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Cainozoic Larger Foraminifers From Dominican Repub
Cainozoic Larger Foraminifers From Dominican Repub
(1) Dept. d’Estratigrafia, Paleontología i Geociències Marines, Fac. de Geol, Univ. de Barcelona. Martí y Franquès s/n, 08028, Barcelona, España.
e-mail: [email protected], [email protected]
(2) Dept. de Geodinàmica i Geofísica, Fac. de Geología, Univ. de Barcelona. Martí y Franquès s/n, 08028, Barcelona, España.
e-mail: [email protected],
(3) INYPSA. General Díaz Porlier 49, 28001 Madrid, España.
e-mail: [email protected]
ABSTRACT
This work describes the Cainozoic larger foraminifers from Dominican Republic and discuses their biostratigraphic application to the cons-
truction of a lithostratigraphic framework. We have differentiated the following associations:
a. Ranikothalia bermudezi and Neodiscocyclina grimsdalei, which characterizes the Late Paleocene (Thanetian).
b. Eoconuloides wellsi, Eoconuloides lopeztrigoi and Neodiscocyclina barkeri from the Early Eocene reworked in Middle Eocene stra-
ta.
c. Coleiconus elongatus, Fallotella cookei, Cushmania puilboreauensis, Cushmania americana, Fabiania cassis, Eoconuloides lopez-
trigoi, Amphistegina parvula, Discocyclina marginata, Polylepidina antillea and Penoperculinoides cubensis, that characterizes the
Middle Eocene
d. Fallotella cookei, Cushmania americana, Fabiania cassis, Amphistegina parvula, Lepidocyclina (Lepidocyclina) macdonaldi,
Lepidocyclina (Nephrolepidina) cf. chaperi, Lepidocyclina (Pliolepidina) cf. peruviana and Nummulites cf. willcoxi, that characteri-
zes the Middle-Late Eocene interval.
e. Nummulites cf. panamensis, Heterostegina cf. antillea, Lepidocyclina (Lepidocyclina) mantelli, Lepidocyclina (Lepidocyclina) cane-
llei, Lepidocyclina (Eulepidina) undosa and Lepidocyclina (Nephrolepidina) tournoueri, characteristic from the Oligocene in middle
shelf environments.
f. Archiasinids, Praerhapydionina cf. delicata and Discorinopsis sp., characteristic from the Oligocene in shallow carbonate shelves.
g. Miogypsina cf. antillea and Miarchaias floridanus that indicates an Early-Middle Miocene age.
h. Miarchaias floridanus, Miosorites americanus, Annulosorites spiralis and Operculinoides cf. bullbrooki, characteristic from the
Early-Late Miocene.
Este trabajo describe los macroforaminíferos del Cenozoico de la República Dominicana y estudia su aplicación biostratigráfica en la cons-
trucción de un marco litostratigráfico. Se han diferenciado las siguientes asociaciones:
a. Ranikothalia bermudezi y Neodiscocyclina grimsdalei, que caracteriza al Paleoceno Superior (Thanetiense).
b. Eoconuloides wellsi, Eoconuloides lopeztrigoi y Neodiscocyclina barkeri del Eoceno Inferior, retrabajados en estratos del Eoceno
Medio.
c. Coleiconus elongatus, Fallotella cookei, Cushmania puilboreauensis, Cushmania americana, Fabiania cassis, Eoconuloides lopeztrigoi,
Amphistegina parvula, Discocyclina marginata, Polylepidina antillea y Penoperculinoides cubensis, que caracteriza al Eoceno Medio.
d. Fallotella cookei, Cushmania americana, Fabiania cassis, Amphistegina parvula, Lepidocyclina (Lepidocyclina) macdonaldi,
Lepidocyclina (Nephrolepidina) cf. chaperi, Lepidocyclina (Pliolepidina) cf. peruviana y Nummulites cf. willcoxi, que caracteriza al
intérvalo Eoceno Medio-Superior.
e. Nummulites cf. panamensis, Heterostegina cf. antillea, Lepidocyclina (Lepidocyclina) mantelli, Lepidocyclina (Lepidocyclina) cane-
llei, Lepidocyclina (Eulepidina) undosa y Lepidocyclina (Nephrolepidina) tournoueri, característica del Oligoceno en ambientes de
plataforma media.
f. Archiasinids, Praerhapydionina cf. delicata y Discorinopsis sp., característica del Oligoceno en ambientes de plataforma somera.
g. Miogypsina cf. antillea y Miarchaias floridanus que indica una edad Mioceno Inferior-Medio.
h. Miarchaias floridanus, Miosorites americanus, Annulosorites spiralis y Operculinoides cf. bullbrooki, característica del Mioceno
Inferior-Superior.
359
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
Fig. 1. Areas sampled in Dominican Republic with location of the chronostratigraphic diagram of Figure 2
Fig.1. Areas muestreadas en la República Dominicana y situación del esquema cronostratigráfico de la Figura 2
360
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
Fig. 2. Chronostratigraphic diagram showing lithology, facies, larger foraminifer associations and depositional sequences in the Cainozoic
of western Dominican Republic (location in Fig. 1)
Fig.2. Diagrama cronostratigráfico que muestra la litología, facies, asociaciones de macroforaminíferos y las secuencias deposicionales
del Cainozoico del oeste de la República Dominicana (situación en Fig. 1)
lected referred to geographical coordinates and the different authors are heterochronous ranging
stratigraphic framework of the geological map and from the Paleocene to the Middle Eocene (Bowin,
deposited in the collections of the Dirección General 1966; Boisseau, 1987; Bourdon, 1985). Samples
de Minería in Santo Domingo. Although the sampling examined in this paper were acquired in the Don
lacks rigorous stratigraphic succession, it is enough Juan basin, Monteplata sheet.
for local biostratigraphic resolution. More sampling is — La Luisa Fm (Hernaiz Huerta, 2004). It is compo-
needed for regional correlation with other Caribbean sed of siltstones, sandstones with chert and micri-
biozones. tic limestones. La Luisa Fm is ascribed by the
The following lithostratigraphic units have been same author to the Middle Eocene. Samples exa-
studied: mined in this paper were acquired in the Don
— Don Juan Fm (Bowin, 1966). It is composed of Juan Basin, Monteplata sheet.
siltstones, sandstones and greywackes with inter- — Loma Peñón Fm (Bourdon, 1985; Lebrón and
calations of conglomerates, breccias, tuffs, cineri- Mann, 1991). It is composed of calcarenites and
tes and grey limestones. The ages provided by calcirudites. These autors recognize reworked
361
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
362
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
fauna of the Paleocene and ascribed the Loma Middle Pliocene (Saunders et al., 1986; Akers in
Peñón Fm to the Middle Eocene. Samples exami- Vokes, 1989; Hottinger, 2001). Samples examined
ned in this paper were acquired in the Bejucal in this paper were acquired in the Cibao basin,
Basin, Rincón Chavón sheet. Mao and Pepillo Salcedo sheets.
