG
EC-Cal, VZ-PE EC-Bañ, VZ-PE EC-Cal, CO-PE
BioButterfly DataBase - Butterflycatalogs.com Perú Theclinae

11
Tribu - Tribe
Eumaeini
9 10 11
Micandra comae ♀ Micandra comae ♀ Micandra comae ♀

DvG DvG DvG
EC-Caj, CO-PE EC-Orq, CO-PE EC-Cal, CO-PE
12 13 14
Micandra comae ♀ Micandra comae ♀ Micandra comae

JAB KK DvG
CO-Oqy, 1850m CO-PE CO-RlT, CO-PE CO-Mzm, 2600m CO-PE
15 16 17
Micandra dignota Micandra ion Micandra platyptera ♂

DvG DvG DvG
PE-Wyq, CO-BO PE-Rctl, CO-PE PE-PNYC, CR-BO,GF
18 19 20
Micandra platyptera ♂ Micandra platyptera ♀ Micandra platyptera ♀

KG AN
CO-SM, CR-BO,GF EC-WS, CR-BO,GF CO-PSJ, 1800m Nov21 CR-BO,GF
BioButterfly DataBase - Butterflycatalogs.com Perú Theclinae Página 2 de 87

12
Tribu - Tribe
Eumaeini
21 22 23
Micandra platyptera ♀ Micandra platyptera ♀ Micandra platyptera ♀
CO-PSJ, 1850m Oct17 CR-BO,GF CO-QEV, 1975m CR-BO,GF CO-QEV, 1975m CR-BO,GF
24 25 26
Micandra platyptera ♀ Micandra platyptera ♀ Mithras nautes ♂

BBt DvG
EC-, CR-BO,GF CO-QEV, 1975m CR-BO,GF PE-ARCC, VZ-PE,BR
27 28 29
Mithras nautes ♀ Mithras nautes Theorema dysmenia

DvG RH FM
PE-ARCC, VZ-PE,BR BR-CJL, VZ-PE,BR CO-, CO-PE
30 31 32
Thestius epopea Thestius epopea Thestius epopea

DvG DvG DvG
CO-Mzm, 1350m NI-PE CO-Mzm, 1350m NI-PE CO-Mzm, 1350m NI-PE

13
Tribu - Tribe
Eumaeini
33 34 35
Thestius meridionalis ♂ Thestius meridionalis ♂ Thestius meridionalis ♂

DvG DvG FM
EC-, 550m PN-BO,BR CO-MMt, 200m PN-BO,BR CO-RP, 560m Ago23 PN-BO,BR
36 37 38
Thestius meridionalis ♂ Thestius meridionalis ♂ Thestius meridionalis ♀

DvG KNc DvG
CO-SM, 750m PN-BO,BR PE-PNM, PN-BO,BR EC-AJL, 400m PN-BO,BR
39
Thestius meridionalis ♀
SS
CO-MMt, 200m Nov22 PN-BO,BR
40 41
Subtribu - Subtribe
Rhammina
Balintus tityrus Balintus tityrus

DvG
EC-WS, CO-BO CO-RlR, 1850m - CO-BO

14
Tribu - Tribe
Eumaeini
42 43 44
Balintus tityrus Johnsonita auda ♂ Johnsonita auda

DvG
EC-WS, CO-BO CO-, - PN-PE,VZ CO-, PN-PE,VZ
45 46 47
Johnsonita auda Johnsonita auda Johnsonita auda

SN
CO-Oqy, 1850m Sep26 PN-PE,VZ CO-SFx, 2600m May18 PN-PE,VZ CO-QEV, 1940m PN-PE,VZ
48 49 50
Johnsonita auda Johnsonita auda Johnsonita chlamydem ♂

KG DvG DvG
PE-Mrd, PN-PE,VZ PE-Rctl, PN-PE,VZ PE-Salb, May31 PE-BO
51 52 53
Johnsonita chlamydem ♂ Johnsonita chlamydem ♀ Johnsonita chlamydem ♀

DvG DvG DvG
PE-Salb, May31 PE-BO PE-Pro, Jun03 PE-BO PE-PMp, Sep28 PE-BO

15
Tribu - Tribe
Eumaeini
54 55 56
Johnsonita chlamydem Johnsonita chlamydem Johnsonita pardoa

KK THn THn
PE-Pro, Jun05 PE-BO PE-, PE-BO CO-Jard, VZ-BO
57 58 59
Johnsonita pardoa Lathecla latagus Lathecla latagus

AN DvG DvG
PE-, VZ-BO CO-Mzm, MX-BO,GF,BR? PE-Mrd, MX-BO,GF,BR?
60 61 62
Lathecla mimula Lathecla mimula Lathecla mimula

DvG DvG AN
CO-Mzm, 1300m CO?EC-PE,BR CO-Mzm, 1300m CO?EC-PE,BR PE-, CO?EC-PE,BR
63 64 65
Lathecla mimula Rhamma arria ♂ Rhamma arria

DvG FM AH
PE-MWC, CO?EC-PE,BR CO-RPF, 3000m VZ-BO,AR CO-Mzm, VZ-BO,AR

16
Tribu - Tribe
Eumaeini
66 67 68
Rhamma aurugo Rhamma commodus Rhamma commodus

DvG
PE-Cus, PE CO-PSJ, 1800m Dic19 VZ-BO CO-PSJ, 1800m Dic19 VZ-BO
69 70 71
Rhamma commodus Rhamma comstocki ♀ Rhamma comstocki

FM FM
CO-PSJ, 1800m Dic19 VZ-BO CO-RPF, 3000m CO-PE CO-, May06 CO-PE
72 73 74
Rhamma familiaris Rhamma familiaris Rhamma hybla

Paralustrus DvG Paralustrus KG Shapiroana DvG
EC-Caj, CO-BO EC-Sidr, CO-BO EC-Caj, CO-PE
75 76 77
Rhamma sp2 Rhamma sp2 Salazaria sala

DvG DvG PB
PE-, PE-, CO-Oqy, 1850m Jun04 CO-PE

17
Tribu - Tribe
Eumaeini
78 79
Salazaria sala Salazaria sala

DvG THn
CO-Mzm, May28 CO-PE EC-, Nov12 CO-PE
80 81
Subtribu - Subtribe
Timaetina
Marachina maraches Marachina maraches

DvG DvG
PE-Rctl, CO-BO CO-Mzm, 1700m CO-BO
82 83 84
Nesiostrymon calchinia Nesiostrymon calchinia Nesiostrymon celona

KG DvG OE
CO-RlT, 1700m Sep22 MX-AR CO-Mzm, 1700m MX-AR CO-, MX-BR
85 86 87
Nesiostrymon celona Nesiostrymon celona Paraspiculatus azul

DvG DvG THn
PE-PNYC, MX-BR PE-Rmp-Mq, MX-BR EC-, E EC-PE

18
Tribu - Tribe
Eumaeini
88 89 90
Paraspiculatus colombiensis Paraspiculatus elis Paraspiculatus elis

Mithras DvG Mithras AN Mithras DvG
CO-Mzm, 1800m CO-BO,BR,GF EC-Ten, SE CO,E EC-PE,BR,SU EC-PmpL, SE CO,E EC-PE,BR,SU
91 92 93
Paraspiculatus oroanna Paraspiculatus orobia ♂ Paraspiculatus orobia

DvG Mithras BBt Mithras DvG
PE-VCm, E EC-PE CO-CRC, 320m Jun20 CO-BO,BR,AR EC-Orq, CO-BO,BR,AR
94 95 96
Paraspiculatus orobia Paraspiculatus orobia Paraspiculatus orobia

Mithras AN Mithras TH Mithras DvG
PE-, CO-BO,BR,AR EC-Atrk, CO-BO,BR,AR PE-MWC, CO-BO,BR,AR
97 98 99
Paraspiculatus orobia Paraspiculatus orocana Paraspiculatus orocana

Mithras DvG Mithras DJF Mithras DJF
PE-Plm, CO-BO,BR,AR CO-, 1300m Jul16 CO-PE CO-, 1300m Jul16 CO-PE

19
Tribu - Tribe
Eumaeini
100 101 102
Penaincisalia amatista Penaincisalia atymna Penaincisalia culminicola

Abloxurina DvG DvG KG
PE-Wyq, EC-PE EC-Loj, EC-BO EC-Pulu, PE-BO
103 104 105
Penaincisalia culminicola Penaincisalia ianusi Penaincisalia loxurina ♂

DvG Ianusanta Mmi Thecloxurina
PE-Cus, PE-BO EC-Baz, EC-PE CO-SSb, 2500m CO-BO,VZ,AR
106 107 108
Penaincisalia loxurina Penaincisalia loxurina Penaincisalia loxurina

Thecloxurina DvG Thecloxurina BBt Thecloxurina
PE-Wyq, CO-BO,VZ,AR CO-Jard, CO-BO,VZ,AR CO-SFx, 2600m May18 CO-BO,VZ,AR
109 110 111
Penaincisalia loxurina Penaincisalia loxurina Penaincisalia loxurina

Thecloxurina Thecloxurina Thecloxurina DvG
CO-SFx, 2800m May16 CO-BO,VZ,AR CO-SFx, 2800m May16 CO-BO,VZ,AR CO-Mzm, 2600m CO-BO,VZ,AR

20
Tribu - Tribe
Eumaeini
112 113 114
Penaincisalia loxurina Penaincisalia loxurina Penaincisalia purpurea

Thecloxurina Thecloxurina Pons DvG
CO-SFx, 2800m May16 CO-BO,VZ,AR CO-SFx, 2800m May16 CO-BO,VZ,AR PE-Wyq, EC-PE
115 116 117
Penaincisalia purpurea Phothecla photismos Timaeta gabriela ♀

Pons KK DvG DvG
PE-Wyq, EC-PE PE-Mrd, VZ-PE EC-WS, EC-BO
118 119 120
Timaeta sp n Timaeta tema Timaeta tema

Temecla DvG Temecla DvG Temecla DvG
PE-Yab, Oct26 PE PE-PntL, CO-BO,BR PE-ARCC, CO-BO,BR
121 122 123
Timaeta tema Timaeta timaeus ♂ Timaeta timaeus ♂

Temecla KK
BO-Vtu, CO-BO,BR CO-, - CO-BO CO-PSJ, 1800m CO-BO

21
Tribu - Tribe
Eumaeini
124 125 126
Timaeta timaeus ♂ Timaeta timaeus ♂ Timaeta timaeus ♂

DvG
CO-, CO-BO CO-, - CO-BO PE-Rctl, CO-BO
127 128 129
Timaeta timaeus ♀ Timaeta timaeus ♀ Timaeta timaeus ♀

DvG
CO-RlR, 1850m - CO-BO CO-, - CO-BO PE-CoR, CO-BO
130
Timaeta trochus ♀
KK
PE-CoR, CO-PE
131 132
Subtribu - Subtribe
Atlidina
Arcas imperialis ♂ Arcas imperialis ♂

DvG DvG
PE-ARCC, MX-AR PE-ARCC, MX-AR

22
Tribu - Tribe
Eumaeini
133 134 135
Arcas imperialis ♂ Arcas imperialis ♀ Arcas tuneta

DvG THn
PE-Qtz, MX-AR CO-, - MX-AR EC-, CO-BO,BR,Guianas
136 137 138
Arcas tuneta tuneta Atlides atys ♂ Atlides havila

DvG GS
CO-Rmo, 1830m Feb26 CO-BO,BR PE-ARCC, PN-BO,BR BO-Cor, CO-BO
139 140 141
Atlides inachus Atlides inachus Atlides inachus

AN Leo AN
PE-, Abr19 MX-BR CO-Mzm, MX-BR PE-PNM, MX-BR
142 143 144
Atlides polybe ♂ Atlides polybe ♂ Atlides polybe ♂

KD DW DvG
MX-HTn, Nov23 MX-AR PN-Bqt, May03 MX-AR CO-Mzm, Sep26 MX-AR

23
Tribu - Tribe
Eumaeini
145 146 147
Atlides rustan ♂ Atlides rustan ♀ Brangas coccineifrons

FM DvG DvG
CO-, May31 MX-BO,BR,AR PE-Jgu, MX-BO,BR,AR PE-ARCC, MX-PE
148 149 150
Brangas felderi Brangas getus Brangas getus

CRm DvG DvG
CO-, 1750m Ago01 CO-PE PE-ARCC, MX-BO,BR,AR CO-SM, 750m MX-BO,BR,AR
151 152 153
Brangas getus Brangas insolitus Brangas neora ♀

DvG DvG
PE-ARCC, MX-BO,BR,AR PE-Rctl, EC-PE CO-GMq, 5m MX-BR
154 155 156
Brangas neora ♀ Brangas polonus Denivia arene

GS DvG
CO-GMq, 5m MX-BR BO-Crv, PE-BO PE-Mrd, EC-PE

24
Tribu - Tribe
Eumaeini
157 158 159
Denivia danaus Denivia danaus Denivia hemon ♂

Theritas/Lucilda DvG Theritas/Lucilda DvG Theritas Mnz
CO-Mzm, 1400m VZ-PE CO-Mzm, 1400m VZ-PE EC-, MX-BO,BR,AR
160 161 162
Denivia hemon ♂ Denivia hemon ♀ Denivia hemon ♀

Theritas Cdr Theritas Theritas GC
EC-, MX-BO,BR,AR CO-HNp, 200m Ene29 MX-BO,BR,AR PA-, MX-BO,BR,AR
163 164 165
Denivia hemon ♀ Denivia lisus ♂ Denivia lisus ♂

Theritas BBt Theritas KG Theritas DvG
CO-CRC, 320m Jun21 MX-BO,BR,AR PE-CoR, MX-BO,BR,AR PE-ARCC, MX-BO,BR,AR
166 167 168
Denivia lisus ♂ Denivia lisus ♀ Denivia lisus ♀

Theritas DvG Theritas DvG Theritas DvG
PE-ARCC, MX-BO,BR,AR PE-Qtz, MX-BO,BR,AR PE-Qtz, MX-BO,BR,AR

25
Tribu - Tribe
Eumaeini
169 170 171
Denivia monica Denivia monica Denivia monica

KG DvG DvG
PE-Qtz, VZ-BO PE-Qtz, VZ-BO EC-Qtz, VZ-BO
172 173 174
Denivia phegeus ♂ Denivia phegeus ♀ Denivia phegeus ♀

Theritas AN Theritas AN Theritas DvG
PE-, CO-BO,VZ,BR PE-, CO-BO,VZ,BR PE-ARCC, CO-BO,VZ,BR
175 176 177
Denivia phegeus Denivia phegeus Denivia viresco ♂

Theritas DvG Theritas DvG Theritas DvG
CO-Bvt, 200m Ene31 CO-BO,VZ,BR PE-Qtz, CO-BO,VZ,BR PE-ARCC, PE-BO,BR(Ma),GF
178 179 180
Denivia viresco ♂ Denivia viresco ♀ Pseudolycaena marsyas ♂

Theritas DvG Theritas DvG KK
PE-ARCC, PE-BO,BR(Ma),GF PE-ARCC, PE-BO,BR(Ma),GF CO-MMt, 200m Oct20 NI?-AR

26
Tribu - Tribe
Eumaeini
181 182 183
Pseudolycaena marsyas ♂ Pseudolycaena marsyas ♂ Pseudolycaena marsyas ♂

SS
CO-Tft, 574m NI?-AR EC-Ten, NI?-AR CO-, - NI?-AR
184 185 186
Pseudolycaena marsyas ♀ Pseudolycaena marsyas ♀ Pseudolycaena marsyas

FM DvG
CO-RP, 560m Nov07 NI?-AR CO-SNc, 474m NI?-AR BO-Volc, NI?-AR
187 188 189
Pseudolycaena marsyas Pseudolycaena marsyas Pseudolycaena marsyas

KK KK
PE-Jgu, NI?-AR PE-Jgu, NI?-AR CO-Sin, 780m Jun14 NI?-AR
190 191 192
Pseudolycaena marsyas Theritas harrietta Theritas harrietta

DvG DvG
CO-Sin, 700m Jul05 NI?-AR PE-Rctl, PE-BO PE-Qtz, PE-BO

27
Tribu - Tribe
Eumaeini
193 194 195
Theritas mavors ♂ Theritas mavors ♂ Theritas mavors ♂

FM FM FM
CO-RP, 560m Feb09 MX-BO,BR,GY CO-, May31 MX-BO,BR,GY CO-RP, 560m Jul25 MX-BO,BR,GY
196 197 198
Theritas mavors ♀ Theritas paupera Theritas paupera

DvG
CO-, MX-BO,BR,GY CO-Mzm, 1800m CO-PE,VZ CO-PSJ, 1800m Abr26 CO-PE,VZ
199
Theritas paupera
CO-SJn, 800m Sep12 CO-PE,VZ
200 201
Subtribu - Subtribe
Evenina
Evenus batesii Evenus batesii

FM DvG
CO-RP, 560m Ago23 MX-BR,AR EC-PmpL, MX-BR,AR

28
Tribu - Tribe
Eumaeini
202 203 204
Evenus coronata Evenus felix Evenus floralia

CRm AN AN
CO-, 2300m Oct02 MX-PE EC-Ten, VZ-PE EC-Atrk, EC-BO,BR,GF
205 206 207
Evenus gabriela Evenus gabriela Evenus regalis

DvG DvG JJ
EC-Atrk, CO-PE,BR,GF EC-Atrk, CO-PE,BR,GF CO-, May06 MX-BR,AR
208 209 210
Evenus regalis Evenus regalis Evenus regalis

MCC KK JJ
BR-, Abr02 MX-BR,AR EC-Atrk, MX-BR,AR CO-, May06 MX-BR,AR
211 212 213
Evenus satyroides ♂ Evenus satyroides ♂ Evenus satyroides ♂

KK DvG GC
EC-Atrk, CO-BO,VZ,BR PE-, CO-BO,VZ,BR BR-CJL, CO-BO,VZ,BR

29
Tribu - Tribe
Eumaeini
214 215 216
Evenus satyroides ♀ Evenus tagyra Evenus tagyra

DvG KK WC
PE-, CO-BO,VZ,BR PE-ARCC, PE-C BR BR-CJL, PE-C BR
217 218
Subtribu - Subtribe
Jantheclina
Allosmaitia myrtusa Allosmaitia myrtusa

DvG DvG
PE-Jgu, PN-BO EC-Orq, PN-BO
219 220 221
Allosmaitia strophius Allosmaitia strophius Allosmaitia strophius

DvG DvG AN
PE-ARCC, US(Tx)-BO,BR,AR PE-ARCC, US(Tx)-BO,BR,AR PE-VCm, US(Tx)-BO,BR,AR
222 223 224
Allosmaitia strophius Allosmaitia strophius Allosmaitia strophius

DvG DvG DvG
PE-Qtz, US(Tx)-BO,BR,AR EC-Atrk, US(Tx)-BO,BR,AR EC-WS, US(Tx)-BO,BR,AR

30
Tribu - Tribe
Eumaeini
225 226 227
Aveexcrenota anna ♂ Contrafacia imma Contrafacia imma

Theritas DPt
CO-PYm, 2100m Nov04 CO-PE CO-VCC, 1650m Mar18 MX-BO,BR,AR CO-VCC, 1650m Mar18 MX-BO,BR,AR
228 229 230
Contrafacia imma Contrafacia imma Contrafacia imma
CO-VCC, 1650m Mar18 MX-BO,BR,AR CO-VPN, 1450m MX-BO,BR,AR CO-VPN, 1450m MX-BO,BR,AR
231 232 233
Enos falerina Enos myrtea Enos polka

RnM FM DvG
VZ-, MX-BO,BR CO-RNM, 150m Jun21 CO-PE,BR PE-Qtz, EC-BO,BR
234 235 236
Enos thara Janthecla leea Janthecla leea

BBn DvG AN
PN-Vant, Jul07 MX-AR BR-RG, CO-BO,BR,GF EC-Atrk, CO-BO,BR,GF

31
Tribu - Tribe
Eumaeini
237 238 239
Janthecla malvina Janthecla rocena Janthecla rocena

DW DvG DvG
PE-Jgu, PE-BO,BR(RJ) PE-ARCC, MX-BO,BR,AR PE-ARCC, MX-BO,BR,AR
240 241 242
Janthecla rocena Janthecla sista Janthecla sista

KK DvG DvG
PE-ARCC, MX-BO,BR,AR PE-Qtz, CO-BO,BR,VZ PE-Qtz, CO-BO,BR,VZ
243 244 245
Janthecla sista Laothus gibberosa ♂ Laothus gibberosa ♂

AN Gibbosa FM Gibbosa DvG
PE-, CO-BO,BR,VZ CO-RP, 560m CO-BO EC-Rho, CO-BO
246 247 248
Laothus gibberosa ♂ Laothus gibberosa ♀ Laothus gibberosa ♀

Gibbosa Gibbosa DvG Gibbosa DvG
CO-, - CO-BO EC-Orq, CO-BO PE-Qtz, CO-BO

32
Tribu - Tribe
Eumaeini
249 250 251
Laothus gibberosa ♀ Laothus gibberosa ♀ Laothus gibberosa

Gibbosa KG Gibbosa Gibbosa DvG
CO-PSJ, 1800m CO-BO CO-PSJ, 1800m CO-BO PE-Qtz, CO-BO
252 253 254
Laothus viridicans Laothus viridicans Laothus viridicans

AN DvG DvG
EC-Loj, VZ-BO PE-Qtz, VZ-BO EC-Cal, VZ-BO
255
Laothus viridicans
CO-, - VZ-BO
256 257
Subtribu - Subtribe
Paiwarriina
Annamaria ganimedes Annamaria ganimedes

Lamasina DvG Lamasina DvG
PE-PMp, Sep28 Guianas-PE PE-Qtz, Nov15 Guianas-PE

33
Tribu - Tribe
Eumaeini
258 259 260
Annamaria rhaptissima ♂ Annamaria rhaptissima ♀ Annamaria rhaptissima ♀

Airamanna DvG Airamanna DvG Airamanna AN
PE-Mrd, CO-BO PE-Qtz, CO-BO PE-PNM, CO-BO
261 262 263
Fasslantonius episcopalis ♂ Fasslantonius episcopalis ♂ Fasslantonius episcopalis ♀

Paiwarria Leo Paiwarria Leo Paiwarria TGz
CO-Mzm, CO-PE CO-Mzm, CO-PE CO-, Oct02 CO-PE
264 265 266
Kolana ergina Kolana ergina Kolana ergina

FM DvG KK
CO-RP, 560m Nov07 CO-PE,BR(Am),GF PE-Qtz, CO-PE,BR(Am),GF BO-Vtu, CO-PE,BR(Am),GF
267 268 269
Kolana ergina Kolana ligurina Kolana ligurina

AN AN KK
PE-Qtz, CO-PE,BR(Am),GF PE-PNM, MX-BO,BR,AR EC-Atrk, MX-BO,BR,AR

34
Tribu - Tribe
Eumaeini
270 271 272
Kolana lyde Paiwarria telemus ♂ Paiwarria telemus ♀

KK GC DvG
EC-Atrk, MX-PE,VZ BR-CJL, CO-BO,BR PE-Qtz, CO-BO,BR
273 274 275
Paiwarria telemus ♀ Paiwarria telemus ♀ Paiwarria venulius

DvG WC DvG
EC-Atrk, CO-BO,BR BR-CJL, CO-BO,BR PE-ARCC, CO-BO,BR,AR
276
Paiwarria venulius
MCC
BR-, Nov19 CO-BO,BR,AR
277 278
Subtribu - Subtribe
Cupatheclina
Cupathecla cupentus Cupathecla cupentus

Megathecla THn Megathecla DvG
EC-, MX-BO,GY,BR CO-Mzm, 1400m Jun23 MX-BO,GY,BR

35
Tribu - Tribe
Eumaeini
279
Cupathecla cupentus
Megathecla DvG
PE-ARCC, Sep28 MX-BO,GY,BR
280 281
Subtribu - Subtribe
Parrhasiina
Apuecla picus Dicya carnica

DvG DvG
PE-MWC, Oct20 PE-BR,GY,GF AR-ChM, MX-AR
282 283 284
Dicya carnica Dicya carnica Dicya dicaea

DvG KG DvG
EC-Cpl, MX-AR CO-Rpau, MX-AR AR-ChM, MX-AR
285 286 287
Dicya dicaea Dicya iambe Dicya iambe

DvG
AR-ChM, Dec23 MX-AR CO-PSJ, 1800m CR-BO,GF CO-PSJ, 1800m CR-BO,GF

36
Tribu - Tribe
Eumaeini
288 289 290
Dicya iambe Dicya iambe Ignata caldas

FM DvG DvG
CO-, CR-BO,GF PE-Qtz, CR-BO,GF EC-Podc, MX-PE
291 292 293
Ignata caldas Ignata caldas Ignata levis

AN AN DvG
PE-, MX-PE PE-, MX-PE PE-PntL, PE-BO,BR(Pa)
294 295 296
Ignata levis Ignata mulsus Ignata norax

AN DvG DvG
PE-, PE-BO,BR(Pa) PE-ARCC, PE-BR(Pa),GF PE-ARCC, MX-AR
297 298 299
Michaelus ira Michaelus ira Michaelus jebus

DvG DvG DvG
PE-Jgu, MX-AR PE-ARCC, MX-AR PE-VCm, MX-AR

37
Tribu - Tribe
Eumaeini
300 301 302
Michaelus jebus Michaelus jebus Michaelus jebus

DvG
CO-VPN, 1450m MX-AR PE-Qtz, MX-AR CO-VPN, 1450m MX-AR
303 304 305
Michaelus jebus Michaelus jebus Michaelus joseph

AN BBt DvG
PE-, MX-AR EC-Podc, MX-AR PE-ARCC, CR-PE,BR
306 307 308
Michaelus joseph Michaelus joseph Michaelus phoenissa

DvG DvG DvG
PE-ARCC, CR-PE,BR PE-ARCC, CR-PE,BR EC-Podc, MX-PE,BR,GF
309 310 311
Michaelus phoenissa Michaelus phoenissa Michaelus phoenissa

DvG
CO-CRC, 320m Ene10 MX-PE,BR,GF CO-Tft, 574m MX-PE,BR,GF PE-PntL, MX-PE,BR,GF

38
Tribu - Tribe
Eumaeini
312 313 314
Michaelus phoenissa Michaelus phoenissa Michaelus thordesa

FM DvG
CO-RP, 560m Nov07 MX-PE,BR,GF CO-Tft, 574m MX-PE,BR,GF PE-ARCC, MX-BO,AR
315 316 317
Michaelus thordesa Michaelus thordesa Oenomaus andi

DvG DvG AN
PE-ARCC, MX-BO,AR PE-Qtz, MX-BO,AR PE-VCm, EC-PE
318 319 320
Oenomaus andi Oenomaus atena Oenomaus atena

DvG DvG DvG
PE-Qtz, EC-PE PE-ARCC, CR-PE,VZ,BR PE-ARCC, CR-PE,VZ,BR
321 322 323
Oenomaus atesa Oenomaus atesa Oenomaus gemma

DvG DvG Porthecla DvG
EC-BBR, MX-PE,BR,GF EC-BBR, MX-PE,BR,GF PE-CJL, CO-PE,BR

39
Tribu - Tribe
Eumaeini
324 325 326
Oenomaus gemma Oenomaus gemma Oenomaus jauffreti

Porthecla KG Porthecla KG DvG
CO-CCri, 280m Ago13 CO-PE,BR CO-CCri, 280m Ago13 CO-PE,BR PE-Qtz, PE-BR(Pa),GF
327 328 329
Oenomaus lea Oenomaus minyia Oenomaus minyia

DvG Porthecla WC Porthecla DvG
PE-ARCC, Oct09 EC-PE BR-CJL, VZ-PE,BR(Am),GF EC-Atrk, VZ-PE,BR(Am),GF
330 331 332
Oenomaus minyia Oenomaus ortygnus Oenomaus ortygnus

Porthecla SDM DvG DW
BR-CJL, VZ-PE,BR(Am),GF PE-ARCC, MX-BO,BR PN-Bqt, MX-BO,BR
333 334 335
Oenomaus ortygnus Oenomaus ortygnus Olynthus essus

DGz RC
CO-RlT, MX-BO,BR CO-, - MX-BO,BR BR-CJL, PE-BR,SU,GF

40
Tribu - Tribe
Eumaeini
336 337 338
Olynthus lorea Olynthus narbal Olynthus narbal

DvG TS
PE-MWC, PE-BR,SU,GF PN-CTwr, PN-PE,VZ,BR,GF CO-Mor, 80m May09 PN-PE,VZ,BR,GF
339 340 341
Olynthus narbal Olynthus narbal Olynthus obsoleta

BB AN MCC
CO-Mor, PN-PE,VZ,BR,GF EC-Atrk, PN-PE,VZ,BR,GF BR-, Mar27 PE-BO,BR(Rd)
342 343 344
Olynthus obsoleta Olynthus ophelia Parrhasius orgia

MCC DvG
BR-, PE-BO,BR(Rd) PE-ARCC, CO-PE,BR,AR,GF CO-Pña, MX-AR
345 346 347
Parrhasius orgia Parrhasius orgia Parrhasius orgia

DvG DvG DvG
PE-Qtz, MX-AR PE-ARCC, MX-AR PE-ARCC, MX-AR

41
Tribu - Tribe
Eumaeini
348 349 350
Parrhasius orgia Parrhasius polibetes Parrhasius polibetes

DvG DvG
CO-MMt, 200m MX-AR CO-HLB, 524m - MX-AR PE-Qtz, MX-AR
351 352 353
Parrhasius polibetes Parrhasius polibetes Parrhasius polibetes

DvG
CO-PSJ, 1800m Feb07 MX-AR PE-Jul, MX-AR CO-PSJ, 1800m Feb07 MX-AR
354 355 356
Parrhasius polibetes Parrhasius polibetes Parrhasius polibetes

FM
CO-, May31 MX-AR CO-Gurc, 690m - MX-AR CO-Jard, MX-AR
357 358 359
Parrhasius selika Parrhasius selika Parrhasius selika

PB
CO-Jard, CO-BO,BR CO-, 2600m Oct13 CO-BO,BR CO-PSJ, 1800m Ago06 CO-BO,BR

42
Tribu - Tribe
Eumaeini
360 361 362
Parrhasius selika Parrhasius selika Parrhasius selika

AN
PE-Qtz, CO-BO,BR CO-PSJ, 1800m Ago06 CO-BO,BR CO-QEV, 1940m CO-BO,BR
363 364 365
Parrhasius selika Symbiopsis aprica Symbiopsis aprica

DvG THn
CO-QEV, 1940m CO-BO,BR PE-ARCC, Sep25 CO?EC-PE,BR,SU PE-, CO?EC-PE,BR,SU
366 367 368
Symbiopsis aprica Symbiopsis perulera Symbiopsis perulera

DvG DvG DvG
PE-Qtz, Nov11 CO?EC-PE,BR,SU PE-Rmpt, Sep30 PE-BR PE-ARCC, Oct07 PE-BR
369 370 371
Symbiopsis perulera Symbiopsis pupilla Symbiopsis pupilla

DvG
PE-Rmpt, Sep27 PE-BR CO-RSm, 400m Jul19 CO-BO,BR,GY,GF CO-RSm, 400m Jul19 CO-BO,BR,GY,GF

43
Tribu - Tribe
Eumaeini
372 373
Thepytus echelta Thepytus thyrea

KG JAR
BR-Rgu, MX-BR,GF BR-Blm, EC-PE,BR,AR,GF
374 375
Subtribu - Subtribe
Ipideclina
Ipidecla crepundia Ipidecla crepundia

AN AN
PE-VCm, PE-BO,BR,GF PE-, PE-BO,BR,GF
376 377 378
Ipidecla crepundia Ipidecla schausi ♂ Ipidecla schausi ♀

DvG MM
BO-Volc, PE-BO,BR,GF CO-Gai, 582m MX-AR CO-Tft, 574m MX-AR
379
Ipidecla schausi ♀
DvG
PE-ARCC, MX-AR

44
Tribu - Tribe
Eumaeini
380 381
Subtribu - Subtribe
Calycopidina
Arzecla albolineata Arzecla arza

DvG
CO-Mzm, 1600m Oct01 CO-PE CO-PSJ, 1850m Oct17 MX-BO,BR,AR
382 383 384
Arzecla arza Arzecla arza Arzecla arza

AN BBt
CO-PSJ, 1800m Nov14 MX-BO,BR,AR PE-, MX-BO,BR,AR CO-Sin, 700m May21 MX-BO,BR,AR
385 386 387
Arzecla arza Arzecla calatia Arzecla calatia

DvG KG DvG
PE-ARCC, MX-BO,BR,AR PE-ARCC, MX,NI-PE,BR,GF PE-MWC, MX,NI-PE,BR,GF
388 389 390
Arzecla canacha Arzecla canacha Arzecla canacha

DvG
CO-Pña, VZ-PE CO-Jard, VZ-PE EC-Loj, VZ-PE

45
Tribu - Tribe
Eumaeini
391 392 393
Arzecla canacha Arzecla canacha Arzecla canacha

RW KK DvG
EC-, VZ-PE BO-Volc, VZ-PE CO-, 1200m Ene12 VZ-PE
394 395 396
Arzecla paralus Arzecla paralus Arzecla taminella

DvG DvG
PE-CoR, GT-PE CO-PSJ, 1800m GT-PE CO-Mzm, 1400m MX-BO,BR,GF,AR
397 398 399
Arzecla taminella Arzecla taminella Arzecla taminella

KK DvG DvG
EC-Ten, MX-BO,BR,GF,AR PE-ARCC, MX-BO,BR,GF,AR PE-ARCC, MX-BO,BR,GF,AR
400 401 402
Arzecla tucumanensis Arzecla tucumanensis Arzecla tucumanensis

DvG MCC KK
PE-TngM, PE-BO,BR,AR BR-, PE-BO,BR,AR PE-Qtz, PE-BO,BR,AR

46
Tribu - Tribe
Eumaeini
403 404 405
Arzecla tucumanensis Badecla badaca ♀ Badecla badaca

DvG Lamprospilus DvG Lamprospilus DvG
PE-TngM, PE-BO,BR,AR PE-PntL, VZ-BO,BR,AR PE-ARCC, VZ-BO,BR,AR
406 407 408
Badecla badaca Badecla badaca Badecla badaca

Lamprospilus AH Lamprospilus MIW Lamprospilus DvG
CO-, Feb01 VZ-BO,BR,AR BR-, VZ-BO,BR,AR BO-Cor, VZ-BO,BR,AR
409 410 411
Badecla lanckena Badecla lanckena Badecla sp

DvG DvG DvG
PE-ARCC, PE-BO PE-CoR, PE-BO PE-ARCC, PE
412 413 414
Calycopis anastasia ♀ Calycopis anastasia Calycopis anthora

DvG DvG KK
BO-LHt, PE-BO BO-Pcy, PE-BO PE-, CO-BO,BR

47
Tribu - Tribe
Eumaeini
415 416 417
Calycopis anthora Calycopis anthora Calycopis anthora

FM JZm SO
CO-RP, 560m CO-BO,BR EC-, Mar09 CO-BO,BR EC-, Abr04 CO-BO,BR
418 419 420
Calycopis atnius Calycopis atnius Calycopis atnius

GC DvG
CO-, - CR-PE,BR,GF BR-CJL, CR-PE,BR,GF PE-PntL, CR-PE,BR,GF
421 422 423
Calycopis atnius Calycopis atnius Calycopis bactra

DvG DvG AH
PE-PntL, CR-PE,BR,GF PE-ARCC, CR-PE,BR,GF CO-SM, NI-PE,VZ,GF
424 425 426
Calycopis bactra Calycopis bellera ♀ Calycopis bellera

DvG KG DvG
PN-PplRd, NI-PE,VZ,GF PE-ARCC, PN-BO,BR,GF,AR EC-Orq, PN-BO,BR,GF,AR

48
Tribu - Tribe
Eumaeini
427 428 429
Calycopis bellera Calycopis bellera Calycopis bellera

AN DvG
EC-Atrk, PN-BO,BR,GF,AR CO-Pña, PN-BO,BR,GF,AR PE-Pro, PN-BO,BR,GF,AR
430 431 432
Calycopis buphonia Calycopis buphonia Calycopis buphonia

DvG AN
CO-CRC, 320m Ene12 CR-PE,BR,GF PE-ARCC, CR-PE,BR,GF PE-, CR-PE,BR,GF
433 434 435
Calycopis buphonia Calycopis caesaries Calycopis caesaries