— Río Yabón Fm (Bourdon, 1985; Lebrón and Mann, — Las Lavas Fm (Zoeten and Mann, 1999). It is com-
1991). It is composed of massive limestones. The posed of calcareous breccias with blocks of volca-
Río Yabón Fm is ascribed by the same authors to nic rocks, marls and limestones (turbidite facies).
the Middle Eocene. Samples examined in this Las Lavas Fm is ascribed to the Late Oligocene-
paper were acquired in the Río Yabón basin, Early Miocene (Zoeten and Mann, 1999) and to
Miches sheet. the Late Oligocene-Middle Miocene by Bernárdez
— Haitises Fm (Brouwer and Brouwer, 1982). It is (2004). Samples examined in this paper were
composed of reefal limestones. The Haitises Fm is acquired in the Septentrional Cordillera, Monte
ascribed to the Middle to Late Miocene on the Cristi sheet.
basis of its stratigraphic position a top of the Early- — Monte Cristi unit (Bernárdez, 2004). It is compo-
Middle Miocene Yanigua Fm (Iturralde-Vinent, sed of greywackes and limolites (turbidite facies).
2001). Samples examined in this paper were acqui- The Monte Cristi unit is ascribed by the same
red in the Hato Mayor and Monte Plata sheets. author to the Middle-Late Miocene. Samples exa-
— Cercado Fm (Maury, 1919). It is composed of luti- mined in this paper were acquired in the Cibao
tes, marls, sandstones, conglomerates, bioclastic basin, Monte Cristi sheet.
and reefal limestones. The Cercado Fm is ascri- — Neiba Fm (Vaughan et al., 1921; Bermúdez, 1949).
bed to the Late Miocene (Saunders et al., 1986). Micritic pelagic limestone with chert. The Neiba
Samples examined in this paper were acquired in Fm is ascribed by these authors to the Middle-
the Cibao basin, Monción sheet. Late Eocene. In the Neiba Sierra, Hernaiz Huerta
— Gurabo Fm (Maury, 1919). It is composed of 2004a, 2004b; expand this range to the Early
marls, lutites, sandy limestones, calcareous Eocene-Early Miocene, and define the following
sandstones, limestones, limestones with corals. informal units (see also Hernaiz Huerta et al.,
The Gurabo Fm is ascribed to the Late Miocene- 2007):
1.- Sulcoperculina globosa DE CIZANCOURT, 1949. Rincón Chavón 6472-III-IG JG 9027, x40
2.- Sulcoperculina globosa DE CIZANCOURT, 1949. Axial section. Rincón Chavón 6472-III-IG JG 9004, x40
3.- Orbitoides sp. Rincón Chavón 6472-III-IG JG 9027, x20
4.- Cubanina sp., transversal section. Monción 5974-II-BR FC 9064, x20
5.- Cubanina sp., oblique section. Mao 5974-I-BR EB 9031, x20.
6, 7.- Discorinopsis sp. Jimaní 5871-III-IG JG 9003, x20
8.- Coleiconus elongatus (COLE, 1942). La Descubierta 5871-I-IG HH 9021, x20
9.- Coleiconus elongatus (COLE, 1942). La Descubierta 5871-I-IG HH 9038, x20
10.- Coleiconus elongatus (COLE, 1942). Galván 5971-IV-IG HH 9030, x20
11.- Fallotella sp. 1. Rincón Chavón 6472-III-IG JG 9002, x20
12.- Cushmania puilboreauensis (WOODRING, 1924). La Descubierta 5871-I-IG HH 9020, x20
13.- Fallotella cookei (MOBERG, 1928). Bánica 5873-II-BR JS 9029, x20
14.- Fallotella cookei (MOBERG, 1928). Arroyo Limón 5973-III-BR EB 9048, x20
15.- Fallotella cookei (MOBERG, 1928). Bánica 5873-II-BR JS 9030, x20
16.- Cushmania puilboreauensis (WOODRING, 1924). La Descubierta 5871-I-IG HH 9020, x20
17.- Cushmania americana (CUSHMAN, 1919). Arroyo Limón 5973-III-BR EB 9048, x20
18.- Cushmania puilboreauensis (WOODRING, 1924). La Descubierta 5871-I-IG HH 9020, x20
19.- Cushmania americana (CUSHMAN, 1919). La Descubierta 5871-I-IG HH 9021, x20
20.- Annulosorites spiralis HOTTINGER, 2001. Bánica 5873-II-BR JS 9017, x20
21, 22.- Praerhapydionina cf. delicata HENSON, 1950. Jimaní 5871-III-IG JG 9003, x20
23.- Miarchaias floridanus (CONRAD, 1846). Bánica 5873-II-BR JS 9017, x20
24.- Annulosorites spiralis HOTTINGER, 2001. Monte Plata 6272-III-IG HH 9037, x20
25.- Miarchaias floridanus (CONRAD, 1846). Pepillo Salcedo 5875-II-EB 9002, x20
26.- Miosorites americanus (CUSHMAN, 1918). Bánica 5873-II-BR JS 9023, x40
27.- Miarchaias floridanus (CONRAD, 1846). Pepillo Salcedo 5875-II-EB 9002, x20
28.- Miarchaias floridanus (CONRAD, 1846). Bánica 5873-II-BR JS 9017, x20
29.- Miosorites americanus (CUSHMAN, 1918). Hato Mayor 6372-III-IG JG 9004, x40
363
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
- Lower Neiba Fm (Early-Middle Eocene). It is among others. Butterlin (1954) expand the age
composed of regular, frecuently thick bedded range to the Late Oligocene. Samples examined in
limestones with chert. Samples examined in this paper were acquired in the Bánica sheet.
this paper were acquired in La Descubierta
and Galván sheets.
- Brecciated Neiba unit (Late Eocene-Early SYSTEMATICS
Miocene). It is composed of brecciated massi-
ve limestones with corals and gastropoda. Family GLOBOTEXTULARIIDAE Cushman, 1927
Samples examined in this paper were acqui- Genus Cubanina Palmer, 1936
red in La Descubierta and Galván sheets. Pl. 1, figs. 4 and 5
- El Aguacate de Neiba Volcanosedimentary
Complex (Middle-¿Late Eocene?). It is compo- Samples. Gurabo Fm (Mao 5974-I-BR EB 9031),
sed of basalt and andesite flows, volcanic Cercado Fm (Monción 5974-II-BR FC 9064).
breccias and tuffs, calcarenites and marls. Remarks. Agglutinated form with initial biserial
Samples examined in this paper were acqui- growth that changes to uniserial. The chambers
red in La Descubierta sheet. have an endoskeleton composed of radial partitions
- Upper Neiba Fm (Late Eocene-Early Miocene). perpendicular to the septum. We found one trans-
It is composed of regular, thin bedded limes- versal section and an oblique one. This form differs
tones with chert, and intercalations of marly to Cubanina alavensis defined in the Oligocene of
limestones and marls to the top. Samples exa- Cuba (Palmer 1936) because of its larger dimensions
mined in this paper were acquired in the and a higher number of exoskeletal partitions in
Jimaní, La Descubierta and Galván Arroyo each chamber.