DvG DvG KG
EC-PmpL, CR-PE,BR,GF PE-ARCC, BZ-PE,BR CO-CRC, 320m Ago31 BZ-PE,BR
436 437 438
Calycopis calus Calycopis calus Calycopis calus
CO-, Dic31 MX-BO,AR CO-, MX-BO,AR CO-PSJ, 1800m Abr26 MX-BO,AR

49
Tribu - Tribe
Eumaeini
439 440 441
Calycopis calus Calycopis calus Calycopis caulonia

DvG DvG
EC-Podc, MX-BO,AR CO-, Dic31 MX-BO,AR AR-ChM, PE-BO,BR,AR
442 443 444
Calycopis caulonia Calycopis centoripa ♂ Calycopis centoripa ♂

DvG DvG AN
AR-ChM, PE-BO,BR,AR EC-AJL, 400m CO-BO,BR PE-, CO-BO,BR
445 446 447
Calycopis centoripa ♀ Calycopis centoripa ♀ Calycopis cerata

DvG GC
PE-Aco, CO-BO,BR BR-CJL, CO-BO,BR CO-HFr, 1100m Mar11 HN-AR
448 449 450
Calycopis cerata Calycopis cerata Calycopis cerata

DvG
CO-HFr, 1100m Mar11 HN-AR CO-CRC, 550m HN-AR CO-RSm, 400m Jul19 HN-AR

50
Tribu - Tribe
Eumaeini
451 452 453
Calycopis cerata Calycopis cerata Calycopis cicero

DvG DvG DvG
PE-ARCC, HN-AR PE-ARCC, HN-AR PE-PNYC, EC-PE
454 455 456
Calycopis cicero Calycopis demonassa Calycopis demonassa

DvG
PE-Jgu, EC-PE CO-Sin, 700m Nov01 MX-BO,BR,GF CO-VPN, 1450m MX-BO,BR,GF
457 458 459
Calycopis demonassa Calycopis demonassa Calycopis demonassa
CO-CRC, 350m Ago23 MX-BO,BR,GF CO-VPN, 1450m Sep21 MX-BO,BR,GF CO-VPN, 1450m Sep21 MX-BO,BR,GF
460 461 462
Calycopis demonassa Calycopis demonassa Calycopis gentilla ♂

DvG
CO-, - MX-BO,BR,GF CO-Sin, 700m Nov01 MX-BO,BR,GF AR-ChM, PE,BR(RJ)-AR

51
Tribu - Tribe
Eumaeini
463 464 465
Calycopis gentilla Calycopis johnsoni Calycopis johnsoni

DvG RRt DvG
BO-Chl, PE,BR(RJ)-AR EC-Cpl, CO-BO PE-Qtz, CO-BO
466 467 468
Calycopis lerbela Calycopis malta Calycopis mimas

DvG DvG DvG
GY-, EC-BR(Pa),GF PE-PH, CO?EC-BO,BR EC-Podc, PN-PE,BR,GF
469 470 471
Calycopis mimas Calycopis nicolayi Calycopis orcilla

DvG DvG DvG
EC-Podc, PN-PE,BR,GF BR-SBr, PE-BO,BR(Pb) PE-Qtz, CO?EC-BO
472 473 474
Calycopis orcillula ♀ Calycopis orcillula Calycopis orcillula

DvG AN DvG
PN-PplRd, HN-PE,BR,VZ EC-Atrk, HN-PE,BR,VZ PN-PplRd, HN-PE,BR,VZ

52
Tribu - Tribe
Eumaeini
475 476 477
Calycopis orcillula Calycopis origo Calycopis origo

DvG DvG DvG
PN-PplRd, HN-PE,BR,VZ PN-PplRd, CR-BO,BR,GF PE-, CR-BO,BR,GF
478 479 480
Calycopis partunda Calycopis petaurister Calycopis pisis

DvG DvG
PE-PNYC, PE-BR CO-VCC, 1640m Oct11 VZ-PE,GY PE-ARCC, PN-PE,BR
481 482 483
Calycopis pisis Calycopis suda Calycopis suda

DvG KK KK
PN-PplRd, PN-PE,BR PE-BSht, PE-BO PE-BSht, PE-BO
484 485 486
Calycopis tamos Calycopis tamos Calycopis trebula

DvG DvG KG
CO-Mzm, 1400m CR-PE CO-Mzm, 1400m CR-PE HN-, MX-PE,BR

53
Tribu - Tribe
Eumaeini
487 488 489
Calycopis trebula Calycopis trebula Calycopis trebula

DvG DvG DvG
PE-ARCC, MX-PE,BR PE-VCm, MX-PE,BR PE-ARCC, MX-PE,BR
490 491 492
Calycopis wolfii Calycopis wolfii Calycopis xeneta ♂

DvG DvG DvG
PE-ARCC, CO-PE,BR,GF PE-ARCC, CO-PE,BR,GF PE-ARCC, NI-PE,VZ,GF
493 494 495
Calycopis xeneta ♀ Calycopis xeneta Calycopis xeneta

KK DvG DvG
PE-Pro, NI-PE,VZ,GF PE-PNYC, NI-PE,VZ,GF PE-ARCC, NI-PE,VZ,GF
496 497 498
Camissecla camissa Camissecla camissa Camissecla cleocha

DvG RC KG
PE-VCm, CO? EC-PE PE-TRC, CO? EC-PE CO-Afl, CO-PE

54
Tribu - Tribe
Eumaeini
499 500 501
Camissecla cleocha Camissecla cleocha Camissecla cleocha

DvG KG
PE-Qtz, 1150m CO-PE PE-Qtz, 1150m CO-PE CO-PSJ, 1800m Nov21 CO-PE
502 503 504
Camissecla pactya Camissecla pactya Camissecla pactya

DvG MC JHg
PE-, CO-BO CO-, 2090m CO-BO EC-WS, CO-BO
505 506 507
Camissecla verbenaca Camissecla verbenaca Camissecla vesper

DvG DvG DvG
CO-Mzm, VZ-PE,BR CO-Mzm, VZ-PE,BR PE-ARCC, CO-BO,BR,GF
508 509 510
Camissecla vesper Electrostrymon endymion Electrostrymon joya

DvG RH DvG
PE-VCm, CO-BO,BR,GF BR-CJL, PE-BO,BR,AR EC-Vilc, Nov30 US(Tx)-PE

55
Tribu - Tribe
Eumaeini
511 512 513
Electrostrymon joya Gigantorubra aruma ♀ Gigantorubra aruma

DvG Arumecla DvG Arumecla DvG
PE-Cus, Aug22 US(Tx)-PE PE-CoR, CO-PE,BR,GF PE-ARCC, CO-PE,BR,GF
514 515 516
Gigantorubra aruma Gigantorubra netesca ♂♀ Gigantorubra orcidia ♂

Arumecla AH Arumecla RnM Lamprospilus DvG
CO-SM, Feb05 CO-PE,BR,GF VZ-, PE-BO,GY PE-ARCC, MX-BO,BR,PY,GF
517 518 519
Gigantorubra orcidia ♂ Gigantorubra orcidia ♀ Gigantorubra orcidia

Lamprospilus DvG Lamprospilus DvG Lamprospilus
PE-ARCC, MX-BO,BR,PY,GF PE-ARCC, MX-BO,BR,PY,GF CO-HLB, 524m - MX-BO,BR,PY,GF
520 521 522
Gigantorubra orcidia Kisutam syllis Kisutam syllis

Lamprospilus THn FM
PE-, MX-BO,BR,PY,GF CO-, US(Tx)-PE,BR,GF CO-, May31 US(Tx)-PE,BR,GF

56
Tribu - Tribe
Eumaeini
523 524 525
Kisutam syllis Kisutam syllis Kisutam syllis

DvG DvG KK
EC-Podc, US(Tx)-PE,BR,GF PE-MWC, US(Tx)-PE,BR,GF CO-MMt, US(Tx)-PE,BR,GF
526 527 528
Lamprospilus coelicolor ♂ Lamprospilus coelicolor ♀ Lamprospilus coelicolor ♀

AN KK DvG
PE-, MX-BO,BR,GF PE-Qtz, MX-BO,BR,GF CO-Mzm, 1300m Sep30 MX-BO,BR,GF
529 530 531
Lamprospilus coelicolor ♀ Lamprospilus decorata ♀ Lamprospilus decorata

AN DvG DvG
PE-Qtz, MX-BO,BR,GF PE-, CO-PE PE-VCm, CO-PE
532 533 534
Lamprospilus draudti ♂ Lamprospilus draudti ♂ Lamprospilus draudti ♀

Leo AN
CO-Mzm, CO-PE CO-QEV, 1850m Feb27 CO-PE CO-PSJ, 1800m Feb07 CO-PE

57
Tribu - Tribe
Eumaeini
535 536 537
Lamprospilus draudti ♀ Lamprospilus genius ♂ Lamprospilus genius ♀

DvG DvG KK
CO-Mzm, 2100m CO-PE BR-SBr, EC-PE,BR,GF BR-, EC-PE,BR,GF
538 539 540
Lamprospilus nicetus Lamprospilus nicetus Rubroserrata ecbatana ♂

DvG THn DvG
PE-ARCC, VZ-BO EC-, VZ-BO PE-ARCC, VZ-PE,BR,GF
541 542 543
Rubroserrata ecbatana ♀ Rubroserrata ecbatana Rubroserrata ecbatana

DvG
PE-TngM, VZ-PE,BR,GF CO-Pña, VZ-PE,BR,GF CO-VCC, 1650m Mar18 VZ-PE,BR,GF
544 545 546
Rubroserrata ecbatana Rubroserrata ecbatana Ziegleria ceromia
CO-CRC, 320m Ene26 VZ-PE,BR,GF CO-SNc, 474m Nov23 VZ-PE,BR,GF CO-Sin, 780m May24 MX-BO,BR,UR,GF

58
Tribu - Tribe
Eumaeini
547 548 549
Ziegleria ceromia Ziegleria ceromia Ziegleria hesperitis

DvG
CO-Sin, 780m May24 MX-BO,BR,UR,GF CO-Sin, 780m May21 MX-BO,BR,UR,GF CO-VCC, 1650m Mar18 MX-BO,BR,AR,GF
550 551 552
Ziegleria hesperitis Ziegleria hesperitis Ziegleria hesperitis

LC
CO-VCC, 1650m Mar18 MX-BO,BR,AR,GF EC-Atrk, Oct31 MX-BO,BR,AR,GF CO-, MX-BO,BR,AR,GF
553 554
Subtribu - Subtribe
Strymonina
Arawacus dolylas Arawacus dolylas
CO-VPN, 1450m PN-BR,AR CO-, - PN-BR,AR
555 556 557
Arawacus dolylas Arawacus ellida Arawacus ellida

KG KG
CO-VPN, 1450m PN-BR,AR PE-Rmp-Mq, Nov14 VZ-BO,BR,AR PE-Rmp-Mq, Nov14 VZ-BO,BR,AR

59
Tribu - Tribe
Eumaeini
558 559 560
Arawacus ellida Arawacus separata Arawacus separata

TH FM KK
BO-Corc, Nov16 VZ-BO,BR,AR CO-RP, 560m Ago01 CO-AR CO-MMt, 200m Oct17 CO-AR
561 562 563
Hypostrymon aepeona Hypostrymon asa Hypostrymon asa

KG DvG DvG
PE-BSht, EC-PE PE-ARCC, GT-PE,BR,GF,AR EC-Podc, GT-PE,BR,GF,AR
564 565 566
Hypostrymon asa Hypostrymon asa Hypostrymon renidens

DvG DvG DvG
EC-Atrk, GT-PE,BR,GF,AR CO-MMt, 200m GT-PE,BR,GF,AR PE-CoR, CO?EC-PE
567 568 569
Hypostrymon renidens Panthiades aeolus Panthiades aeolus

DvG FM DvG
CO-RAA, CO?EC-PE CO-RNM, 150m Jun21 PN-BO,BR CO-MMt, 200m Mar06 PN-BO,BR

60
Tribu - Tribe
Eumaeini
570 571 572
Panthiades aeolus Panthiades aeolus Panthiades bitias ♂

DvG DvG
PE-Pro, PN-BO,BR PE-Rmp-Mq, PN-BO,BR CO-RSm, 400m Ago02 MX-BO,BR,GF
573 574 575
Panthiades bitias ♂ Panthiades bitias ♂ Panthiades bitias ♂
CO-HLB, 524m - MX-BO,BR,GF CO-RSm, 400m Jul19 MX-BO,BR,GF CO-QLH, 750m Jul25 MX-BO,BR,GF
576 577 578
Panthiades bitias ♀ Panthiades bitias ♀ Panthiades bitias ♀

SS
CO-, - MX-BO,BR,GF CO-HLB, 524m - MX-BO,BR,GF CO-MMt, 200m MX-BO,BR,GF
579 580 581
Panthiades bitias ♀ Panthiades bitias ♀ Panthiades paphlagon

DvG
CO-HLB, 524m Ago18 MX-BO,BR,GF CO-Rch, 1000m Feb06 MX-BO,BR,GF PE-PNYC, CO-PE,VZ

61
Tribu - Tribe
Eumaeini
582 583 584
Panthiades paphlagon Panthiades paphlagon Panthiades paphlagon

KK LC
BO-Volc, 1400m CO-PE,VZ CO-PSJ, 1800m CO-PE,VZ CO-Mzm, CO-PE,VZ
585 586 587
Panthiades paphlagon Panthiades phaleros Panthiades phaleros

KG KG DvG
PE-, CO-PE,VZ PA-, MX-AR PE-ARCC, MX-AR
588 589 590
Panthiades phaleros Panthiades phaleros Panthiades phaleros

DvG KG KG
PE-ARCC, MX-AR PA-, MX-AR PE-ARCC, MX-AR
591 592 593
Rekoa marius Rekoa marius Rekoa marius

DvG PS
CO-, US(Tx)-AR PE-PH, US(Tx)-AR PE-PNYC, US(Tx)-AR

62
Tribu - Tribe
Eumaeini
594 595 596
Rekoa marius Rekoa meton Rekoa meton

KK DvG
PE-Pro, US(Tx)-AR CO-HLB, 524m - MX-AR EC-Podc, MX-AR
597 598 599
Rekoa meton Rekoa meton Rekoa palegon

DvG DvG
PE-VCm, MX-AR CO-Sin, 780m Nov15 MX-AR CO-Mzm, 1350m US(Tx)-AR
600 601 602
Rekoa palegon Rekoa palegon Rekoa palegon

DvG DvG
PE-Qtz, US(Tx)-AR EC-Pro, US(Tx)-AR CO-Jard, 1700m May30 US(Tx)-AR
603 604 605
Rekoa palegon Rekoa palegon Rekoa palegon

DvG
CO-Mzm, 1350m US(Tx)-AR CO-, US(Tx)-AR CO-, - US(Tx)-AR

63
Tribu - Tribe
Eumaeini
606 607 608
Rekoa palegon Rekoa stagira ♀ Rekoa stagira

KK DvG
EC-Atrk, US(Tx)-AR CO-, - MX-AR PE-AMZ, Oct26 MX-AR
609 610 611
Strymon astiocha Strymon astiocha Strymon astiocha

KG DW KG
BO-Chl, Nov15 MX-AR PE-Hual, Nov16 MX-AR BO-Chl, Nov15 MX-AR
612 613 614
Strymon astiocha Strymon astiocha Strymon bazochii

DW KG MG
PE-Hual, Nov16 MX-AR PE-Hual, Nov16 MX-AR CO-, Ene02 US(Tx)-AR
615 616 617
Strymon bazochii bazochii Strymon bazochii bazochii Strymon bazochii bazochii
CO-, - US(Tx)-AR CO-, - US(Tx)-AR CO-Sin, 780m Jun14 US(Tx)-AR

64
Tribu - Tribe
Eumaeini
618 619 620
Strymon bazochii bazochii Strymon bazochii bazochii Strymon bazochii bazochii
CO-Gai, 600m US(Tx)-AR CO-Sin, 780m Jun14 US(Tx)-AR CO-, US(Tx)-AR
621 622 623
Strymon bazochii bazochii Strymon bubastus bubastus ♂ Strymon bubastus bubastus ♂
CO-VPN, 1450m Feb19 US(Tx)-AR CO-, VZ-PE,BR,AR CO-, VZ-PE,BR,AR
624 625 626
Strymon bubastus bubastus Strymon bubastus bubastus Strymon bubastus bubastus

DvG
CO-HLB, 524m Oct24 VZ-PE,BR,AR CO-Sin, 700m May21 VZ-PE,BR,AR CO-HLB, 524m - VZ-PE,BR,AR
627 628 629
Strymon bubastus bubastus Strymon bubastus bubastus Strymon bubastus bubastus
CO-HLB, 524m Oct24 VZ-PE,BR,AR CO-CCri, 300m Ago06 VZ-PE,BR,AR CO-Tft, 574m VZ-PE,BR,AR

65
Tribu - Tribe
Eumaeini
630 631 632
Strymon bubastus bubastus Strymon bubastus bubastus Strymon cestri
CO-, - VZ-PE,BR,AR CO-, - VZ-PE,BR,AR CO-, US(Az,Tx)-BO,BR,AR
633 634 635
Strymon cestri Strymon cestri Strymon crambusa

DvG
CO-, US(Az,Tx)-BO,BR,AR CO-, US(Az,Tx)-BO,BR,AR PE-Rmpt, Sep24 PE-BO,BR,AR
636 637 638
Strymon crambusa Strymon crambusa Strymon crambusa

KG DvG DvG
BR-RG, Nov21 PE-BO,BR,AR PE-Rmpt, Sep24 PE-BO,BR,AR PE-Rmpt, Sep24 PE-BO,BR,AR
639 640 641
Strymon crambusa Strymon crambusa Strymon daraba

KG DvG DvG
BR-RG, Nov21 PE-BO,BR,AR PE-Rmpt, Sep24 PE-BO,BR,AR PE-, EC-PE

66
Tribu - Tribe
Eumaeini
642 643 644
Strymon daraba Strymon daraba Strymon daraba

DvG KK DvG
PE-, EC-PE PE-, EC-PE PE-, EC-PE
645 646 647
Strymon daraba Strymon davara Strymon gabatha ♀

KK AN DvG
PE-, EC-PE PE-, EC-BO,BR CO-Mzm, MX-PE,VZ
648 649 650
Strymon gabatha ♀ Strymon gabatha Strymon gabatha

DvG DvG
CO-Mzm, MX-PE,VZ CO-Mzm, 1350m Sep25 MX-PE,VZ CO-CRC, 320m MX-PE,VZ
651 652 653
Strymon istapa istapa Strymon istapa istapa Strymon istapa istapa