Limón and Bánica sheets. Distribution. This form occurs in Gurabo Fm associat-
— Catanamatías Fm (Bernárdez and Soler, 2004). ed with Operculinoides cf. bullbrooki, Miosorites amer-
Defined in the southwest margin of the Central icanus and Sphaerogypsina globula, and in Cercado
Cordillera as a transitional unit between the Neiba Fm associated with Operculinoides cf. bullbrooki.
and Sombrerito Fms, it is composed of marls, cal- Age. The associated foraminifers indicate a Miocene age.
carenites, sandy and bioclastic limestones, calca-
renitic turbidites, breccias, conglomerates. The
Family VALVULAMMINIDAE Loeblich and Tappan,
Catanamatías Fm is ascribed by the same authors
1987
to the Late Oligocene. Samples examined in this
Genus Discorinopsis Cole, 1941
paper were acquired in the Arroyo Limón and
Pl. 1, figs. 6 and 7
Bánica sheets.
— Sombrerito Fm (Olson, en Bermúdez, 1949). It is Samples. Upper Neiba Fm (Jimaní 5871-III-IG JG
composed of marls/marly limestones, calcareni- 9003).
tes, conglomerates, breccias with calcareous and Remarks. Agglutinated form with a high-trochospiral
volcanic clasts. McLaughlin et al. (1991) ascribed test, with an endoskeleton composed of pillars. In our
the Sombrerito Formation to the late Early material we have only two oblique sections and the
Miocene-earliest part of the Late Miocene. The specific determination is not possible.
base of the Sombrerito crosses time lines, beco- Distribution. This form occurs in the Upper Neiba Fm
ming older (Oligocene) toward the northwest associated with archaiasinids, peneroplids and
(Hernaiz Huerta et al., 2007). Samples examined Praerhapydionina cf. delicata.
in this paper were acquired in the Bánica sheet. Age. According to Cole (1941) and Robinson and
— Barahona Mb of Sombrerito Fm (Hernaiz Huerta Wright (1993) stratigraphic range of the genus
et al., 2007). It is composed of massive limestones Discorinopsis is Middle Eocene to Oligocene.
with sporadic corals. The Barahona Mb is ascri-
bed by the same author to the Miocene. Samples
examined in this paper were acquired in the Family COSKINOLIDAE Moullade, 1965
Jimaní and La Descubierta sheets. Genus Coleiconus Hottinger and Drobne, 1980
— Madame Joie Fm (Woodring, 1922). It is composed Coleiconus elongatus (COLE, 1942)
of coral and bioclastic limestones, sandy and con- Pl. 1, figs. 8, 9 and 10
glomeratic limestones. The Madame Joie Fm is
ascribed to the Early Miocene by Woodring (1922), 1956 Coskinolina elongata COLE, Cole, p. 215; Pl. 24,
Woodring et al. (1924) and Maurrasse (1982) figs. 6-11; Pl. 31, figs. 1-2
364
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
1960 Coskinolina elongata COLE, Butterlin and Fallotella cookei (MOBERG, 1928)
Bonet, p. 9-14; Pl. 1, figs. 1-3 Pl. 1, figs. 13, 14 and 15
1964 Coskinolina elongata COLE, Cole and Applin, p.
24-25; Pl. 1, fig. 12; Pl. 2, fig. 9; Pl. 3, figs. 4, 8 1941 Dictyoconus cookei (MOBERG), Cole, Pl. 18,
1980 Coskinolina (Coleiconus) elongata COLE, figs. 11-12
Hottinger and Drobne, p. 233, 235; Pl. 13, figs. 1947 Dictyoconus cookei (MOBERG), Cole and
7-14; Text figs. 11A, B, C, D Bermúdez, p. 7; Pl. 1, fig. 6
1993 Coskinolina sp. cf. C. douvillei (DAVIES), 1952 Dictyoconus cookei (MOBERG), Cole and
Robinson and Wright, p. 289; Figs 7.1-3 Gravell, p. 711-712; Pl. 90, figs. 5, 8, 13, 15
1956 Dictyoconus cookei (MOBERG), Cole, Pl. 25,
Samples. Upper Neiba Fm (Galván 5971-IV-IG HH figs. 6-7
9030), Brecciated Neiba Fm (La Descubierta 5871-I-IG 1968 Heterodictyoconus cookei (MOBERG), Butterlin
HH 9021 and 9038). and Moullade, p. 13; Pl. 2, figs. 1-11; Pl. 3, figs.
Remarks. The morphology of the test is high-conical 1-3
to conical. The growth is pfenderinid-type (Pl. 1, fig. 9) 1980 Fallotella (Fallotella) cookei (MOBERG),
in the early stages and uniserial at the adult stage. Hottinger and Drobne, p. 53, 54-56; Pl. 1, fig. 3;
The basal diameter reaches 1.25 mm in the 7th unise- Pl. 16, figs. 1-14; Text fig. 9D, E, 12A
rial chamber. The endoskeleton consists of a few pil- 1993 Fallotella cookei (MOBERG), Robinson and
lars, 7 pillars in the 8th uniserial chamber. The
Wright, p. 281; Figs. 9.1-6; 10.3-4
exoskeleton consists of simple radial beams. In the
marginal zone the chambers are inflated.
Samples. Upper Neiba Fm (Galván 5971-IV-IG HH
Distribution. Coleiconus elongata occurs in the Upper
9030, Bánica 5873-II-BR JS 9029 and 9030; Arroyo
Neiba Fm associated with Fallotella cookei,
Limón 5973-III-BR EB 9052), Catanamatías Fm (Arroyo
Eoconuloides lopeztrigoi, Discocyclina marginata and
Limón 5973-III-BR EB 9048), Rio Yabón Fm (Miches
Amphistegina parvula. In the Brecciated Neiba Fm it
6372-I-IG AD 9016).
is associated with Cushmania americana, Fabiania
Remarks. The test is conical to high-conical. The basal
cassis, Amphistegina parvula and Eoconuloides
lopeztrigoi. diameter reaches 1.35 mm in the 12th uniserial cham-
Age. According to Cole (1956), Cole and Applin ber. In basal section of 1.25 mm of diameter shows 12
(1964), and Hottinger and Drobne (1980) Coleiconus partitions per quadrant. The endoskeleton is com-
elongata ranges from the latest Early Eocene to the posed of pillars and the exoskeleton of beams and
Middle Eocene. rafters (Pl.1, fig. 14).