DvG KG DvG
BR-, MX-AR MX-Plq, MX-AR PE-, MX-AR

67
Tribu - Tribe
Eumaeini
654 655 656
Strymon istapa istapa Strymon megarus Strymon megarus

KG FM
MX-Cat, MX-AR CO-RP, 560m Ago01 MX-BO,BR,AR CO-CRC, 320m Ene12 MX-BO,BR,AR
657 658 659
Strymon megarus Strymon michelle Strymon michelle

DvG DvG
CO-CRC, 320m Ene12 MX-BO,BR,AR PE-, EC-PE EC-, Dic08 EC-PE
660 661 662
Strymon mulucha Strymon mulucha Strymon mulucha
CO-VCC, 1650m Mar18 MX-BO,BR,AR CO-VCC, 1650m Mar18 MX-BO,BR,AR CO-, - MX-BO,BR,AR
663 664 665
CO-SNc, 474m Dic07 MX-BO,BR,AR CO-PSJ, 1800m MX-BO,BR,AR CO-Tft, 574m MX-BO,BR,AR

68
Tribu - Tribe
Eumaeini
666 667 668
CO-RSD, MX-BO,BR,AR CO-, MX-BO,BR,AR CO-, MX-BO,BR,AR
669 670 671
Strymon mulucha Strymon mulucha Strymon rufofusca
CO-RSm, 400m Ago02 MX-BO,BR,AR CO-PSJ, 1800m Dic05 MX-BO,BR,AR CO-SNc, 474m Dic14 US(Tx)-AR
672 673 674
Strymon rufofusca Strymon rufofusca Strymon rufofusca
CO-Gai, 582m Ago24 US(Tx)-AR CO-SNc, 474m Nov16 US(Tx)-AR CO-Gai, 582m Ago24 US(Tx)-AR
675 676 677
CO-, US(Tx)-AR CO-, US(Tx)-AR CO-Gai, 582m Ago24 US(Tx)-AR

69
Tribu - Tribe
Eumaeini
678 679 680

LEM
CO-Sin, 780m Ago23 US(Tx)-AR CO-HLB, 524m - US(Tx)-AR CO-Sin, 780m Ago23 US(Tx)-AR
681 682 683
Strymon yojoa Strymon yojoa Strymon yojoa

DvG DvG
CO-Mnc, 600m US(Tx,Az)-PE,BR,GF CO-SNc, 474m Nov23 US(Tx,Az)-PE,BR,GF CO-Mnc, 600m US(Tx,Az)-PE,BR,GF
684 685 686
Strymon yojoa Strymon yojoa Strymon ziba
CO-SNc, 474m Nov23 US(Tx,Az)-PE,BR,GF CO-SNc, 474m Nov23 US(Tx,Az)-PE,BR,GF CO-Sin, 700m Sep20 MX-BO,AR,GF
687 688 689
Strymon ziba Strymon ziba Strymon ziba
CO-Sin, 780m Ago23 MX-BO,AR,GF CO-Sin, 780m Ago23 MX-BO,AR,GF CO-, MX-BO,AR,GF

70
Tribu - Tribe
Eumaeini
690 691 692
Strymon ziba Strymon ziba Thereus cithonius

Noreena FM
CO-, - MX-BO,AR,GF CO-Mor, 80m May01 MX-BO,AR,GF CO-RP, 560m Nov07 MX-AR
693 694 695
Thereus cithonius Thereus cithonius Thereus endera

Noreena FM Noreena FM DvG
CO-, May31 MX-AR CO-RP, 560m Ago01 MX-AR CO-MMt, CO-PE,BR(Am)
696 697 698
Thereus endera Thereus ortalus Thereus ortalus

RW KG BBn
PE-, CO-PE,BR(Am) MX-Ctm, MX-BO,BR,PY MX-Ctm, MX-BO,BR,PY
699 700 701
Thereus pedusa Thereus pedusa Thereus timoclea

Pedusa AN Pedusa JJ GS
PE-, PN-PE,BR CO-, 250m May12 PN-PE,BR BO-, PN-BO,GF

71
Tribu - Tribe
Eumaeini
702 703
Subtribu - Subtribe
Strephonotina
Aubergina alda Aubergina hesychia

AN AN
PE-, CO-BO,BR(Pa),GF PE-, CR-PE,BR
704 705 706
Aubergina hesychia Celmia anastomosis Celmia celmus

DvG THn
CO-Mzm, 1400m Jun09 CR-PE,BR PE-, PE-BR,Guianas CO-VPN, 1450m Sep21 MX-AR
707 708 709
Celmia celmus Celmia celmus Celmia celmus

KK
CO-VPN, 1450m Sep21 MX-AR CO-MMt, 200m Oct16 MX-AR CO-Mor, 80m May01 MX-AR
710 711 712
Celmia color Celmia conoveria Celmia conoveria

DvG AN DvG
PE-, PE-BO,BR(Am),GY EC-Atrk, MX-PE,BR,GF BR-, MX-PE,BR,GF

72
Tribu - Tribe
Eumaeini
713 714 715
Celmia mecrida Gargina caninius ♂ Gargina caninius ♂

JB DvG DvG
BR-CJL, VZ-PE,BR,Guianas PE-PntL, MX-BO,BR,GF PE-PntL, MX-BO,BR,GF
716 717 718
Gargina caninius ♂ Gargina caninius Gargina caninius

DvG KK
PE-, MX-BO,BR,GF CO-VPN, 1450m MX-BO,BR,GF EC-Ten, MX-BO,BR,GF
719 720 721
Gargina emessa ♀ Gargina emessa ♀ Gargina emessa ♀
CO-RSm, 400m Jul19 NI-PE,BR,GF CO-RSm, 400m Ago02 NI-PE,BR,GF CO-RSm, 400m Jul19 NI-PE,BR,GF
722 723 724
Gargina emessa Gargina gargophia Gargina gargophia

GM
CO-CRC, 320m Ago21 NI-PE,BR,GF CO-VCC, 1650m Mar18 MX-BO,BR,GF CO-PSJ, 1800m May15 MX-BO,BR,GF

73
Tribu - Tribe
Eumaeini
725 726 727
Gargina gargophia Gargina gargophia Gargina gnosia ♀

DvG FM DW
CO-Mzm, 1400m MX-BO,BR,GF CO-RP, 560m Feb09 MX-BO,BR,GF BO-VTn, Nov23 MX-BO,AR,GF
728 729 730
Gargina gnosia Gargina panchaea Gargina panchaea

DvG DvG DvG
BO-Chl, Nov08 MX-BO,AR,GF PE-PMp, PE-BR,AR PE-Jgu, PE-BR,AR
731 732 733
Gargina panchaea Gargina thyesta ♀ Gargina thyesta ♀

DvG DvG DvG
PE-Jul, PE-BR,AR CO-PGt, CR-BR CO-PGt, CR-BR
734 735 736
Gossenia lycabas Gossenia lycabas Gossenia lycabas

Thestius DvG Thestius DvG Thestius DvG
PE-AMZ, Oct17 CO-PE,BR PE-VCm, Oct09 CO-PE,BR PE-AMZ, CO-PE,BR

74
Tribu - Tribe
Eumaeini
737 738 739
Gossenia lycabas Iaspis castitas Iaspis temesa

Thestius DvG DW DvG
PE-ARCC, CO-PE,BR BR-RG, PE-BR(Pa,ES),GF GY-, MX-PE,BR,GY
740 741 742
Iaspis temesa Ministrymon arthuri Ministrymon arthuri

DvG DvG DvG
PE-ARCC, MX-PE,BR,GY PE-VCm, EC-PE,BR(RJ) PE-ARCC, EC-PE,BR(RJ)
743 744 745
Ministrymon arthuri Ministrymon azia Ministrymon azia

DvG KG
PE-PntL, EC-PE,BR(RJ) MX-, US(Az)-BO,BR,AR CO-PSJ, 1800m US(Az)-BO,BR,AR
746 747 748
Ministrymon azia Ministrymon azia Ministrymon azia

DvG DvG DvG
PE-ARCC, US(Az)-BO,BR,AR EC-Vilc, US(Az)-BO,BR,AR EC-Vilc, US(Az)-BO,BR,AR

75
Tribu - Tribe
Eumaeini
749 750 751
Ministrymon azia Ministrymon azia Ministrymon azia
CO-PSJ, 1800m Dic05 US(Az)-BO,BR,AR CO-PSJ, 1800m Abr26 US(Az)-BO,BR,AR CO-PSJ, 1800m US(Az)-BO,BR,AR
752 753 754
Ministrymon azia Ministrymon azia Ministrymon cleon

WC DvG
CO-PSJ, 1800m Feb07 US(Az)-BO,BR,AR MX-, US(Az)-BO,BR,AR PE-Qtz, CR-BO,BR,AR
755 756 757
Ministrymon cleon Ministrymon cleon Ministrymon cruenta

DvG DvG
EC-Orq, CR-BO,BR,AR CO-PSJ, 1800m CR-BO,BR,AR EC-Vilc, CO-BO,AR,GF
758 759 760
Ministrymon cruenta Ministrymon megacles Ministrymon megacles

DvG DvG DvG
EC-Vilc, CO-BO,AR,GF PE-ARCC, VZ-BO,BR,SU,GF PE-ARCC, VZ-BO,BR,SU,GF

76
Tribu - Tribe
Eumaeini
761 762 763
Ministrymon phrutus Ministrymon phrutus Ministrymon phrutus

LEM
CO-HLB, 524m - MX-BO,BR CO-, - MX-BO,BR CO-Gai, 582m MX-BO,BR
764 765 766
Ministrymon phrutus Ministrymon phrutus Ministrymon una una
CO-Sin, 780m Ago23 MX-BO,BR CO-Sin, 780m Oct10 MX-BO,BR CO-, - CO-BO,BR,AR
767 768 769
Ministrymon una una Ministrymon una una Ministrymon una una

KK
CO-Mor, 80m May01 CO-BO,BR,AR CO-, - CO-BO,BR,AR CO-SM, Oct11 CO-BO,BR,AR
770 771 772
Ministrymon una una Ministrymon una una Ministrymon una una
CO-Rmo, 1830m CO-BO,BR,AR CO-, - CO-BO,BR,AR CO-Mor, 80m May01 CO-BO,BR,AR

77
Tribu - Tribe
Eumaeini
773 774 775
Ministrymon zilda Ministrymon zilda Nicolaea besidia

DvG DvG DvG
PE-ARCC, MX-BO,BR,GF PE-ARCC, MX-BO,BR,GF PE-, VZ-PE,BR,GF
776 777 778
Nicolaea dolium Nicolaea munditia Nicolaea munditia

AH DvG DvG
CO-SM, Feb15 MX-BO,BR PE-VCm, EC-BO,Guianas PE-VCm, EC-BO,Guianas
779 780 781
Nicolaea munditia Nicolaea pyxis Nicolaea pyxis

DvG DvG DvG
PE-VCm, EC-BO,Guianas PE-Qtz, CO-PE,BR,GF PE-PNYC, CO-PE,BR,GF
782 783 784
Nicolaea velina Ostrinotes empusa Ostrinotes empusa

Bsu DvG DvG
MX-, MX-BR,GF CO-Mzm, 1400m Sep23 PN-AR PE-, 650m PN-AR

78
Tribu - Tribe
Eumaeini
785 786 787
Ostrinotes empusa Ostrinotes empusa Ostrinotes gentiana

DvG KK KK
PE-, 650m PN-AR PE-Jgu, PN-AR PE-Rmp-Mq, CO-BO,BR
788 789 790
Ostrinotes halciones Ostrinotes halciones Ostrinotes sospes

DvG KK DvG
CO-Mzm, 1400m Sep23 MX-PE,BR PE-Rmp-Mq, MX-PE,BR PE-ARCC, CO-PE
791 792 793
Ostrinotes tarena Ostrinotes tympania Ostrinotes tympania

KK DvG BB
PE-Qtz, VZ-PE,BR,GF PE-PMp, EC-PE,BR(Pa),GF CO-, EC-PE,BR(Pa),GF
794 795 796
Ostrinotes tympania Ostrinotes tympania Ostrinotes tympania

DvG KK DvG
PE-ARCC, EC-PE,BR(Pa),GF PE-Jgu, EC-PE,BR(Pa),GF EC-Podc, EC-PE,BR(Pa),GF

79
Tribu - Tribe
Eumaeini
797 798 799
Siderus athymbra Siderus athymbra Siderus bouvieri

DvG DvG DvG
PE-ARCC, CO-PE,BR,VZ,GF GY-, CO-PE,BR,VZ,GF PE-, EC-PE
800 801 802
Siderus leucophaeus ♀ Siderus leucophaeus ♀ Strephonota acameda

DvG DvG DvG
PE-ARCC, Nov11 GT-PE,BR,GF PE-ARCC, Nov11 GT-PE,BR,GF PE-VCm, PE-BO,BR(Pa)
803 804 805
Strephonota adela Strephonota agrippa Strephonota ambrax

GC AN DvG
BR-CJL, PE-BR(Rd),GF EC-Atrk, Oct24 VZ-BO,BR(Pa) CO-Sin, 700m May21 MX-PE,BR,AR
806 807 808
Strephonota ambrax Strephonota ambrax Strephonota azurinus

Jpe DvG DvG
CO-Sin, 700m Jul05 MX-PE,BR,AR PE-ARCC, MX-PE,BR,AR PE-Qtz, CR-PE

80
Tribu - Tribe
Eumaeini
809 810 811
Strephonota carteia ♂ Strephonota carteia ♀ Strephonota carteia ♀

DvG AN DvG
PE-ARCC, CO-PE,BR,GY EC-PmpL, CO-PE,BR,GY PE-ARCC, CO-PE,BR,GY
812 813 814
Strephonota carteia Strephonota cyllarissus Strephonota cyllarissus

DvG KK KK
PE-ARCC, CO-PE,BR,GY PE-Qtz, 1050m Nov22 CO-PE,BR,SU CO-MMt, 200m Oct15 CO-PE,BR,SU
815 816 817
Strephonota cyllarissus Strephonota cyllarissus Strephonota elika

GC AN DvG
BR-CJL, CO-PE,BR,SU PE-VCm, CO-PE,BR,SU BR-PNI, Feb03 PE-BR,AR
818 819 820
Strephonota foyi Strephonota foyi Strephonota foyi

DvG DvG KK
PE-, PE-BO,BR PE-Jgu, PE-BO,BR PE-Rmp-Mq, PE-BO,BR

81
Tribu - Tribe
Eumaeini
821 822 823
Strephonota foyi Strephonota jactator Strephonota metanira

DvG JAR Exorbaetta DvG
PE-ARCC, PE-BO,BR BR-, PE-BO,BR,PY,AR PE-Qtz, CO-BO,BR(Am)
824 825 826
Strephonota metanira Strephonota parvipuncta Strephonota parvipuncta

Exorbaetta DvG Robustana DvG Robustana DvG
PE-Qtz, CO-BO,BR(Am) PE-VCm, CO-PE,BR,GF PE-ARCC, CO-PE,BR,GF
827 828 829
Strephonota porphyritis ♀ Strephonota porphyritis Strephonota porphyritis

Serratonotes DvG Serratonotes DvG Serratonotes SDM
PE-ARCC, PE-BR(Pa),GY GY-, PE-BR(Pa),GY BR-, PE-BR(Pa),GY
830 831 832
Strephonota prina Strephonota sphinx Strephonota sphinx

DvG DvG DvG
PE-ARCC, CO-PE PE-Qtz, Nov26 PN-AR AR-ChM, Jan20 PN-AR

82
Tribu - Tribe
Eumaeini
833 834 835
Strephonota sphinx Strephonota sphinx Strephonota strephon

KP FM DvG
PN-Vant, Mar07 PN-AR CO-, May31 PN-AR PE-ARCC, CO-PE,BR,Guianas
836 837 838
Strephonota strephon Strephonota strephon Strephonota strephon

DvG DvG DvG
PE-ARCC, CO-PE,BR,Guianas PE-ARCC, CO-PE,BR,Guianas PE-MWC, CO-PE,BR,Guianas
839 840 841
Strephonota syedra Strephonota syedra Strephonota syedra

Syedranota DvG Syedranota DvG Syedranota GC
PE-, MX-PE,BR,GF GY-, MX-PE,BR,GF BR-CJL, MX-PE,BR,GF
842 843 844
Strephonota tephraeus Strephonota tephraeus Strephonota tephraeus

DvG
CO-VCC, 1650m Ene24 US(Tx)-PE,BR,AR CO-Sin, 780m Ago16 US(Tx)-PE,BR,AR EC-Atrk, US(Tx)-PE,BR,AR

83
Tribu - Tribe
Eumaeini
845 846 847
Strephonota tephraeus Strephonota tephraeus Strephonota tephraeus

KK DvG AN
PE-Qtz, US(Tx)-PE,BR,AR PE-ARCC, US(Tx)-PE,BR,AR PE-PntL, US(Tx)-PE,BR,AR
848 849 850
Strephonota trebonia Strephonota tyriam Terenthina terentia ♂

Treboniana LC Diminutina RH DvG
EC-Yach, Apr11 EC-PE BR-CJL, PE-BR(Pa),GF EC-Atrk, CR-PE,BR,GF
851 852 853
Terenthina terentia ♂ Terenthina terentia ♀ Terenthina terentia ♀

FM
CO-RP, 560m Nov07 CR-PE,BR,GF CO-CRC, 320m CR-PE,BR,GF CO-CRC, 320m CR-PE,BR,GF
854 855 856
Terenthina terentia ♀ Theclopsis gargara Theclopsis gargara

DvG DvG DvG
EC-Atrk, CR-PE,BR,GF PE-Pro, CO-PE,BR,VZ,GF PE-MWC, CO-PE,BR,VZ,GF

84
Tribu - Tribe
Eumaeini
857 858 859
Theclopsis gargara Theclopsis gargara Theclopsis lydus ♂

DvG DvG DvG
EC-Orq, CO-PE,BR,VZ,GF PE-Jgu, CO-PE,BR,VZ,GF PE-PntL, VZ-BO,BR,GY,GF
860 861 862
Theclopsis lydus ♀ Theclopsis lydus ♀ Theclopsis lydus ♀

DvG DvG
EC-Cpl, VZ-BO,BR,GY,GF CO-RSD, VZ-BO,BR,GY,GF CO-Bvt, 200m VZ-BO,BR,GY,GF
863 864 865
Theclopsis lydus Theclopsis lydus Theclopsis lydus