Distribution. This form occurs in the Upper Neiba Fm
associated with Coleiconus elongatus, Cushmania
americana, Eoconuloides lopeztrigoi, Amphistegina
Family ORBITOLINIDAE Martin, 1980 parvula, Fabiania cassis, Eodictyoconus cubensis,
Genus Fallotella Mangin, 1954 Discocyclina marginata, Polylepidina antillea,
Fallotella sp. 1 Lepidocyclina (Lepidocyclina) macdonaldi,
Pl. 1, fig. 11 Lepidocyclina (Nephrolepidina) cf. chaperi and
Lepidocyclina (Pliolepidina) cf. peruviana. In the Rio
Samples. Loma Peñón Fm (Rincón Chavón 6472-III-IG Yabón Fm Fallotella cookei occurs associated with
JG 9002). Fabiania cassis, Eoconuloides lopeztrigoi,
Remarks. We found a basal section of a small speci- Amphistegina parvula and Lepidocyclina
men of Fallotella sp. The diameter of this section is 0.9 (Lepidocyclina) macdonaldi. Finally, in the breccias of
mm and shows 8 marginal partitions per quadrant. the Catanamatías Fm this form is associated with
Distribution. This form occurs, together with Cushmania americana, Fabiania cassis, Amphistegina
Ranikothalia bermudezi, as Paleocene forms reworked parvula and Lepidocyclina (Nephrolepidina) cf. chaperi.
into the Loma Peñón Fm, associated to Eocene larger Age. According to Hottinger and Drobne (1980)
foraminifera as Eoconuloides lopeztrigoi and Fallotella cookei occurs from the latest Early to the
Discocyclina marginata. Middle Eocene, whereas according to Robinson and
Age. We consider this species as Paleocene in age Wright (1993), and Robinson (1995) this species
because of its small size and because it occurs with occurs from the early Middle Eocene to, possibly, the
other Paleocene species. earliest Oligocene.
365
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
Genus Cushmania Silvestri, 1925 Age. Cushmania americana occurs in the Middle
Cushmania americana (CUSHMAN, 1919) Eocene according to Hottinger and Drobne (1980),
Pl. 1, figs. 17 and 19 and in Jamaica it ranges from the latest Early Eocene
to the earliest Late Eocene according to Robinson and
1928 Cushmania americana (CUSHMAN), Vaughan, Wright (1993).
281-282; Pl. 44, figs. 1-2
1944 Dictyoconus americanus (CUSHMAN), Cole, p.
36-37; Pl. 4, figs 1-6; Pl. 18, fig. 11 Cushmania puilboreauensis (WOODRING, 1924)
1944 Dictyoconus americanus (CUSHMAN), Cole and Pl. 1, figs. 12, 16 and 18
Bermúdez, p. 6; Pl. 3, figs. 6-10
1947 Dictyoconus americanus (CUSHMAN), Cole and 1928 Dictyoconus puilboreauensis WOODDRING,
Bermúdez, p. 195; Pl. 14, figs. 5, 8 Vaughan, p. 281; Pl. 43, fig. 6
1952 Dictyoconus americanus (CUSHMAN), Cole and 1944 Dictyoconus americanus (CUSHMAN), Cole, Pl.
Gravell, p. 711; Pl. 90, figs. 1-2, 12, 14 8, figs 12-13
1956 Dictyoconus americanus (CUSHMAN), Cole, p. 1952 Dictyoconus americanus (CUSHMAN), Cole and
217-218; Pl. 25, figs. 10-11 Gravell, p. 711, Pl. 90, figs. 6-8
1964 Dictyoconus americanus (CUSHMAN), Cole and 1956 Dictyoconus americanus (CUSHMAN), Cole, Pl.
Applin, p. 25; Pl. 2, figs. 3, 6-7 non 5, 7-8 25, figs. 8-9
1968 Heyerodictyoconus americanus (CUSHMAN), 1980 Dictyoconus (Cushmania) puilboreauensis
Butterlin and Moullade, p. 12-13; Pl. 1, figs. 4-9 (WOODDRING), Hottinger and Drobne, p. 62,
1980 Dictyoconus (Cushmania) americanus 64; Pl. 1, fig. 2; Pl. 21, fig. 1-15; text fig. 9C;
(CUSHMAN), Hottinger and Drobne, p. 61-62; Pl. 12D
1, figs. 1, 4-9; Pl. 20, figs. 1-10; Text figs. 9A, B,
12C Samples. Brecciated Neiba Fm (La Descubierta 5871-
1993 Cushmania americana (CUSHMAN), Robinson I-IG HH 9020).
and Wright, p. 293, 295; Figs. 10.5-8; 11.2, Remarks. Robinson and Wright (1993) and Butterlin
11.4-5 and Moullade (1968) consider this form as synonymy
1999 Cushmania Americana (CUSHMAN), Blanco- of Cushmania americana. Following Hottinger and
Bustamante et al., Pl. 2, fig. 5 Drobne (1980) Cushmania puilboreauensis differs
from Cushmania americana because of the high-con-
Samples. Upper Neiba Fm (Bánica 5873-II-BR JS ical morphology of its test. When the test reaches a
9013, 9029 and 9030), Brecciated Neiba Fm (La diameter of 1.7 mm the axial section shows 14-17 pil-
Descubierta 5871-I-IG HH 9021), Catanamatías Fm lars in the last chamber.
(Arroyo Limón 5973-III-BR EB 9048). Distribution. This form occurs in the Brecciated Neiba
Remarks. The endoskeleton in Cushmania consists Fm associated with Amphistegina parvula and
of pillars (Pl. 1, figs. 12, 17, 18). The exoskeleton con- Fabiania cassis.
sists of beams and rafters, forming a subepidermal Age. According to Hottinger and Drobne (1980)
network in the marginal zone (Pl. 1, fig. 12). In mega- Cushmania puilboreauensis occurs in the Middle
lospheric forms the proloculus is positioned at the Eocene.
apex. In Cushmania americana the test has a low
conical form. When the test reaches a diameter of
3.5 mm the axial section shows 18-22 pillars in the Family SORITIDAE Ehrenberg, 1839
last chamber. Genus Praerhapydionina van Wessem, 1943
Distribution. Cushmania americana occurs in the Praerhapydionina cf. delicata HENSON, 1950
Brecciated Neiba Fm associated with Coleiconus elon- Pl. 1, figs. 21 and 22
gatus and Penoperculinoides cubensis. In the Upper
Neiba Fm this form occurs associated with 1963 Praerhapydionina delicata H ENSON ,
Amphistegina parvula, Fabiania cassis, Eodictyoconus Hottinger, p. 964-965; Pl. 1, fig. 3; Pl. 2, figs.
cubensis, Lepidocyclina (Lepidocyclina) macdonaldi, 1-10
Lepidocyclina (Pliolepidina) cf. peruviana and 1993 Praerhapydionina delicata HENSON, Robinson
Polylepidina antillea. Finally in the breccias of the and Wright, p. 303, 305; Figs. 15.1-6
Catanamatías Fm this form is associated with Fallotella
cookei, Fabiania cassis, Amphistegina parvula and Samples. Upper Neiba Fm (Jimaní 5871-III-IG JG
Lepidocyclina (Nephrolepidina) cf. chaperi. 9003).