MM FM AN
CO-Gvr, Ene13 VZ-BO,BR,GY,GF CO-RP, 560m Ago01 VZ-BO,BR,GY,GF EC-Atrk, VZ-BO,BR,GY,GF
866 867 868
Theclopsis lydus Theclopsis lydus Tmolus cydrara

AN KNc DvG
EC-Atrk, Oct30 VZ-BO,BR,GY,GF PE-PNM, VZ-BO,BR,GY,GF PE-ARCC, MX-BO,S BR,AR,GF

85
Tribu - Tribe
Eumaeini
869 870 871
Tmolus cydrara Tmolus cydrara Tmolus cydrara

DvG FM KK
PE-ARCC, MX-BO,S BR,AR,GF CO-RP, 560m Ago01 MX-BO,S BR,AR,GF CO-MMt, 200m Oct20 MX-BO,S BR,AR,GF
872 873 874
Tmolus echion Tmolus echion Tmolus echion

DvG
CO-Mzm, MX-BO,S BR,AR,GF CO-PSJ, 1850m Oct17 MX-BO,S BR,AR,GF CO-Mor, 80m May01 MX-BO,S BR,AR,GF
875 876 877
CO-Sin, 780m Sep06 MX-BO,S BR,AR,GF CO-PSJ, 1800m Feb22 MX-BO,S BR,AR,GF CO-VCC, 1650m Ene25 MX-BO,S BR,AR,GF
878 879 880
CO-PSJ, 1850m Oct17 MX-BO,S BR,AR,GF CO-PSJ, 1850m Oct17 MX-BO,S BR,AR,GF CO-GMq, 5m MX-BO,S BR,AR,GF

86
Tribu - Tribe
Eumaeini
881 882 883
Tmolus echion Tmolus echion Tmolus mutina

DvG
CO-GMq, 5m Dic29 MX-BO,S BR,AR,GF CO-HLB, 524m - MX-BO,S BR,AR,GF PE-ARCC, CR-PE,N BR,GY,GF
884 885 886
Tmolus mutina Tmolus mutina Tmolus ufentina

DvG GC DvG
PE-ARCC, CR-PE,N BR,GY,GF BR-, CR-PE,N BR,GY,GF PE-Qtz, Nov21 CO-BO,BR(Am),GF
887
Tmolus ufentina
DvG
PE-Qtz, Nov21 CO-BO,BR(Am),GF
888
Subtribu - Subtribe
Trichonidina
Trichonis immaculata ♂
GC
BR-CJL, EC-PE,BR,GF

87
Tribu - Tribe
Eumaeini
889 890
Subtribu - Subtribe
Callophryidina
Cyanophrys amyntor Cyanophrys amyntor
CO-Sin, 700m Jun14 MX-PE,BR CO-HLB, 524m - MX-PE,BR
891 892 893
Cyanophrys amyntor Cyanophrys amyntor Cyanophrys amyntor

DvG DvG DvG
PE-ARCC, MX-PE,BR PE-ARCC, MX-PE,BR PE-ARCC, MX-PE,BR
894 895 896
Cyanophrys amyntor Cyanophrys amyntor Cyanophrys amyntor
CO-SJn, 800m Sep12 MX-PE,BR CO-Jard, MX-PE,BR CO-HLB, 524m - MX-PE,BR
897 898 899
Cyanophrys amyntor Cyanophrys banosensis Cyanophrys herodotus

KG
CO-HLB, 524m - MX-PE,BR PE-BSht, EC-PE CO-CRC, 320m Abr05 MX-AR

88
Tribu - Tribe
Eumaeini
900 901 902
Cyanophrys herodotus Cyanophrys herodotus Cyanophrys herodotus

THn DvG
CO-, MX-AR CO-CRC, 320m MX-AR EC-Podc, MX-AR
903 904 905
Cyanophrys pseudolongula Cyanophrys pseudolongula Cyanophrys pseudolongula
CO-PSJ, 1800m Dic05 CO-BO,VZ,AR CO-PSJ, 1800m CO-BO,VZ,AR CO-PSJ, 1800m CO-BO,VZ,AR
906 907 908
Cyanophrys pseudolongula Chalybs hassan Chalybs hassan

DvG DvG
CO-PSJ, 1800m CO-BO,VZ,AR EC-Orq, MX-PE,BR,GF,AR PE-ARCC, MX-PE,BR,GF,AR
909 910 911
Chalybs hassan Chalybs hassan Chalybs janias

KK AN DvG
PE-Qtz, 1050m MX-PE,BR,GF,AR PE-, MX-PE,BR,GF,AR EC-Cpl, MX-PE,BR,GF

89
Tribu - Tribe
Eumaeini
912 913 914
Chalybs janias Chalybs janias Chalybs janias

GC FM DvG
BR-CJL, MX-PE,BR,GF CO-RP, 560m Ago23 MX-PE,BR,GF CO-Mzm, MX-PE,BR,GF
915 916 917
Chlorostrymon simaethis Chlorostrymon simaethis Chlorostrymon telea

KK
CO-Gai, 582m US(Tx)-BR,AR CO-, - US(Tx)-BR,AR PE-, US(Tx)-BO,BR,AR
918 919 920
Chlorostrymon telea Chlorostrymon telea Erora badeta

DvG DvG
PE-ARCC, US(Tx)-BO,BR,AR PE-ARCC, US(Tx)-BO,BR,AR CO-RSm, 400m Ago02 CR-PE,BR,AR
921 922 923
Erora badeta Erora badeta Erora badeta

AN
CO-RSm, 400m Ago02 CR-PE,BR,AR CO-QLH, 750m Jul25 CR-PE,BR,AR PE-, CR-PE,BR,AR

90
Tribu - Tribe
Eumaeini
924 925 926
Erora badeta Erora biblia Erora caespes ♀

KK MCC THn
EC-Atrk, CR-PE,BR,AR BR-, Ago13 VZ-BO,BR(Pa),PY CO-CRC, 320m CO?EC-BO
927 928 929
Erora caespes Erora carla ♀ Erora carla ♀

AN
PE-, CO?EC-BO CO-PSJ, 1800m MX-BR CO-PSJ, 1800m MX-BR
930 931 932
Erora carla Erora carla Erora carla

RRt
CO-Sin, 780m Ago16 MX-BR CO-Mzm, MX-BR CO-PSJ, 1800m Nov21 MX-BR
933 934 935

DvG DvG
CO-PSJ, 1800m Nov21 MX-BR CO-Mzm, 1350m MX-BR CO-Mzm, 1350m MX-BR

91
Tribu - Tribe
Eumaeini
936 937 938

RRt
CO-PSJ, 1800m Nov21 MX-BR CO-Mzm, MX-BR CO-PSJ, 1800m Ago13 MX-BR
939 940 941
Erora carla Erora carla Erora gabina

DvG
CO-Sin, 780m Ago16 MX-BR CO-, MX-BR PE-Qtz, MX-BO,BR,AR
942 943 944
Erora gabina Erora nitetis Erora phrosine

DvG BBn DvG
PE-Qtz, MX-BO,BR,AR MX-Oax, MX-PE PE-Qtz, EC-BO
945 946 947
Erora phrosine Ocaria aholiba Ocaria aholiba

KG KG DvG
PE-PNYC, EC-BO CO-RlT, CR-BO,VZ PE-Rctl, CR-BO,VZ

92
Tribu - Tribe
Eumaeini
948 949 950
Ocaria aholiba Ocaria aholiba Ocaria aholiba

AN DvG DvG
EC-Cpl, CR-BO,VZ EC-BBR, CR-BO,VZ CO-Mzm, 1600m CR-BO,VZ
951 952 953
Ocaria aholiba Ocaria aholiba Ocaria arpoxais

DvG DvG DvG
CO-RlT, CR-BO,VZ PE-CoR, CR-BO,VZ CO-Mzm, MX-BO
954 955 956
Ocaria calesia Ocaria calesia Ocaria calesia

DvG
CO-Mzm, 2050m CO-PE,AR CO-RlR, 1850m - CO-PE,AR CO-PSJ, 1800m CO-PE,AR
957 958 959
Ocaria calesia Ocaria calesia Ocaria calesia

DvG DvG DvG
CO-Mzm, Jun12 CO-PE,AR PE-Rctl, CO-PE,AR EC-WS, CO-PE,AR

93
Tribu - Tribe
Eumaeini
960 961 962
Ocaria cinerea Ocaria cinerea Ocaria cinerea

DvG DvG DvG
PE-Qtz, EC,PE-BR(RS) EC-Atrk, EC,PE-BR(RS) PE-Jgu, EC,PE-BR(RS)
963 964 965
Ocaria clepsydra Ocaria elongata Ocaria elongata

DvG KK BBt
CO-Mzm, 1600m Jun24 CO-PE,VZ PE-CoR, CO-BO EC-Baz, CO-BO
966 967 968
Ocaria elongata Ocaria elongata Ocaria elongata

Leo
CO-Mzm, CO-BO CO-Ang, 1900m CO-BO CO-Ang, 1900m CO-BO
969 970 971
Ocaria ocrisia Ocaria ocrisia Ocaria ocrisia

DvG
CO-, - MX-AR CO-VPN, 1450m MX-AR EC-JSR, MX-AR

94
Tribu - Tribe
Eumaeini
972 973 974
CO-, MX-AR CO-, MX-AR CO-Sin, 780m Nov01 MX-AR
975 976 977
CO-, - MX-AR CO-Ang, 1900m MX-AR CO-, - MX-AR
978 979 980

FM
CO-RP, 560m Nov07 MX-AR CO-QEV, 1975m MX-AR CO-QEV, 1975m MX-AR
981 982 983

DvG DvG DvG
EC-Cpl, MX-AR PE-ARCC, MX-AR PE-ARCC, MX-AR

95
Tribu - Tribe
Eumaeini
984 985 986
Ocaria ocrisia Ocaria ocrisia Ocaria sadei

DvG
CO-Sin, 780m Nov01 MX-AR CO-RlR, 1850m - MX-AR PE-Rmpt, CO-BO
987 988 989
Ocaria sadei Ocaria sadei Ocaria thales

OE KG GC
CO-, Feb06 CO-BO PE-Rmp-Mq, CO-BO BR-CJL, Ago20 MX-AR
990 991 992
Ocaria thales Ocaria thales Ocaria thales

DW KG
PA-PplRd, MX-AR PE-PNYC, 1100m MX-AR CO-QLH, 750m Jul25 MX-AR
993 994 995
Ocaria thales Semonina ares Semonina ares

DvG DvG DvG
PE-PNYC, 1000m MX-AR PE-Jgu, PN-PE,BR,GF PE-Jgu, PN-PE,BR,GF

96
Tribu - Tribe
Eumaeini
996 997 998
Thaeides muela Thaeides theia ♀ Thaeides theia

DvG DvG
PE-Wyq, EC-BO,CH CO-RAD, May17 MX-BO,BR,AR CO-RlT, Ene06 MX-BO,BR,AR
999
Thaeides theia
KNc
PE-PNM, MX-BO,BR,AR
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Esta versión del catálogo se generó el lunes, 9 de mayo de 2022 Date of report
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97
98
Lycaenidae
Subfamilia Polyommatinae
Tribu - Tribe
Polyommatini
1 2
Subtribu - Subtribe
Sin Subtribu
Hemiargus hanno hanno ♂ Hemiargus hanno hanno ♀♂
CO-, - CO-AR CO-VCC, 1650m Oct11 CO-AR
3 4 5
Hemiargus hanno hanno ♀ Hemiargus hanno hanno Hemiargus hanno hanno
CO-VCC, 1650m Oct11 CO-AR CO-VCC, 1650m Oct11 CO-AR CO-VCC, 1650m Oct11 CO-AR
6 7 8
Hemiargus hanno hanno Hemiargus hanno hanno Hemiargus hanno hanno
CO-VCC, 1650m Oct11 CO-AR CO-PSJ, 1800m Feb07 CO-AR CO-, CO-AR
BioButterfly DataBase - Butterflycatalogs.com Perú Polyommatinae

99
Tribu - Tribe
Polyommatini
9 10 11
CO-VCC, 1650m Oct11 CO-AR CO-PSJ, 1800m Feb07 CO-AR CO-, CO-AR
12 13 14
CO-, - CO-AR CO-VCC, 1650m Oct11 CO-AR CO-VCC, 1650m Oct11 CO-AR
15 16 17
Hemiargus hanno hanno Hemiargus hanno hanno Hemiargus ramon

DvG
CO-, - CO-AR CO-, - CO-AR EC-Vilc, EC-PE
18 19 20
Hemiargus ramon Hemiargus ramon Hemiargus ramon

DvG DvG DvG
PE-Lim, EC-PE EC-Vilc, EC-PE EC-Vilc, EC-PE
BioButterfly DataBase - Butterflycatalogs.com Perú Polyommatinae Página 2 de 5

100
Tribu - Tribe
Polyommatini
21 22 23
Itylos pacis Itylos pacis Leptotes bathyllos

Madeleinea AN Madeleinea AN KK
PE-, PE-BO PE-, PE-BO EC-, Oct25 EC-PE
24 25 26
Leptotes bathyllos Leptotes callanga Leptotes callanga

KK AN DvG
EC-Ten, EC-PE PE-, PE-BO PE-Chtk, PE-BO
27 28 29
Leptotes callanga Leptotes callanga Leptotes cassius

DvG AN
PE-, PE-BO EC-PaP, PE-BO CO-, - US-VZ,BO,BR,AR,SU
30 31 32
Leptotes cassius Leptotes cassius Leptotes cassius
CO-, - US-VZ,BO,BR,AR,SU CO-GMq, 5m US-VZ,BO,BR,AR,SU CO-, US-VZ,BO,BR,AR,SU

101
Tribu - Tribe
Polyommatini
33 34 35
Leptotes cassius Leptotes cassius Leptotes lamasi

DvG AN
CO-VPN, 1450m US-VZ,BO,BR,AR,SU CO-Mzm, 1450m US-VZ,BO,BR,AR,SU PE-, EC-PE
36 37 38
Leptotes lamasi Nabokovia cuzquenha Nabokovia cuzquenha

DvG DvG DvG
EC-Vilc, EC-PE PE-, 3100m Jul16 PE PE-, 3100m Jul16 PE
39 40 41
Zizula cyna Zizula cyna Zizula cyna
CO-, - US-AR CO-, US-AR CO-Sin, 780m Jun14 US-AR
42
Zizula cyna
Leo
CO-Mzm, Jun05 US-AR

102
Tribu - Tribe
Polyommatini
infGUIDE Pais SUBFAMILIA Photo Guide
Esta versión del catálogo se generó el lunes, 9 de mayo de 2022 Date of report
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103
Mariposas de Perú familia Lycaenidae
Neotropical butterflies, family Lycaenidae
Nota: especies señaladas en la casilla tienen foto en el catálogo. Agradecemos colaboracion para completar las fotos que nos faltan
Note: species with a checked box have a photo in the catalog. We appreciate collaboration to complete the missing photos.
Lycaenidae - Perú
Subfamilia Tribu
Eumaeini
Theclinae
Subtribu
Tribu
Rhammina
Eumaeini
Balintus 7
Subtribu
15 Balintus tityrus (C. Felder & R. Felder,
Eumaeina 1865)
Brevianta 1 Johnsonita 8
1 Brevianta ematheon (Cramer, 1777) 16 Johnsonita auda (Hewitson, 1867)

2 Brevianta undulata (Hewitson, 1867) 17 Johnsonita catadupa (Hewitson, 1869)
3 Brevianta undulella (Strand, 1918) 18 Johnsonita chlamydem (H. Druce, 1907)
Eumaeus 2 19 Johnsonita iacinta Bálint, et al, 2021
4 Eumaeus minyas (Hübner, [1809]) 20 Johnsonita johnbanksi Bálint, 2003
5 Eumaeus toxana (Boisduval, 1870) 21 Johnsonita oxalita Bálint, et al, 2021
Micandra 3 22 Johnsonita pardoa (D'Abrera, 1995)
6 Micandra aegides (C. Felder & R. Felder, 23 Johnsonita subcunicula Bálint, et al, 2021
1865) 24 Johnsonita zubkova Bálint, et al, 2021
7 Micandra comae (H. Druce, 1907) Lathecla 9
8 Micandra dignota (Draudt, 1919) Lathecla latagus (Godman & Salvin, 1887)
25
9 Micandra ion (Druce, 1890)