366
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Remarks. We found two oblique sections where the Genus Annulosorites Hottinger, 2001
endoskeletal partitions could be seen. The dimen- Annulosorites spiralis HOTTINGER, 2001
sions of the uniserial chambers are similar to those Pl. 1, figs. 20 and 24
in Praerhapydionina delicata, but neither the pat-
tern of apertures nor the early growth stages can be 2001 Annulosorites spiralis n. sp., Hottinger, p. 489,
seen, and we prefer to maintain this form as confer. 491; Figs. 5.3-5.11, 6.1-6.5
Distribution. This form occurs associated with archa-
iasinids, peneroplids and Discorinopsis sp. in the Samples. Haitises Fm (Monte Plata 6272-III-IG HH
Upper Neiba Fm. 9037; Hato Mayor 6372-III-IG JG 9004), Madame Joie
Age. In Jamaica, according to Robinson and Wright Fm (Bánica 5873-II-BR JS 9017), Barahona Mb (Jimaní
(1993) and Robinson (1995), Praerhapydionina delica- 5871-III-IG JG 9002), Gurabo Fm (Pepillo Salcedo
ta occurs in beds of Oligocene age. 5875-II-EB 9002).
Remarks. Porcellaneous test with discoidal-biconvex
morphology. The growth changes from planispiral,
Genus Miosorites Seiglie and Rivera, 1976 evolute or involute, in the initial stages to annular.
Miosorites americanus (CUSHMAN, 1918) The annular chambers are subdivided by alternating
Pl. 1, figs. 26 and 29 partitions (septula) in radial position. The septula of
consecutive chambers are continuous in early stages
1929 Amphisorus americanus (CUSHMAN), Vaughan, and alternate later. The initial, spiral chambers show
p. 380-382; Pl. 41, fig. 5 only one row of apertures, whereas the annular
1977 Miosorites americanus (CUSHMAN), Seiglie et chambers have two rows. The diameter of the pro-
al., p. 867; Pl. 1, figs. 1-4, 5?, 6; Pl. 3, figs. 2?, loculus ranges 0.250-0.275 mm.
non Pl. 3, fig.1 Distribution. Annulosorites spiralis occurs in the
1981 Archaias (Archaias) floridanus (CONRAD), Haitises, Barahona, Madame Joie and Gurabo forma-
Butterlin, p. 133; Pl. 8, fig. 1 tions associated with Miosorites americanus,
2001 Miosorites americanus (CUSHMAN), Hottinger, Miarchaias floridanus and Sphaerogypsina globula.
491, 495, 499; Figs. 7-8, 9.1-9.4, 10.1-10.5 Age. According to Hottinger (2001) this species is
Late Miocene in age, but the presence of the
Operculinoides cf. bullbrooki in Gurabo Fm extends
Samples. Haitises Fm (Hato Mayor 6372-III-IG JG
down its range to the Middle Miocene (according to
9004), Madame Joie Fm (Bánica 5873-II-BR JS 9016,
Butterlin, 1981) or to the Early Miocene (according to
9017, 9018 and 9023), Barahona Mb (Jimaní 5871-III-
Caudri, 1996).
IG JG 9040), Gurabo Fm (Mao 5974-I-BR EB 9031).
Remarks. Porcellaneous test with discoidal biplanar-
biconvex morphology. The diameter of the test reaches
Genus Miarchaias Hottinger, 2001
1 cm. The annular chambers are evolute except for the
Miarchaias floridanus (CONRAD, 1846)
initial stage, where they may be involute. In each annu- Pl. 1, figs. 23, 25, 27 and 28
lar chamber the endoskeleton is made up of partitions
interpreted as septula (Hottinger, 2001), alternating in 2001 Miarchaias floridanus (CONRAD), Hottinger, p.
radial position from one chamber to the next. The inner 506; Figs. 19.1-19.5, 20.1-20.5, 21.1-21.6
chambers have two alternating apertures, the external
chambers present multiples apertures. The diameter of Samples. Haitises Fm (Monte Plata 6272-III-IG HH
the proloculus ranges 0.300-0.400 mm. 9037; Hato Mayor 6372-III-IG JG 9004), Madame Joie
Distribution. Miosorites americanus occurs in the Fm (Bánica 5873-II-BR JS 9016, 9017 and 9018),
Haitises, Barahona and Madame Joie formations Gurabo Fm (Pepillo Salcedo 5875-II-EB 9002),
associated with Miarchaias floridanus and Barahona Mb (Jimaní 5871-III-IG JG 9002 and 9040;
Annulosorites spiralis. In the Gurabo Fm it is associ- La Descubierta 5871-I-IG HH 9061).
ated with Sphaerogypsina globula, Cubanina sp. and Remarks. Porcellaneous test with planispiral-involute
Operculinoides cf. bullbrooki. growth. The test morphology, lenticular in the early
Age. According to Hottinger (2001) this species is Late chambers, changes to discoidal. The chambers pres-
Miocene in age, but the presence of the ent pillars in radial position and the exoskeleton con-
Operculinoides cf. bullbrooki in Gurabo Fm extends sists in short radial partitions (beams). There are mar-
down its range to the Early-Middle Miocene accord- ginal apertures between beams in the lateral walls
ing to Butterlin (1981) and Caudri (1996). (Hottinger, 2001). The diameter of the test for 2 whorls
367
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
is 1.25-1.60 mm, for 3 whorls is 1.7-2.0 mm. The pro- ciated with Fallotella cookei, Cushmania americana,
loculus is about 0.250 mm in diameter. Amphistegina parvula and Lepidocyclina
Distribution. Miarchaias floridanus occurs associated (Nephrolepidina) cf. chaperi.
with Miosorites americanus, Annulosorites spiralis Age. According to Butterlin (1981) and Robinson and
and Sphaerogypsina globula in the follow lithostrati- Wright (1993) the range of this species is from Middle
graphic units: Haitises, Barahona, Madame Joie and Eocene to Late Eocene in age.
Gurabo formations.
Age. According to Hottinger (2001) this species is Late
Miocene in age, but the presence of the Genus Eodictyoconus COLE and BERMÚDEZ, 1944
Operculinoides cf. bullbrooki in Gurabo Fm extends Eodictyoconus cubensis (CUSHMAN and BERMÚDEZ,
down its range to the Middle Miocene (according to 1944)
Butterlin, 1981) or to the Early Miocene (according to Pl. 2, figs. 3, 4 and 5
Caudri, 1996).