26 Lathecla mimula (Draudt, 1920)
10 Micandra platyptera (C. Felder & R. Felder, Podanotum 10
1865)
27 Podanotum glorissimum Bálint &
Mithras 4
Wojtusiak, 2002
11 Mithras nautes (Cramer, 1779) Podanotum wayquecha Torres & Johnson, Lamprospilus
28
Theorema 5 1996
12 Theorema dysmenia Draudt, 1919 Rhamma 11
Thestius 6 29 Rhamma adriana (K. Johnson, 1992)
13 Thestius epopea (Hewitson, 1870) 30 Rhamma arria (Hewitson, 1870)
14 Thestius meridionalis (Draudt, 1920) 31 Rhamma aurugo (Draudt, 1919)
ButterflyBase-Butterflycatalogs.com Página 1
104
Lycaenidae - Perú
32 Rhamma commodus (C. Felder & R. Felder, 54 Paraspiculatus sine Busby & Robbins, 2017
1865)
Penaincisalia 16
33 Rhamma comstocki K. Johnson, 1992
55 Penaincisalia alatus (H. Druce, 1907)
34 Rhamma familiaris (K. Johnson, 1991) Paralustrus
56 Penaincisalia albalineata (K. Johnson,
35 Rhamma hybla (Druce, 1907) Shapiroana 1992)
36 Rhamma inexpectata K. Johnson, 1992 57 Penaincisalia amatista (Dognin, 1895) Abloxurina
37 Rhamma livida Torres & K. Johnson, 1997 58 Penaincisalia amazona (Bálint &
Wojtusiak, 2003)
38 Rhamma magenta K. Johnson, 1992
59 Penaincisalia atymna (Hewitson, 1870)
39 Rhamma nigrasarotina K. Johnson, 1992
60 Penaincisalia aurulenta K. Johnson, 1990
40 Rhamma tarma K. Johnson, 1992
Salazaria 61 Penaincisalia biophot Bálint & Wojtusaik,
12
2008
41 Salazaria sala (Hewitson, 1867)
62 Penaincisalia caudata K. Johnson, 1990
Tribu
Eumaeini 63 Penaincisalia culminicola (Staudinger,
1894)
Subtribu 64 Penaincisalia chachapoya (Bálint &
Timaetina Wojtusiak, 2003)
65 Penaincisalia descimoni K. Johnson, 1990
Marachina 13
66 Penaincisalia dominiquae (Bálint, Boyer &
42 Marachina maraches (H. Druce, 1912)
Prieto, 2007)
Nesiostrymon 14
67 Penaincisalia duviolsi (Bálint, Boyer & Abloxurina
43 Nesiostrymon calchinia (Hewitson, 1868) Wojtusaik, 2006)
44 Nesiostrymon celona (Hewitson, 1874) 68 Penaincisalia falcatamba (Bálint, Boyer & Abloxurina
Wojtusaik, 2006)
Paraspiculatus 15
69 Penaincisalia felizitas Bálint, 2004
45 Paraspiculatus azul Busby, Robbins, &
Faynel, 2017 70 Penaincisalia ianusi (Bálint, 2011) Ianusanta
46 Paraspiculatus colombiensis (K. Johnson & Mithras 71 Penaincisalia lamasi Bálint, 2001
Constantino, 1997) 72 Penaincisalia libertada J. Hall, 2005
47 Paraspiculatus cosmo Busby, Robbins, & 73 Penaincisalia loxurina (C. Felder & R. Thecloxurina
Faynel, 2017 Felder, 1865)
48 Paraspiculatus elis (Cramer, 1779) Mithras
74 Penaincisalia magnifica (K. Johnson, 1992) Thecloxurina
49 Paraspiculatus emma Busby & Robbins, 75 Penaincisalia molipampa (Bálint, Boyer & Abloxurina
2017 Wojtusaik, 2006)
50 Paraspiculatus oroanna Bálint, 2002 76 Penaincisalia oxapampa (Bálint, Boyer & Abloxurina
51 Paraspiculatus orobia (Hewitson, 1867) Mithras Wojtusaik, 2006)
52 Paraspiculatus orobiana Hewitson, 1867 Mithras 77 Penaincisalia perezi Bálint, 2001
53 Paraspiculatus orocana (H. Druce, 1912) Mithras 78 Penaincisalia purpurea (K. Johnson, 1992) Pons
105
Lycaenidae - Perú
79 Penaincisalia rawlinsi K. Johnson, 1990 100 Arcas jivaro Nicolay, 1971
80 Penaincisalia tegulina (Bálint & Boyer, 101 Arcas tuneta (Hewitson, 1865)
2008)
Atlides 20
Phothecla 17
102 Atlides atys (Cramer, 1779)
81 Phothecla margarita (Draudt, 1920)
103 Atlides bacis (Godman & Salvin, 1887)
82 Phothecla photismos (H. Druce, 1907)
104 Atlides carpophora (Hewitson, 1868)
83 Phothecla thespia (Hewitson, 1870)
105 Atlides halljasoni Bálint, Kertész &
Timaeta 18 Wojtusiak, 2006
84 Timaeta aepea (Hewitson, 1874) 106 Atlides havila (Hewitson, 1865)
85 Timaeta bennetti (Dyar, 1913) Temecla 107 Atlides inachus (Cramer, 1775)
86 Timaeta boyeropampa (Bálint & 108 Atlides polybe (Linnaeus, 1763)

Wojtusiak, 2010)
109 Atlides rustan (Stoll, 1790)
87 Timaeta cospata Robbins & R. Busby, 2008
Brangas 21
88 Timaeta christina Robbins & R. Busby,
110 Brangas caranus (Stoll, 1780)
2008
111 Brangas coccineifrons (Godman & Salvin,
89 Timaeta gabriela Robbins & R. Busby,
1887)
2008
112 Brangas dydimaon (Cramer, 1777)
90 Timaeta heraclides (Godman & Salvin, Temecla
1887) 113 Brangas felderi (Goodson, 1945)
91 Timaeta molinopampa (Bálint & Jagiello 114 Brangas getus (Fabricius, 1787)
Wojtusiak, 2000)
115 Brangas insolitus Bálint & Faynel, 2008
92 Timaeta pilosa Robbins & R. Busby, 2008
116 Brangas neildonatus Bálint & Faynel, 2008
93 Timaeta roberti Robbins & R. Busby, 2008 Trochusinus
117 Brangas neora (Hewitson, 1867)
94 Timaeta sergius (Godman & Salvin, 1887) Temecla
118 Brangas polonus Bálint, 2008
95 Timaeta tema (Hewitson, 1867) Temecla
119 Brangas teucria (Hewitson, 1868) Dabreras
96 Timaeta timaeus (C. Felder & R. Felder,
1865) 120 Brangas torfrida (Hewitson, 1867)
Denivia 22
97 Timaeta trochus (H. Druce, 1907)
Tribu 121 Denivia adamsi H. Druce, 1909
Eumaeini 122 Denivia arene Goodson, 1945
Subtribu 123 Denivia boliboyerus Bálint & Wojtusiak, Lucilda

2006
Atlidina
124 Denivia dabreus Bálint, 2002 Lucilda
Arcas 19
125 Denivia danaus (C. Felder & R. Felder, Theritas/Lucilda
98 Arcas alleluia Bálint, 2002 1865)
99 Arcas imperialis (Cramer, 1775) 126 Denivia hemon (Cramer, 1775) Theritas
106
Lycaenidae - Perú
127 Denivia lisus (Stoll, 1790) Theritas 147 Allosmaitia myrtusa (Hewitson, 1867)
128 Denivia monica Hewitson, 1867 148 Allosmaitia strophius (Godart, [1824])
129 Denivia orsina (Hewitson, 1877) Aveexcrenota 27
130 Denivia phegeus (Hewitson, 1865) Theritas

149 Aveexcrenota anna (H. Druce, 1907) Theritas
131 Denivia silma Martins, Faynel, & Robbins, Contrafacia 28

2016
150 Contrafacia imma (Prittwitz, 1865)
132 Denivia viresco (H. Druce, 1907) Theritas
Enos 29
Pseudolycaena 23
151 Enos falerina (Hewitson, 1867)
133 Pseudolycaena marsyas (Linnaeus, 1758)
152 Enos myrtea (Hewitson, 1867)
Theritas 24
153 Enos polka Lamas & Robbins, 2009
134 Theritas harrietta A.G. Weeks, 1901
154 Enos thara (Hewitson, 1867)
135 Theritas mavors Hübner, 1818
Janthecla 30
136 Theritas paupera (C. Felder & R. Felder,
1865) 155 Janthecla adrienne Robbins, 2009
Tribu 156 Janthecla leea Venables & Robbins, 1991
Eumaeini Janthecla malvina (Hewitson, 1867)
157
Subtribu 158 Janthecla rocena (Hewitson, 1867)

Evenina 159 Janthecla sista (Hewitson, 1867)
Evenus 25 Laothus 31
137 Evenus batesii (Hewitson, 1865) 160 Laothus gibberosa (Hewitson, 1867) Gibbosa
138 Evenus candidus (H. Druce, 1907) 161 Laothus viridicans (C. Felder & R. Felder,
1865)
139 Evenus coronata (Hewitson, 1865)
Tribu
140 Evenus felix Neild & Bálint, 2014 Eumaeini
141 Evenus floralia (H. Druce, 1907)
Subtribu
142 Evenus gabriela (Cramer, 1775)
Paiwarriina
143 Evenus regalis (Cramer, 1775)
Annamaria 32
144 Evenus satyroides (Hewitson, 1865)
162 Annamaria ganimedes (Cramer, 1775) Lamasina
145 Evenus tagyra (Hewitson, 1865)
Tribu 163 Annamaria lathyi (Bálint, 2005) Lamasina
Eumaeini 164 Annamaria rhaptissima (K. Johnson, 1991) Airamanna

Fasslantonius 33
Subtribu
Jantheclina 165 Fasslantonius episcopalis (Fassl, 1912) Paiwarria
Allosmaitia Kolana 34
26
Allosmaitia fidena (Hewitson, 1867) 166 Kolana ergina (Hewitson, 1867)

146
107
Lycaenidae - Perú
167 Kolana ligurina (Hewitson, 1874) 184 Michaelus phoenissa (Hewitson, 1867)
168 Kolana lyde (Godman & Salvin, 1887) 185 Michaelus thordesa (Hewitson, 1867)
Paiwarria 35 Oenomaus 42
169 Paiwarria telemus (Cramer, 1775) 186 Oenomaus andi R. Busby & Faynel, 2012
170 Paiwarria venulius (Cramer, 1779) 187 Oenomaus atena (Hewitson, 1867)
Tribu Oenomaus atesa (Hewitson, 1867)
188
Eumaeini
189 Oenomaus barba (H. Druce, 1907) Porthecla
Subtribu 190 Oenomaus curiosa Faynel & Moser, 2008

Cupatheclina 191 Oenomaus gemma (H. Druce, 1907) Porthecla
Cupathecla 36 192 Oenomaus jauffreti Faynel & Moser, 2008

171 Cupathecla cupentus (Stoll, 1781) Megathecla 193 Oenomaus lea Faynel & Robbins, 2012
Tribu Oenomaus mentirosa Faynel & Robbins,
Eumaeini
194
2012
195 Oenomaus minyia (Hewitson, 1867) Porthecla
Subtribu
Parrhasiina 196 Oenomaus nigra Faynel & Moser, 2008
Apuecla 37
197 Oenomaus ortygnus (Cramer, 1779)
198 Oenomaus peruensis Faynel & Moser, Porthecla
172 Apuecla picus (H. Druce, 1907)
2011
Beatheclus 38
199 Oenomaus taua Faynel & Moser, 2008
173 Beatheclus beatrizae Bálint & Dahners, Thepytus
Oenomaus wilmotti R. Busby, Faynel & Porthecla
200
2006 Moser, 2011
Dicya 39
Olynthus 43
174 Dicya carnica (Hewitson, 1873) Olynthus essus (Herrich-Schäffer, [1853])
201
175 Dicya dicaea (Hewitson, 1874) Olynthus lorea (Möschler, 1883)

202
176 Dicya iambe (Godman & Salvin, 1887) Olynthus narbal (Stoll, 1790)
203
Ignata 40
204 Olynthus obsoleta (Lathy, 1926)
177 Ignata caldas Robbins, 2010 205 Olynthus ophelia (Hewitson, 1867)
178 Ignata levis (H. Druce, 1907) Olynthus porphyreticus (H. Druce, 1907)
206
179 Ignata mulsus (H. Druce, 1907) Parrhasius 44

180 Ignata norax (Godman & Salvin, 1887) Parrhasius orgia (Hewitson, 1867)
207
Michaelus 41
Parrhasius polibetes (Stoll, 1781)
208
181 Michaelus ira (Hewitson, 1867) Parrhasius selika (Hewitson, 1874)

209
182 Michaelus jebus (Godart, [1824]) Symbiopsis 45

183 Michaelus joseph Robbins, 2010 Symbiopsis aprica (Möschler, 1883)
210
108
Lycaenidae - Perú
211 Symbiopsis perulera Robbins, 2004 230 Calycopis anthora (Hewitson, 1877)
212 Symbiopsis pupilla (Draudt, 1920) 231 Calycopis atnius (Herrich-Schäffer, [1853])
Thepytus 46 232 Calycopis bactra (Hewitson, 1877)
213 Thepytus echelta (Hewitson, 1867) 233 Calycopis barza (W. D. Field, 1967)
214 Thepytus jennifer R. Busby & Robbins, 234 Calycopis bellera (Hewitson, 1877)
2010
235 Calycopis boliviensis (K. Johnson, 1991)
215 Thepytus thyrea (Hewitson, 1867)
236 Calycopis buphonia (Hewitson, 1868)
Tribu
Eumaeini 237 Calycopis caesaries (H. Druce, 1907)
238 Calycopis calus (Godart, [1824])
Subtribu
239 Calycopis caulonia (Hewitson, 1877)
Ipideclina
240 Calycopis centoripa (Hewitson, 1868)
Ipidecla 47
241 Calycopis cerata (Hewitson, 1877)
216 Ipidecla crepundia (H. Druce, 1909)
242 Calycopis cicero Robbins & Duarte, 2005
217 Ipidecla schausi (Godman & Salvin, 1887)
243 Calycopis cyanus (Draudt, 1920)
Tribu
Eumaeini 244 Calycopis demonassa (Hewitson, 1868)
245 Calycopis fractunda W. D. Field, 1967
Subtribu
246 Calycopis gentilla (Schaus, 1902)
Calycopidina
247 Calycopis gizela (Hewitson, 1877)
Arzecla 48
248 Calycopis johnsoni (Salazar, 2000)
218 Arzecla albolineata (Lathy, 1936)
249 Calycopis lerbela W.D. Field, 1967
219 Arzecla arza (Hewitson, 1874)
250 Calycopis malta (Schaus, 1902)
220 Arzecla calatia (Hewitson, 1873)
251 Calycopis mimas (Godman & Salvin, 1887)
221 Arzecla canacha (Hewitson, 1877)
252 Calycopis naka (W. D. Field, 1967)
222 Arzecla paralus (Godman & Salvin, 1887)
253 Calycopis nicolayi W. D. Field, 1967
223 Arzecla straboris Bálint, 2019
254 Calycopis orcilla (Hewitson, 1874)
224 Arzecla taminella (Schaus, 1902)
255 Calycopis orcillula (Strand, 1916)
225 Arzecla tucumanensis (K. Johnson &
256 Calycopis origo (Godman & Salvin, 1887)
Kroenlein, 1993)
257 Calycopis partunda (Hewitson, 1877)
Badecla 49
258 Calycopis petaurister (H. Druce, 1907)
226 Badecla badaca (Hewitson, 1868) Lamprospilus
259 Calycopis pisis (Godman & Salvin, 1887)
227 Badecla lanckena (Schaus, 1902)
260 Calycopis puppius (Godman & Salvin,
Calycopis 50 1887)
228 Calycopis anastasia W. D. Field, 1967 261 Calycopis spadectis (K. Johnson &
229 Calycopis anfracta (H. Druce, 1907) Kroenlein, 1993)
109
Lycaenidae - Perú
262 Calycopis suda (Draudt, 1920) Rubroserrata 56
263 Calycopis tamos (Godman & Salvin, 1887) 288 Rubroserrata ecbatana (Hewitson, 1868)
264 Calycopis tifla (W. D. Field, 1967) Ziegleria 57
265 Calycopis trebula (Hewitson, 1868) 289 Ziegleria ceromia (Hewitson, 1877)
266 Calycopis vibulena (Hewitson, 1877) 290 Ziegleria hesperitis (A. Butler & H. Druce,
267 Calycopis vitruvia (Hewitson, 1877) 1872)
Tribu
Calycopis wolfii (K. Johnson, 1993)
268
Eumaeini
269 Calycopis xeneta (Hewitson, 1877)
Camissecla Subtribu
51
Strymonina
270 Camissecla camissa (Hewitson, 1870)
Arawacus 58
271 Camissecla cleocha (Hewitson, 1870)
291 Arawacus aetolus (Sulzer, 1776)
272 Camissecla pactya (Hewitson, 1874)
292 Arawacus binangula (Schaus, 1902)
273 Camissecla verbenaca (H. Druce, 1907)
293 Arawacus dolylas (Cramer, 1777)
274 Camissecla vesper (H. Druce, 1909)
294 Arawacus ellida (Hewitson, 1867)
Electrostrymon 52
295 Arawacus separata (Lathy, 1926)
275 Electrostrymon endymion (Fabricius, 1775)
Hypostrymon 59
276 Electrostrymon joya (Dognin, 1895)
296 Hypostrymon aepeona (Draudt, 1920)
Gigantorubra 53
297 Hypostrymon asa (Hewitson, 1868)
277 Gigantorubra aruma (Hewitson, 1877) Arumecla
298 Hypostrymon renidens (Draudt, 1920)
278 Gigantorubra calycopinotis (K. Johnson & Arumecla
Kroenlein, 1993) Panthiades 60
279 Gigantorubra netesca (Draudt, 1920) Arumecla 299 Panthiades aeolus (Fabricius, 1775)
280 Gigantorubra orcidia (Hewitson, 1874) Lamprospilus 300 Panthiades bitias (Cramer, 1777)
Kisutam 54 301 Panthiades paphlagon (C. Felder & R.
Felder, 1865)
281 Kisutam syllis (Godman & Salvin, 1887)
302 Panthiades phaleros (Linnaeus, 1767)
Lamprospilus 55
Rekoa 61
282 Lamprospilus aunus (Cramer, 1775)
303 Rekoa marius (Lucas, 1857)
283 Lamprospilus coelicolor (A. Butler & H.
Druce, 1872) 304 Rekoa meton (Cramer, 1779)
284 Lamprospilus decorata Lathy, 1926 305 Rekoa palegon (Cramer, 1780)
285 Lamprospilus draudti Lathy, 1932 306 Rekoa stagira (Hewitson, 1867)
286 Lamprospilus genius Geyer, 1832 Strymon 62
287 Lamprospilus nicetus C. Felder & R. Felder, 307 Strymon ahrenholzi Nicolay & Robbins,
1865 2005
110
Lycaenidae - Perú
308 Strymon astiocha (Prittwitz, 1865) Subtribu
Strephonotina
309 Strymon bazochii (Godart, [1824])
Strymon bubastus (Stoll, 1780) Aubergina 64
310
311 Strymon cestri (Reakirt, [1867]) 337 Aubergina alda (Hewitson, 1868)
312 Strymon crambusa (Hewitson, 1874) 338 Aubergina hesychia (Godman & Salvin,
1887)
313 Strymon cryptodes Balint et al, 2019
Celmia 65
314 Strymon daraba (Hewitson, 1867)
339 Celmia anastomosis (Draudt, [1918])
315 Strymon davara (Hewitson, 1868)
340 Celmia celmus (Cramer, 1775)
316 Strymon flavaria (Ureta, 1956)
341 Celmia color (H. Druce, 1907)
317 Strymon gabatha (Hewitson, 1870)
342 Celmia conoveria (Schaus, 1902)
318 Strymon heodes (H. Druce, 1909)
343 Celmia mecrida (Hewitson, 1867)
319 Strymon istapa (Reakirt, [1867])
Gargina 66
320 Strymon jacqueline Nicolay & Robbins,
2005 344 Gargina caninius (H. Druce, 1907)
321 Strymon lamasi Nicolay & Robbins, 2005 345 Gargina emessa (Hewitson, 1867)
322 Strymon lucena (Hewitson, 1868) 346 Gargina gargophia (Hewitson, 1877)
323 Strymon megarus (Godart, [1824]) 347 Gargina gnosia (Hewitson, 1868)
324 Strymon michelle Nicolay & Robbins, 2005 348 Gargina panchaea (Hewitson, 1869)
325 Strymon mulucha (Hewitson, 1867) 349 Gargina thyesta (Hewitson, 1869)
326 Strymon rufofusca (Hewitson, 1877) Gossenia 67
327 Strymon yojoa (Reakirt, [1867]) 350 Gossenia lycabas (Cramer, 1777) Thestius
328 Strymon ziba (Hewitson, 1868) Iaspis 68
Thereus 63 351 Iaspis castitas (H. Druce, 1907)

329 Thereus brocki Robbins, Heredia & Busby, 352 Iaspis temesa (Hewitson, 1868)
2015
Ministrymon 69
330 Thereus cithonius (Godart, [1824]) Noreena
353 Ministrymon arthuri Bálint, K. Johnson &
331 Thereus columbicola (Strand, 1916) Austin, [1999]
332 Thereus endera (Hewitson, 1867) 354 Ministrymon azia (Hewitson, 1873)
333 Thereus gabathana (Strand, 1918) 355 Ministrymon cleon (Fabricius, 1775)
334 Thereus ortalus (Godman & Salvin, 1887) 356 Ministrymon cruenta (Gosse, 1880)
335 Thereus pedusa (Hewitson, 1867) Pedusa
357 Ministrymon megacles (Stoll, 1780)
336 Thereus timoclea (Hewitson, 1870) 358 Ministrymon ollantaitamba (K. Johnson, L.
Tribu Miller & Herrera, 1992)
Eumaeini 359 Ministrymon phrutus (Geyer, 1832)
111
Lycaenidae - Perú
360 Ministrymon una (Hewitson, 1873) 388 Strephonota buechei Faynel & Moser,
2011
361 Ministrymon zilda (Hewitson, 1873)
389 Strephonota caeruleus Faynel & Moser,
Nicolaea 70
2011
362 Nicolaea besidia (Hewitson, 1868) Strephonota carteia (Hewitson, 1870)
390
363 Nicolaea dolium (H. Druce, 1907) Strephonota cyllarissus (Herbst, 1800)
391
364 Nicolaea heraldica (Dyar, 1914) Strephonota dindymus (Cramer, 1775)

392
365 Nicolaea laconia (Hewitson, 1868) Strephonota elika (Hewitson, 1867)

393
366 Nicolaea munditia (H. Druce, 1907) Strephonota ericeta (Hewitson, 1867)
394
367 Nicolaea pyxis (K. Johnson, 1993) Strephonota foyi (Schaus, 1902)
395
368 Nicolaea salki (Faynel & Robbins, 2014) Strephonota hightoni (Bálint, 2014) Meridaria
396
369 Nicolaea velina (Hewitson, 1868) Strephonota jactator (H. Druce, 1907)
397
Ostrinotes 71 Strephonota metanira (Hewitson, 1867) Exorbaetta

398
370 Ostrinotes empusa (Hewitson, 1867) 399 Strephonota parvipuncta (Lathy, 1926) Robustana
371 Ostrinotes gentiana (H. Druce, 1907) 400 Strephonota perola (Hewitson, 1867)
372 Ostrinotes halciones (A. Butler & H. Druce, 401 Strephonota porphyritis (H. Druce, 1907) Serratonotes
1872)
402 Strephonota prina K. Johnson, Austin, Le
373 Ostrinotes purpuriticus (H. Druce, 1907) Crom & Salazar, 1997
374 Ostrinotes silva (Faynel & Robbins, 2014) 403 Strephonota purpurantes (H. Druce, 1907)
375 Ostrinotes sospes (Draudt, 1920) 404 Strephonota sphinx (Fabricius, 1775)
376 Ostrinotes tarena (Hewitson, 1874) 405 Strephonota strephon (Fabricius, 1775)
377 Ostrinotes tympania (Hewitson, 1869) 406 Strephonota syedra (Hewitson, 1867) Syedranota
Siderus 72 407 Strephonota tephraeus (Geyer, 1837)

378 Siderus athymbra (Hewitson, 1867) 408 Strephonota trebonia (Hewitson, 1870) Treboniana
379 Siderus bouvieri (Lathy, 1936) 409 Strephonota tyriam (H. Druce, 1907) Diminutina
380 Siderus leucophaeus (Hübner, 1818) Terenthina 74
381 Siderus philinna (Hewitson, 1868) 410 Terenthina terentia (Hewitson, 1868)
Strephonota 73 Theclopsis 75
382 Strephonota acameda (Hewitson, 1867) 411 Theclopsis gargara (Hewitson, 1868)
383 Strephonota adela (Staudinger, 1888) 412 Theclopsis lydus (Hübner, [1819])
384 Strephonota agrippa (Fabricius, 1793) Tmolus 76
385 Strephonota ambrax (Westwood, 1852) Tmolus cydrara (Hewitson, 1868)
413
386 Strephonota azurinus (A. Butler & H. Tmolus echion (Linnaeus, 1767)
414
Druce, 1872)
415 Tmolus mutina (Hewitson, 1867)
387 Strephonota berardi Faynel & Moser, 2011
112
Lycaenidae - Perú
416 Tmolus ufentina (Hewitson, 1868) 435 Erora nitetis (Godman & Salvin, 1887)
Tribu Erora phrosine (H. Druce, 1909)
436
Eumaeini
437 Erora senta (Draudt, 1920)
Subtribu Ocaria 83
Trichonidina 438 Ocaria aholiba (Hewitson, 1867)
Bistonina 77 439 Ocaria arpoxais (Godman & Salvin, 1887)
417 Bistonina biston (Möschler, 1877) 440 Ocaria calesia Hewitson, 1870
418 Bistonina feretria (Hewitson, 1878) 441 Ocaria cinerea (Lathy, 1936)
Trichonis 78 442 Ocaria clepsydra (H. Druce, 1907)
419 Trichonis immaculata Lathy, 1930 443 Ocaria elongata (Hewitson, 1870)
Tribu Ocaria elvira (K. Johnson, 1992)
444
Eumaeini
445 Ocaria ocrisia (Hewitson, 1868)
Subtribu 446 Ocaria sadei (Weeks, 1901)
Callophryidina 447 Ocaria thales (Fabricius, 1793)
Cyanophrys 79 Semonina 84
420 Cyanophrys amyntor (Cramer, 1775) 448 Semonina ares (Godman & Salvin, 1887)
421 Cyanophrys argentinensis (Clench, 1946) 449 Semonina semones (Godman & Salvin,
Cyanophrys banosensis (Clench, 1944) 1887)
422
Cyanophrys herodotus (Fabricius, 1793) Thaeides 85

423
424 Cyanophrys pseudolongula (Clench, 1944) 450 Thaeides muela (Dyar, 1913)
Chalybs 80 451 Thaeides theia (Hewitson, 1870)

Subfamilia
425 Chalybs hassan (Stoll, 1790)
426 Chalybs janias (Cramer, 1779) Polyommatinae
Chlorostrymon 81
Tribu
427 Chlorostrymon simaethis (Drury, 1773) Polyommatini
428 Chlorostrymon telea (Hewitson, 1868)
Subtribu
Erora 82 Sin Subtribu
429 Erora badeta (Hewitson, 1873) Eldoradina 86
430 Erora biblia (Hewitson, 1868)
452 Eldoradina cyanea (Balletto, 1993)
431 Erora caespes (H. Druce, 1907)
453 Eldoradina sylphis (Draudt, 1921)
432 Erora carla (Schaus, 1902)
Hemiargus 87
433 Erora gabina (Godman & Salvin, 1887)
454 Hemiargus hanno (Stoll, 1790)
434 Erora nana (C. Felder & R. Felder, 1865)
113
Lycaenidae - Perú
455 Hemiargus ramon (Dognin, 1887)
Itylos 88
456 Itylos ardisensis (Bálint & Lamas, [1997]) Madeleinea
457 Itylos bella (Bálint & Lamas, [1997]) Madeleinea
458 Itylos cobaltana (Bálint & Lamas, 1994) Madeleinea
459 Itylos colca (Bálint & Lamas, [1997]) Madeleinea
460 Itylos fumosus (Balletto, 1993) Madeleinea
461 Itylos gradoslamasi (Bálint, 1997) Madeleinea
462 Itylos huascarana (Bálint & Lamas, 1994) Madeleinea
463 Itylos koa (H. Druce, 1876) Madeleinea
464 Itylos lolita (Bálint, 1993) Madeleinea
465 Itylos mashenka (Bálint, 1993) Madeleinea
466 Itylos mira Bálint & Lamas, [1999] Madeleinea
467 Itylos pacis Draudt, 1921 Madeleinea
468 Itylos pasco Bálint & Lamas, 1994 Madeleinea
469 Itylos pnin Bálint, 1993 Madeleinea
470 Itylos tintarrona (Bálint & K. Johnson, Madeleinea

1995)
471 Itylos titicaca (Weymer, 1890) Madeleinea
Leptotes 89
472 Leptotes bathyllos Tessmann, 1928

473 Leptotes callanga (Dyar, 1913)
474 Leptotes cassius (Cramer, 1775)
475 Leptotes lamasi Bálint & K. Johnson, 1995
Nabokovia 90
476 Nabokovia cuzquenha Bálint & Lamas,

[1997]
477 Nabokovia faga (Dognin, 1895)
Paralycaeides 91
478 Paralycaeides inconspicua (Draudt, 1921)

479 Paralycaeides shade Bálint, 1993
Zizula 92
480 Zizula cyna (W. H. Edwards, 1881)
114
Lycaenidae - Perú
Nota: especies señaladas en la casilla tienen foto en el catálogo.

Note: species with highlighted box have a photo in the catalog. Printed:
GUIDE Especies por FAMILIA PAIS LIST TXT lunes, 9 de mayo de 2022
115
Bibliografía Bibliography
El Neotrópico por su diversidad de mariposas tiene The Neotropics, due to its diversity of butterflies,
mucha información publicada en libros y revistas have a lot of information published in books and
especializadas, esta información nos ha ayudado a specialized magazines. This information has helped
construir el catálogo de BioButterfly Database y a us to build the BioButterfly Database catalog and to
identificar muchas de nuestras fotos. identify many of our photos.
Inicialmente estábamos colocando la bibliografía en Initially we were placing the bibliography in each PDF
cada PDF tratando de asociar los artículos relevantes trying to associate the relevant articles to each
a cada PDF especifico, básicamente si se relaciona specific PDF, basically if it related to a family, a
con una familia, un país o una localidad. Decidimos country or a locality. We decided to make a general
hacer una bibliografía general combinada en formato combined bibliography as a separate PDF with
PDF independiente con citas para todas las citations for all the publications we have been using
publicaciones que hemos estado usando de tal in such a way that it is useful for all our PDFs and at
manera que sea útil para todos nuestros PDF y al the same time it frees up space in all the PDFs.
mismo tiempo libere espacio en todos los PDF. With this link access the PDF with the bibliography:
Con el siguiente vinculo acceda al PDF con la https://www.butterflycatalogs.com/neotropical-
bibliografía.: bibliography.html
https://www.butterflycatalogs.com/neotropical-
bibliography.html If you want to delve into the Neotropical
bibliography, with articles on butterflies from Mexico
Si se quiere profundizar en la bibliografía del to Argentina, we recommend an excellent document
Neotrópico, con artículos de mariposas desde compiled by Dr. Gerardo Lamas 2022 found on the
México hasta Argentina, recomendamos un Butterflies of America BOA page called: An Annotated
excelente documento recopilado por el Dr. Gerardo Bibliography of the Neotropical Butterflies and
Lamas 2022 que se encuentra en la página de Skippers (Lepidoptera: Papilionoidea and
Butterflies of America BOA llamado: An Annotated Hesperioidea) Revised Electronic Edition which can
Bibliography of the Neotropical Butterflies and be consulted at the following link:
Skippers (Lepidoptera: Papilionoidea and
Hesperioidea) Revised Electronic Edition que se http://www.butterfliesofamerica.com/L/Biblio.htm
puede consultar en el siguiente vinculo: We are constantly looking for information to help us
http://www.butterfliesofamerica.com/L/Biblio.htm complete the database, especially changes in the
taxonomy of species, new species and species by
Permanentemente estamos buscando información country or region. We appreciate all the references
que nos ayude a completar la base de datos, that you can send us.
especialmente cambios en la taxonomía de las
especies, nuevas especies y especies por país o
regiones. Agradecemos todas las referencias que nos
puedan enviar.
116
Páginas web recomendadas Recommended web pages
Existen muchísimas fuentes de información sobre There are many sources of information about
mariposas en Internet, sin embargo, presentamos butterflies on the internet, however, we present
acá una selección de aquellas recomendadas que here a selection of those recommended that can
puedan dar información complementaria a la provide complementary information to the
bibliografía, las listas de chequeo o las guías, o bibliography, the checklists, or the guides, or may
pueden tener información útil para la identificación have useful information for the identification or
o el conocimiento de las mariposas. knowledge of butterflies.
Identificación de mariposas - Butterfly Otras - Other

identification https://v3.boldsystems.org (DNA barcodes)
https://www.butterflycatalogs.com (libros de http://tolweb.org/tree/
identificación para la región neotropical) https://www.inaturalist.org/
www.neotropicalbutterflies.com (Mariposas del https://www.nic.funet.fi/
Neotrópico)
https://www.butterfliesofamerica.com Tour de mariposas – Butterfly tours
fotografías de todos los tipos www.butterfliesofcolombia.com
http://www.mariposabutterflytours.com/
Información general de mariposas - General
butterfly information Specific Families or Genera:
https://www.learnaboutbutterflies.com http://parides.genus.free.fr/syst.html
http://www.heliconius.org/2011/neil-rossers-
Nombres - Names maps-preliminary-versions/
https://www.nhm.ac.uk/our- Euptychiina Checklist:
science/data/lepindex/ https://www2.ib.unicamp.br/labor/site/?page_i
https://www.nic.funet.fi/pub/sci/bio/life/insect d=1020
a/lepidoptera/
Plantas hospederas - Host plants

https://www.nhm.ac.uk/our-
science/data/hostplants/search/index.dsml
Inventarios regionales - Regional inventories