1944 Eodictyoconus cubensis (CUSHMAN and
BERMÚDEZ), Cole and Bermúdez, p. 6-10; Pl. 1,
Family CYMBALOPORIDAE Cushman, 1927
fig.1; Pl. 2, figs. 1-12; Pl. 3, figs. 1, 4-5
Genus Fabiania Silvestri, 1924
Samples. Upper Neiba Fm (Bánica 5873-II-BR JS
Fabiania cassis (OPPENHEIM, 1896)
9029), Catanamatías Fm (Bánica 5873-II-BR JS 9009).
Pl. 2, figs. 1 and 2
Remarks. According to Robinson and Wright (1993)
Eodictyoconus cubensis is a synonymy of Fabiania
1944 Pseudorbitolina cubensis CUSHMAN and
cassis. We prefer to maintain it as a different genera
BERMÚDEZ, Cole, p. 35-36; Pl. 13, figs. 1-2
and species because it can be easily distinguished by
1944 Eodictyoconus cubensis (CUSHMAN and
its thicker wall and more simple exo- and endoskele-
BERMÚDEZ), Cole and Bermúdez, Pl. 3, figs. 2-3
tal structures.
1981 Fabiania cassis (OPPENHEIM), Butterlin, p. 35-36;
Pl. 13, figs. 1-3 Distribution. Eodictyoconus cubensis occurs in the
1993 Fabiania cassis (OPPENHEIM), Robinson and Upper Neiba Fm associated with Fallotella cookei,
Wright, p. 307; Figs. 17. 1, 17. 3-6 Fabiania cassis and Polylepidina antillea. In the brec-
cias of the Cantanamatías Fm it is associated with
Samples. Lower Neiba Fm (Bánica 5873-II-BR JS Victoriella conoidea, Lepidocyclina (Lepidocyclina)
9026), Brecciated Neiba Fm (La Descubierta 5871-I-IG macdonaldi, Lepidocyclina (Nephrolepidina) cf.
HH 9038), Upper Neiba Fm (Bánica 5873-II-BR JS chaperi and Polylepidina antillea.
9013, 9029 and 9030), Catanamatías Fm (Arroyo Age. According to Robinson and Wright (1993)
Limón 5973-III-BR EB 9048), Rio Yabón Fm (Miches Eodictyoconus cubensis is a synonymy of Fabiania
6372-I-IG AD 9016). cassis and occurs from Middle to Late Eocene in
Remarks. Typified by having a conical test with thin Jamaica.
wall, and with exo- and endoskeleton structures with-
in each chamber. See Deloffre and Hamaoui (1973) for
an extended description. Fabianiinae indeterminated
Distribution. Fabiania cassis occurs in Lower Neiba Pl. 2, figs. 6 and 7
Fm associated with Eoconuloides wellsi. In the
Brecciated Neiba Fm it is associated with Coleiconus Samples. Las Lavas Fm (Monte Cristi 5875-I-EB 9010),
elongatus, Amphistegina parvula, Cushmania puil- Monte Cristi Fm (Monte Cristi 5875-I-EB 9012),
boreauensis and Eoconuloides lopeztrigoi. In the Barahona Mb (La Descubierta 5871-I-IG HH 9009 and
Upper Neiba Fm it is associated with Amphistegina 9061).
parvula, Cushmania americana, Fallotella cookei, Remarks. Hyaline attached form. It is similar to
Polylepidina antillea, Eodictyoconus cubensis, Gunteria floridana, defined by Cushman and Ponton
Lepidocyclina (Pliolepidina) cf. peruviana and (1933) in Eocene rocks of Cuba, but is structurally
Lepidocyclina (Lepidocyclina) macdonaldi. In the Rio more complex. We found only three sections that do
Yabón Fm it is associated with Fallotella cookei, not allow observing the chamber arrangement. The
Amphistegina parvula, Eoconuloides lopeztrigoi and wall is perforated and has exoskeletal structures,
Lepidocyclina (Lepidocyclina) macdonaldi. Finally in probably beams.
the breccias of the Catanamatías Fm this form is asso- Distribution. This form occurs in the Barahona Mb
368
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369
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
370
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
Family BORELOIDIDAE Reiss, 1963 ventral sides. The diameter ranges from 1.15 to 1.40
Genus Eoconuloides Cole and Bermúdez, 1944 mm and the height from 0.625 to 0.825 mm.
Eoconuloides lopeztrigoi (PALMER, 1934) Distribution. Eoconuloides lopeztrigoi occurs in the
Pl. 2, figs. 16, and 17; Pl. 3, fig. 3 Brecciated Neiba Fm associated with Coleiconus
elongatus, Fabiania cassis and Amphistegina parvula;
1934 Amphistegina lopeztrigoi n. sp., Palmer, p. this form occurs also in the Lower Neiba Fm associ-
255; Pl. 15, figs. 6, 8 ated with Eoconuloides wellsi. In the Upper Neiba Fm
1936 Amphistegina lopeztrigoi PALMER, Baker and this species is associated with Coleiconus elongatus,
Grimsdale, p. 233; Pl. 32, figs. 1-3 Fallotella cookei, Amphistegina parvula and
1944 Amphistegina lopeztrigoi PALMER, Cole, p. 55; Discocyclina marginata. In the Rio Yabón Fm it is
Pl. 1, fig. 17; Pl. 8, fig. 16; Pl. 9, figs. 10-13 associated with Fallotella cookei, Fabiania cassis,
1952 Amphistegina lopeztrigoi PALMER, Cole and Amphistegina parvula and Lepidocyclina
Gravell, p. 714; Pl. 91, figs. 6-8 (Lepidocyclina) macdonaldi. In the Loma Peñón Fm
1957 Amphistegina lopeztrigoi PALMER, Levin, p. this form is associated with Discocyclina marginata
146-147; Pl. 4, figs. 8-9, 13-14 and Eoconuloides wellsi and also with reworked
1981 Eoconuloides lopeztrigoi (PALMER), Butterlin, Paleocene larger foraminifers such as Ranikothalia
Pl. 15, figs. 9-10 bermudezi and Fallotella sp.
1993 Eoconuloides lopeztrigoi (PALMER), Robinson Age. According to Robinson and Wright (1993) and
and Wright, p. 311; Figs. 19.6-9 Blanco-Bustamante et al. (1999) Eoconuloides
1996 Eoconuloides lopeztrigoi (PALMER), Caudri, p. lopeztrigoi occurs from the Late Paleocene to the
1229-1230; Pl. 27, fig. 1 Middle Eocene.
2003 Amphistegina lopeztrigoi PALMER, De Mello e
Sousa et al., Pl. 3, fig. 4
Eoconuloides wellsi COLE and BERMÚDEZ, 1944
Samples. Lower Neiba Fm (Galván 5971-IV-IG HH Pl. 2, figs. 13, and 14
9005 and 9032), Upper Neiba Fm (Galván 5971-IV-IG
HH 9030), Brecciated Neiba Fm (La Descubierta 5871- 1944 Eoconuloides wellsi n. gen. n. sp., Cole and
I-IG HH 9038), Rio Yabón Fm (Miches 6372-I-IG AD Bermúdez, p. 11; Pl. 1, figs. 4-10
9016), Loma Peñón Fm (Rincón Chavón 6472-III-IG JG 1952 Eoconuloides wellsi COLE and BERMÚDEZ, Cole
9002 and 9023). and Gravell, p. 713; Pl. 92, figs. 1-10
Remarks. This form has a lenticular test with low-tro- 1981 Eoconuloides wellsi COLE and BERMÚDEZ,
chospiral growth and strong piles in both dorsal and Butterlin, Pl. 17, figs. 3-4
Plate 2. Systematics II
Plancha 2. Sistemática II
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1993 Eoconuloides wellsi COLE and BERMÚDEZ, Samples. Upper Neiba Fm (Bánica 5873-II-BR JS
Robinson and Wright, Figs. 18.1-2, 19.1-5 9020, 9048 and 9049).
1999 Eoconuloides wellsi COLE and BERMÚDEZ, Remarks. This species shows a considerable variabil-
Blanco-Bustamante et al., Pl. 2, fig. 4 ity in size and shape, with or without piles. It is char-
acterised by a thick walled isolepidine embryo, of
Samples. Lower Neiba Fm (Galván 5971-IV-IG HH 150-300 µm, with two embryonic chambers of a simi-
9032; Bánica 5873-II-BR JS 9026), Loma Peñón Fm lar size, separated by a thinner flat wall. After study-
(Rincón Chavón 6472-III-IG JG 9023). ing the variability of this species, some authors rec-
Remarks. This form varies from high conical to low ognized many synonymies. Thus, for example, Cole
conical trochospiral growth with flat base and sub- (1957a, 1961a) considered that Lepidocyclina
acuate peripheral angle. In the ventral side it has (Lepidocyclina) giraudi R. DOUVILLÉ 1907,
short piles. The diameter of the test ranges from 1.60 Lepidocyclina miraflorensis VAUGHAN 1923,
to 1.90 mm and its height from 1.0 to 1.3 mm Lepidocyclina asterodisca NUTALL 1932, and
Distribution. Eoconuloides wellsi occurs in the Loma Lepidocyclina (Lepidocyclina) waylandvaughani COLE
Peñón Fm associated with Eoconuloides lopeztrigoi 1928, are junior synonyms of Lepidocyclina
and Discocyclina marginata and Neodiscocyclina (Lepidocyclina) canellei. See Cole (1961a) for an
barkeri. In the Lower Neiba Fm this species is asso- extended description and figuration of this species.
ciated with Eoconuloides lopeztrigoi and Fabiania Distribution. This form occurs in the Upper Neiba Fm
cassis. associated with Lepidocyclina (Eulepidina) undosa
Age. According to Blanco-Bustamante et al. (1999) and Nummulites cf. panamensis.
this species occurs in Cuba from the Morozovella sub- Age. According to Butterlin (1981, 1984), Robinson
botinae plancktonic foraminiferal biozone (P 6) of and Wright (1993), Caudri (1996), De Mello e Sousa et
Berggren et al. (1995) through the Morozovella arago- al. (2003) the range of this species is Oligocene to
nensis biozone (P 8), thus being Early Eocene in age. Early Miocene.
According to Robinson and Wrigth (1993) in Jamaica
Eoconuloides wellsi ranges from the latest Early
Eocene to the early Middle Eocene. Lepidocyclina (Lepidocyclina) macdonaldi CUSHMAN, 1919
Pl. 2, fig. 19
Family LEPIDOCYCLINIDAE Scheffen, 1932
1944 Lepidocyclina (Pliolepidina) ariana COLE and
Genus Lepidocyclina Gümbel, 1970
PONTON, Cole, p. 61; Pl. 1, fig. 14; Pl. 14, figs. 1-
Subgenus Lepidocyclina Gümbel, 1970
8; Pl. 16, figs. 11-13; Pl. 17, fig. 13.
Lepidocyclina (Lepidocyclina) canellei LEMOINE and R.
1974 Lepidocyclina (Lepidocyclina) macdonaldi
DOUVILLÉ, 1904
CUSHMAN, Frost and Langenheim, p. 143; Pl.
Pl. 2, figs. 20 and 21
38, figs 1-9; Pl. 39, figs. 1-7; Pl. 40, figs. 1-13;
Pl. 41, figs. 1-4; Pl. 42, figs. 1-6
1933 Lepidocyclina (Lepidocyclina) canellei,
1981 Lepidocyclina (Neolepidina) macdonaldi
LEMOINE and R. DOUVILLÉ, Vaughan, p. 14; Pl. 6,
CUSHMAN, Butterlin, p. 72, 75; Pl. 47, figs. 1-2
figs. 1-5
1935 Lepidocyclina canellei LEMOINE and R. 1981 Lepidocyclina (Neolepidina) ariana COLE and
DOUVILLÉ, van de Geyn and van der Vlerk, p. PONTON, Butterlin, p. 72, 75; Pl. 47, figs. 3-5
230, figs. 43-45 1990 Lepidocyclina macdonaldi CUSHMAN, Butterlin,
1961a Lepidocyclina (Lepidocyclina) canellei, Pl. 3, figs.3-6
LEMOINE and R. DOUVILLÉ, Cole, Pl. 30, figs. 1-13; 1993 Lepidocyclina (Lepidocyclina) macdonaldi
Pl. 31, figs. 1-5; Pl. 32, figs. 1-4; Pl. 33, figs. 1- CUSHMAN, Robinson and Wright, p. 317; Figs.
4; Pl. 34, figs. 1-8; Pl. 35, figs. 1, 2, 4, 5; Pl. 36, 23.5-7
figs. 1-5; Pl. 37, figs. 1-5; Pl. 38, figs. 1-7
1968 Lepidocyclina (Lepidocyclina) canellei Samples. Upper Neiba Fm (Arroyo Limón 5973-III-BR
LEMOINE and R. DOUVILLÉ, Eames et al., p. 297; EB 9053; Bánica 5873-II-BR JS 9030), Catanamatías
Pl. 59, fig. 7 Fm (Bánica 5873-II-BR JS 9009), Rio Yabón Fm
1996 Lepidocyclina (Lepidocyclina) canellei LEMOINE (Miches 6372-I-IG AD 9016).
and R. DOUVILLÉ, Caudri, p. 1227 ; Pl. 18, figs. 1-2 Remarks. Lepidocyclina (Lepidocyclina) macdonaldi
2003 Lepidocyclina (Lepidocyclina) canellei LEMOINE is very similar to Lepidocyclina ariana COLE and
and R. DOUVILLÉ, De Mello e Sousa et al., Pl. 2, PONTON 1934. Cole (1944, 1945) thought that the two
figs. 9-10 species could be synonymous, although latter (Cole,
372
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
1956) he considered them as two valid species. and Neolepidina BRÖNNIMANN 1947. Both genera have
Butterlin (1981) remarks that this two species are been discussed and considered to be synonyms of
sometimes difficult to distinguish, but considers them Lepidocyclina by different authors. Pliolepidina is
to be two separate species, statistically different, that characterized by a megalospheric embryo subdivided
he called Lepidocyclina (Neolepidina) ariana and in three or more chambers, with no further differ-
Lepidocyclina (Lepidocyclina) macdonaldi. ences from Lepidocyclina. Some authors (e.g.