http://calydna.com excelente fotografía/E
http://www.sangay.eu/es (Parque
Nacional Sangay, Ecuador)
http://www.cotacachi.eu (Región de Cotacachi,
W Ecuador)
http://www.loja-zamora.eu/en (Ecuador)
https://www.noreste.eu/en/les-papillons (NE
Ecuador)
http://www.butterfliesofecuador.com/
https://papillons-de-guyane.fr/papillons/
117
Localidades de las fotografías de la familia
Photo localities for the family
Abreviacion y Nombre localidad Ubicación Altura Min- Max m
Argentina AR
ChM Chacra Mariposa Obera, Misiones 300 300
Yab Yaboti Biosphere Reserve San Pedro, Misiones 200 200
Bolivia BO
Cor Coroico Nor Yungas, La Paz 1800 2000
Corc Coroico Nor Yungas, La Paz 1800 2000
Crv Caranavi Nor Yungas, La Paz 800 1200
Chl Chulumani Chulumani, La Paz 1200 1800
Volc Refugio Los Volcanes Florida, Santa Cruz 1300 1300
VTn Villa Tunari Chapare, Cochabamba 500 900
Vtu above Villa Tunari Chipiri, Cochabamba 800 800
Brasil BR
Blm Blumenau Blumenau, Santa Catarina
CJL Cristalino Jungle Lodge Alta Floresta, Mato Grosso 200 200
Mdb Massaranduba Massaranduba, Santa Catarina
Plm Reserva Natural Palmari Atalaia do Norte, Amazonas Am 80 80
PNI Parque Nacional do Itatiaia Resende, Rio de Janeiro 9500 3000
RG Rancho Grande Cacaulandia, Rondonia 300 300
Rgu Reserva Guapi Assu Bird Lodge Cachoeiras de Macacu, Rio de Janeir 0 200
SBc Serra do Bocaina Sao Jose do Barreiro, Sao Paulo 1150 1200
SBr Southeast Brazil Ubatuba, Sao Paulo 0
Colombia CO
- -, o
Afl Altos Florencia Florencia, Caquetá
Ang Carretera Caldas - Angelópolis Angelópolis, Antioquia 1800 2000
Bvt Sendero Buena vista Guaviare, Guainía
Cal Caluma Caluma, Bolivar 1200
118
Colombia CO
CCri Caño Cristales La Macarena, Meta 250 260
CdC Cueva del Condor San Luis, Antioquia 400 540
CRC Cañón del Río Claro - Reserva el Refugio San Francisco, Antioquia 350 600
FCy Finca El Caney Puerto Berrío, Antioquia
Gai Piscicola el Gaitero Sopetrán, Antioquia
Glp Municipio de Galapa Galapa, Atlántico
GMq Golfo de Morrosquillo Tolú, Sucre 0 10
Gurc Quebrada La Guaracu San Jerónimo, Antioquia 650 680
Gvr Varias Guaviare San José del Guaviare, Guaviare
HFr Hacienda Farias Concordia, Antioquia 1000 1100
HLB Humedal La Bramadora Sopetrán, Antioquia
HNp Parque tematico Hacienda Napoles Puerto Triunfo, Antioquia 200 250
Jard Municipio Jardín - Fundación Colibrí Jardín, Antioquia 1500 3000
LPC La Piñuela - Reserva la Cristalina Cocorná, Antioquia 800 900
Mco Municipio Mocoa Mocoa, Putumayo
MMt Municipio de Mitú Mitú, Vaupés 180 200
Mnc Minca Santa Marta, Magdalena
Mor Reserva Natural Moru Cumaribo, Vichada 80 100
Mzm Carretera Montezuma - PNN Tatamá Pueblo Rico, Risaralda 1100 2600
Oqy Santuario de F y F Otún Quimbaya Pereira, Risaralda 1800 1800
PGt Santa Cecilia- Piscina Gito Pueblo Rico, Risaralda 400 400
Pña Peñalisa Tarso, Antioquia 600 700
PSJ Pueblito de San Jose Amagá, Antioquia 1800 1900
PYm Parcelacion El Yarumo La Ceja, Antioquia
QEV Quebrada El Viao Cocorná, Antioquia 1870 1980
QLH Quebrada La Habana San Luis, Antioquia 700 750
RAA Reserva natural de las aves Arrierito Antioqueño Anorí, Antioquia 1400 1850
(Piha)
RAD Reserva natural de las aves El Dorado Santa Marta, Magdalena 950 2600
Rch Finca La Rochela Santa Rosa de Osos, Antioquia 1000 1100
RlR Reserva La Romera Sabaneta, Antioquia 1800 2200
RlT Reserva Las Tangaras El Carmen de Atrato, Chocó 1492 1800
Rmo Rincones de Monticello Medellín, Antioquia 1800 1850
119
Colombia CO
RNM Reserva Natural Mururito Puerto Gaitán, Meta 130 180
RP Río Pepino Mocoa, Putumayo 550 580
Rpau El Paujil - Las Quinchas Puerto Boyacá, Boyacá
RPF Reserva Paz y Flora Guasca, Cundinamarca
RRm Reserva Rocamadre Toluviejo, Sucre 92 320
RSD Río Santodomingo Cocorná, Antioquia
RSm Río Samana San Luis, Antioquia 380 420
SFx San Felix - Las Antenas Bello, Antioquia 2500 3100
Sidr San Isidro - Alto de la Ceja La Ceja, Antioquia 2000 2200
Sin Sinifaná - La Holanda Titiribí, Antioquia 600 790
SJn Municipio de San Jeronimo San Jerónimo, Antioquia
SLz San Lorenzo Santa Marta, Magdalena
SM Santa María Santa María, Boyacá
SNc San Nicolas - Parcelación Arreboles Sopetrán, Antioquia
SSb San Sebastian - Alto del Escobero Retiro, Antioquia
Tft Vereda Tafetanes Sopetrán, Antioquia
Tkk Reserva Natural Takarkuna Capurganá - Sapzurro Acandí, Chocó 0 50
VCC Vereda Cristalina Chamuscados Fredonia, Antioquia 1550 1650
VPN Vereda Paso Nivel - Finca Zanzibar Amagá, Antioquia 1450 1500
Costa Rica CR
Lsel La Selva Biological Station Puerto Viejo de Sarapiqui, Heredia
Ecuador EC
AJL Arajuno Jungle Lodge Tena, Napo 350 350
Atrk Apuya Track Tena, Napo 550 600
Bañ Baños Baños, Tungurahua 1500 2800
Baz Baeza Baeza, Orellana 1800 1800
BBR Rio Bigal Biological Reserve Loreto, Orellana 900 1100
Caj Cajanuma Entrance, PN Podocarpus, EC (Refugio Loja, Loja 2800
Cajanuma)
Cpl Copalinga Lodge Zamora, Zamora-Chinchipe 950 950
JSR Jatun Sacha Reserve Misahualli, Napo 400 400
120
Ecuador EC
Loj Loja area, EC (Villonaco Ridge,top of old Loja road ) Loja, Loja 2400 2750
Mnd Mindo San Miguel de Los Bancos, Pichincha 1000 1400
Orq Las Orquideas Yantaza, Zamora-Chinchipe 900 900
PaP Papallacta Pass Papallacta, Pichincha 4100
PmpL Pimpilala Lodge Tena, Napo 600 600
Podc Bombuscaro Ranger Station, Parque Nacional Zamora, Zamora-Chinchipe 1300 1300
Podocarpus
Pulu Pululahua Crater Calicali, Pichincha 2850
Rho Rio Hollin, Reserva Narupa Jondache, Napo 1050 1050
Ten Area de Tena Tena, Napo 510 550
Vilc Vilcabamba Loja, Loja 1500
WS WildSumaco Lodge Loreto, Orellana 1350 1350
Yach Napo lowlands(Yachana, La Selva, Napo Wildlife Coca, Orellana
Center)
Estados Unidos de US
RGv Rio Grande Valley Hidalgo, Texas
México MX
Cat Catemaco Catemaco, Veracruz 0 1100
CNv Cañon Novillo Ciudad Victoria, Tamaulipas 460 460
Ctm Catemaco Catemaco, Veracruz 200 200
GF Reserva de la Biosfera El Cielo Gomez Farias, Tamaulipas 350 1400
HCcd Hotel Cascades Orizaba, Veracruz 1200 1200
HTn Hotel Taninul Ciudad Valles, San Luis Potosí 45 45
LHt La Huerta Cortazar, Guanajuato 1800 1800
Mue Mueller's Xalapa, Veracruz 500
Oax Oaxaca Oaxaca, Oaxaca 1700 1700
Ord Ocampo road Ocampo, San Luis Potosí 1150 1150
Plq Palenque Joaquin Miguel Gutierrez, Chiapas 70 70
Teo Teotitlan de Valle Teotitlan, Oaxaca 1700 1700
Panamá PN
121
Panamá PN
Boc Bocas Bocas, Bocas del Toro
Bqt Boquete Boquete, Chiriquí 1400 1400
CTwr Canopy Tower Gamboa, Panamá 200 200
Fort Fortuna Field Station Hornito, Chiriquí 1000 1400
PplRd Pipeline Road Gamboa, Panamá 200 200
Tot Mt. Totumas Cloud Forest Volcan, Chiriquí 1800 2000
Vant Valle Antón El Valle de Anton, Coclé
Perú PE
Aco Aconabikh Tarapoto, San Martín 750 750
AMZ Amazonia Lodge Atalaya, Madre de Dios 400 400
ARCC Amazon Research and Conservation Center (Lago Las Piedras, Madre de Dios 230 230
Soledad)
BSht Bosque Shollet Oxapampa, Pasco 1800 2250
CoR Cock of the Rock Lodge/Paradise Lodge, Manu Road San Pedro, Cusco 1350 1400
Cus Cusco area, PE (Abra Malaga,etc) Ollantayambo, Cusco 4200
Chtk Chontachaka, Manu Road Pillcopata, Cusco 850 850
Hual Huallaca Valley Tarapoto, Áncash
Jgu Juan Guerra Tarapoto, San Martín 350 350
Jul La Julianita Moyabamba, San Martín 950 950
Lim Lima Lima, Lima 5 50
Mrd San Pedro, Manu Road San Pedro, Cusco 1350 1400
MWC Manu Wildlife Center Rio Madre de Díos, Madre de Dios 270 270
Pcy Picuroyacu Punchana, Loreto 150 150
PH Pampa Hermosa Chanchamayo, Junín 1200 1300
PMp Playa Mariposa Moyabamba, San Martín 1000 1050
PNM Parque Nacional del Manu San Pedro, Cusco 300 4000
PntL Pantiacolla Lodge Rio Madre de Díos, Madre de Dios 400 400
PNYC Parque Nacional Yanachaga-Chemillen Pozuzo, Pasco 1000 1450
Pro La Promisora San Ramon, Junín 1050 1050
Qtz Quitacalzon, Lower Manu Road San Pedro, Cusco 1050 1100
Rctl Rocotal, Manu road Pillahuata, Cusco 2000 2200
Rmp- Rumipata-Mishquiyacu Moyabamba, San Martín 950 1000
122
Perú PE
Rmpt Rumipata-Mishquiyacu Moyobamba, San Martín 950 1000
Salb San Alberto Oxapampa, Pasco 1800 2300
TngM Tingo Maria area Tingo Maria, Huánuco 650 650
TRC Tambopata Research Center Tambopata, Madre de Dios 240 240
VCm Villa Carmen, Manu Road Pillcopata, Cusco 500 550
Wyq Wayquecha Research Station Pillahuata, Cusco 2900 2900
123
Fotógrafos - Photographers
Este documento es el esfuerzo acumulativo de muchas personas. Los siguientes son los fotógrafos
que han contribuido con sus fotos para generar la guía. Muchas gracias a todos.
This list is the result of the cumulative efforts of a large number of people. The following are the
photographers who have contributed their photos to this work, thank you very much.
AWr Adam Winer JK John Kemner

AH Adrian Hoskins JMD John MacDonald
AM Amy McAndrews JOG John Ortega Galvan
AN Andrew Neild JAB Johnnier Arango Bermudez
Bsu Barry Sullender JJ Jorge J Restrepo
BBc Bill Beck Jpe Jorge Perez
BBt Bill Berthet JFG Jose Francisco Garcia
BBn Bill Bouton JMz Jose Monzon
CRm Cristóbal Ríos-Málaver BB Juan Guillermo Jaramillo Velasquez
DW Dan Wade JW Julia Whitmann
DJF Daniel Jaramillo Ferrer KNc Karen Nichols
DPt Daniel Piedrahita Thiriez KW Kay Wade
DvG David Geale KK Ken Kertell
DGz David Guzman KP Kevin Painter
FM Fredy Montero KD Kim Davis
GMr German Muriziasco KG Kim Garwood
GC Gill Carter KWs Kristine Wallstrom
GM Glenn Mahler LG Leif Gabrielsen
GS Gottfried Siebel LBd Leonel Baldoni
HB Hank Brodkin Leo Leopoldina Tapasco
JZn Jhonny Zambrano LC Les Catchick
JB Jim Brock LuL Luc Legal
JHg Jim Hengeveld LEM Luis Eduardo Mejia
JSn Jim Snyder MCC Marcos Cesar Campis
JAR João Amarildo Ranguetti MIW Maria Isabel Weyermans
JSl Joe Schelling Mmi Maris Midgely
124
MZ Maristela Zamoner THn Tom Horton
MM Martin Moreno TH Tony Hoare †
MR Martin Reed WC Will Carter
MG Michael Garcia
MC Mike Chapman
MRk Mike Rickard
MS Mike Stangeland
Cdr n Calderon
Mnz n Mendez
MHN Natural History Museum London
OE Oscar Enciso
PS Pamela Sai
PM Pat Martin
PT Paul Thompson
PL Pingchung Lee
PB Priscilla Brodkin
RnM Renato Mattei
RC Rick Cech
RH Rich Hoyer
RL Richard Lehman
RW Rob Westerduijn
RGd Robert Gallardo
RGl Robert Gilson
RSb Robert Straub
RRt Roger Rittmaster
RD Rudy Dodero
Smd Santiago Mejía Dugand
SO Scott Olmstead
SFn Shawneen Finnegan
SN Sherry Nelson
SS Shirley Sekarajasingham
SDM Sidney Dantas de Melo
TGz Terry Gonzales
TS Tinio Sanchez
125
Agradecimientos Acknowledgments
Estas guías y listados de chequeo son el resultado del These guides and checklist are the result of the effort
esfuerzo de un gran número de colaboradores, of a large number of collaborators, scientists, fans
científicos, aficionados y fotógrafos a quienes damos and photographers to whom we thank, because
las gracias, pues sin ellos, estas publicaciones no without them, these publications would not have
hubieran sido posible de completar. been possible to complete.
En la familia Hesperiidae agradecemos In the Hesperiidae family we especially thank the
especialmente al experto Bernard Hermier por las expert Bernard Hermier for the countless hours and
innumerables horas y páginas de explicaciones y pages of explanations and corrections without which
correcciones sin las cuales habría sido imposible it would have been impossible to make this
hacer este documento. document.
Al Dr. Keith Willmott por sus consejos en la To Dr. Keith Willmott for his advice in preparing the
elaboración de la presentación de este trabajo e presentation of this work and identification of
identificación de fotos. photos.
Agradecemos a los siguientes expertos que We thank the following experts who have generously
generosamente han ayudado con información y en la helped with information and in the identification of
identificación de algunas fotos que han servido de some photos that have served as support for the list
soporte al listado (en orden alfabético): Stéphane (in alphabetical order): Stéphane Attal, Dr. Andrew
Attal, Dr. Andrew Brower, Prof. James Mallet, Brower, Prof. James Mallet, Andrew Neild, Gregory
Andrew Neild, Gregory Nielsen, Dr. Tomasz Pyrcz, Dr. Nielsen, Dr. Tomasz Pyrcz, Dr. Robert Robbins, Julián
Robert Robbins, Julián Salazar y Aaron Soh. Salazar and Aaron Soh.
A Andrés Arenas por sus consejos y ayuda en la To Andrés Arenas for his advice and help in preparing
elaboración de la base de datos. A Andrew Neild por the database. To Andrew Neild for his
sus recomendaciones y sugerencias. A David Geale recommendations and suggestions. To David Geale
por sus innumerables aportes fotográficos. for his countless photographic contributions.
A Gabriel Rodríguez por sus recomendaciones en To Gabriel Rodríguez for his recommendations on
cómo distribuir las zonas biogeográficas y por los how to distribute the biogeographic zones and for
múltiples datos geográficos de su colección de the multiple geographic data of his collection of
mariposas. Al Dr. Juan José Arango, Rodrigo Bernal y butterflies. To Dr. Juan José Arango, Rodrigo Bernal
Daniel Jaramillo F. por sus consejos y colaboración en and Daniel Jaramillo F. for their advice and
la consecución de datos. A Maristela Zamoner por la collaboration in obtaining data. To Maristela
traducción al portugués. A Jean-Claude Petit por las Zamoner for the portuguese translation. To Jean-
múltiples adiciones de especies y correcciones para Claude Petit for the multiple additions of species and
Ecuador. corrections for Ecuador
Al Museo de Historia Natural de Londres (Natural
History Museum) por permitirnos el acceso a sus To the Natural History Museum of London for
colecciones, ejemplares y datos. A Blanca Huertas allowing us access to their collections, specimens
por todos sus concejos y recomendaciones. and data. To Blanca Huertas for all her advice and
Especial agradecimiento a las familias de los autores, recommendations.
por su apoyo durante la realización de este proyecto. Special thanks to the families of the authors, for their
support during the realization of this project.
126
Esta guía es un legado porque varios de los This guide is a legacy because several of the
sitios donde fueron registradas las especies ya sites where the species were recorded have
están destruidos y por lo tanto estas especies already been destroyed and therefore these
ya no existen en esa localidad específica. El species no longer exist in that specific locality.
listado y la guía ilustran la asombrosa The checklists and the guides illustrate the
diversidad de mariposas del Neotrópico para amazing diversity of butterflies of the
motivar su conocimiento y conservación. Neotropics to motivate their knowledge and
conservation.
“Solo podemos conservar “We can only conserve

y proteger lo que and protect what we
conocemos” know" JGJ

Peru Lycaenidae Butterfly Guide 09may2022 Butterflycatalogs

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Peru Lycaenidae Butterfly Guide 09may2022 Butterflycatalogs

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Formatos disponibles

“Este listado está dedicado a todas las personas que nos han ayudado a construirlo y a

todos aquellos que están trabajando por el conocimiento y la conservación de la

Para citar este documento utilice: To cite this document use:

GARWOOD K., JARAMILLO J.G. Guía de mariposas Lycaenidae de Perú – Butterfly

Annamaria ganimedes Symbiopsis perulera

BioButterfly Database – Butterflycatalogs.com

1- Nombre científico 1- Scientific name

BioButterfly Database – Butterflycatalogs.com

BioButterfly Database – Butterflycatalogs.com

BioButterfly Database – Butterflycatalogs.com

BioButterfly Database – Butterflycatalogs.com

Subtribu - Subtribe Jantheclina

BioButterfly Database - Butterflycatalogs

BioButterfly Database - Butterflycatalogs

BioButterfly Database - Butterflycatalogs

Brevianta ematheon Brevianta ematheon

Brevianta undulata Eumaeus toxana Eumaeus toxana

Micandra aegides ♂ Micandra aegides ♀ Micandra comae ♂

BioButterfly DataBase - Butterflycatalogs.com Perú Theclinae

Micandra comae ♀ Micandra comae ♀ Micandra comae ♀

Micandra comae ♀ Micandra comae ♀ Micandra comae

Micandra dignota Micandra ion Micandra platyptera ♂

Micandra platyptera ♂ Micandra platyptera ♀ Micandra platyptera ♀

BioButterfly DataBase - Butterflycatalogs.com Perú Theclinae Página 2 de 87

Micandra platyptera ♀ Micandra platyptera ♀ Micandra platyptera ♀

Micandra platyptera ♀ Micandra platyptera ♀ Mithras nautes ♂

Mithras nautes ♀ Mithras nautes Theorema dysmenia

Thestius epopea Thestius epopea Thestius epopea

BioButterfly DataBase - Butterflycatalogs.com Perú Theclinae Página 3 de 87

Thestius meridionalis ♂ Thestius meridionalis ♂ Thestius meridionalis ♂

Thestius meridionalis ♂ Thestius meridionalis ♂ Thestius meridionalis ♀

Balintus tityrus Balintus tityrus

BioButterfly DataBase - Butterflycatalogs.com Perú Theclinae Página 4 de 87

Balintus tityrus Johnsonita auda ♂ Johnsonita auda

Johnsonita auda Johnsonita auda Johnsonita auda

Johnsonita auda Johnsonita auda Johnsonita chlamydem ♂

Johnsonita chlamydem ♂ Johnsonita chlamydem ♀ Johnsonita chlamydem ♀

BioButterfly DataBase - Butterflycatalogs.com Perú Theclinae Página 5 de 87

Johnsonita chlamydem Johnsonita chlamydem Johnsonita pardoa

Johnsonita pardoa Lathecla latagus Lathecla latagus

Lathecla mimula Lathecla mimula Lathecla mimula

Lathecla mimula Rhamma arria ♂ Rhamma arria

BioButterfly DataBase - Butterflycatalogs.com Perú Theclinae Página 6 de 87

Rhamma aurugo Rhamma commodus Rhamma commodus

Rhamma commodus Rhamma comstocki ♀ Rhamma comstocki

Rhamma familiaris Rhamma familiaris Rhamma hybla

Rhamma sp2 Rhamma sp2 Salazaria sala

BioButterfly DataBase - Butterflycatalogs.com Perú Theclinae Página 7 de 87

Salazaria sala Salazaria sala

Marachina maraches Marachina maraches

Nesiostrymon calchinia Nesiostrymon calchinia Nesiostrymon celona

Nesiostrymon celona Nesiostrymon celona Paraspiculatus azul

BioButterfly DataBase - Butterflycatalogs.com Perú Theclinae Página 8 de 87

Paraspiculatus colombiensis Paraspiculatus elis Paraspiculatus elis

Paraspiculatus oroanna Paraspiculatus orobia ♂ Paraspiculatus orobia

Paraspiculatus orobia Paraspiculatus orobia Paraspiculatus orobia

Paraspiculatus orobia Paraspiculatus orocana Paraspiculatus orocana

BioButterfly DataBase - Butterflycatalogs.com Perú Theclinae Página 9 de 87

Penaincisalia amatista Penaincisalia atymna Penaincisalia culminicola

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