However, after studying more than 600 specimens, Vaughan and Cole, 1940, 1941; Butterlin, 1981) con-
Frost and Langenheim (1974) concluded that sidered that the type species of Pliolepidina H.
Lepidocyclina (Lepidocyclina) ariana, and DOUVILLÉ 1915, Lepidocyclina (Pliolepidina) tobleri H.
Lepidocyclina (Pliolepidina) proteiformis VAUGHAN DOUVILLÉ 1917, as a teratological form of
1924, are synonyms of Lepidocyclina (Lepidocyclina) Lepidocyclina pustulosa H. DOUVILLÉ 1917, and thus
macdonaldi, because they found a morphological that Pliolepidina was not a valid genus. This caused
gradation between these three forms (even in a single Brönnimann (1947, p. 376) to consider the genus
specimen) in size, number and arrangement of lateral Pliolepidina as “nomen caducum”, proposing anoth-
chamberlets and piles. Robinson and Wright (1993) er new genus, Neolepidina, with Lepidocyclina pustu-
reached the same conclusion. Both Frost and losa as the type species. Vaughan and Cole (1941) put
Langenheim (1974) and Robinson and Wright (1993) Lepidocyclina trinitatis H. DOUVILLÉ 1924 in the syn-
related this variability to the environmental condi- onymy of Lepidocyclina pustulosa, which is the type
tions from observations of the facies where they find species of Isorbitoina THALMANN 1938, for what,
the specimens, in Chiapas and Jamaica respectively. according to Grimsdale and van der Vlerk (1959),
Thus, the specimens of the types “ariana” and “mac- Neolepidina is a synonym of Isorbitoina, which was
donaldi s.s.”, with low lateral chamberlets with thick proposed as a subgenus of Orbitoina, a subjective
walls, tend to predominate in populations associated synonym of Lepidocyclina (Eames et al, 1962).
to shelf carbonates, particularly algal-foraminiferal Eames et al. (1962) regarded Neolepidina a junior
biosparites, whereas the specimens of the type “pro- synonym of Lepidocyclina (s.s) because the “four
teiformis”, with high lateral chamberlets with thin stolon system” which characterizes this genus is
walls, are most common in basin-slope facies, with found, together with the “six stolon system”, in dif-
planktonic and benthonic foraminifers, in biomicrite ferent ontogenetic stages of the same specimens of
or calcareous clays or shales. According to Frost and some Lepidocyclina species.
Langenheim (1974), the depositional environment Following Cole (1960a, 1961b, 1962, 1963) multi-
would affect basically the morphology of the lateral locular embryos (which characterize the genera
chamberlets and the thickness of the walls, affecting Pliolepidina and Multilepidina) are due to irregulari-
very few the size or number of piles and nothing the ties of the reproductive cycle, sometimes are found
features of the embryonic chambers. together with bilocular embryos, and thus cannot be
Following Cole (1944, p.62) Lepidocyclina used to characterize genera. However, Eames et al.
(Lepidocyclina) claibornensis GRAVELL and HANNA (1962) claim that Pliolepidina is “in no way monstrous
1940, is a synonym of Lepidocyclina (Pliolepina) ari- or teratoid” and consider it a valid subgenus of
ana COLE and PONTON 1934, for what the former would Lepidocyclina.
be another synonym of Lepidocyclina (Lepidocyclina) Distribution. Lepidocyclina (Lepidocyclina) macdonaldi
macdonaldi. occurs in the Upper Neiba Fm associated with
Following Frost and Langenheim (1974) the group Fallotella cookei, Cushmania americana, Fabiania cas-
of Lepidocyclina (Lepidocyclina) macdonaldi-ariana- sis, Amphistegina parvula, Nummulites cf. wilcoxi and
proteiformis is a species lineage. It is morphological- Lepidocyclina (Pliolepidina) cf. peruviana. In the brec-
ly similar to the Lepidocyclina ocalana-montgome- cias of the Cantanamatías Fm it is associated with
riensis-gubernacula lineage, both groups having Eodictyoconus cubensis, Victoriella conoidea,
isolepidine embryos. They differ in the shape of the Polylepidina antillea and Lepidocyclina
equatorial chamberlets (arcuate to short-spatulate in (Nephrolepidina) cf. chaperi. Finally in the Rio Yabón
the Lepidocyclina (Lepidocyclina) macdonaldi group; Fm it is associated with Fallotella cookei, Amphistegina
rhombic, spatulate, or hexagonal in the Lepidocyclina parvula, Eoconuloides lopeztrigoi and Fabiania cassis.
ocalana group, and in the lateral chamberlets (with Age. According to Butterlin (1963, 1984), Butterlin and
fewer layers and smaller chamberlets not arranged in Moullade (1968), Frost and Langenheim (1974),
tiers in the Lepidocyclina ocalana group). Robinson and Wright (1993) and De Mello e Sousa et
Remarks-ariana. Lepidocyclina ariana has also been al. (2003) this species ranges from the Middle Eocene
assigned to the genera Pliolepidina H. DOUVILLÉ 1915 to the Late Eocene.
373
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Lepidocyclina (Lepidocyclina) mantelli MORTON, 1883 1974 Lepidocyclina (Eulepidina) undosa CUSHMAN,
Pl. 2, fig. 18 Frost and Langenheim, p. 165; Pl. 46, figs. 1, 2;
Pl 47, figs. 1-7; Pl. 48, figs. 1-4; Pl. 49, figs. 1-4;
1935 Lepidocyclina mantelli (MORTON) Lemoine and Pl. 50, figs. 1-4
R. Douvillé, van de Geyn and van der Vlerk, p. 1993 Lepidocyclina (Eulepidina) undosa CUSHMAN,
237; Figs. 37 (after Vaughan 1928), 38 Robinson and Wright, p. 323, 325; Figs. 22.1-
1944 Lepidocyclina (Lepidocyclina) mantelli 2
(MORTON), Cole, p. 70; Pl. 22, figs. 13-15 1996 Lepidocyclina undosa CUSHMAN, Caudri, p.
1957a Lepidocyclina (Lepidocyclina) mantelli 1224; Pl. 20, fig. 1; Pl. 21, fig. 1
(MORTON), Cole, p. 38-40 ; Pl. 1, figs. 1-5, 7-9 ; 1996 Lepidocyclina favosa CUSHMAN, Caudri, p.
Pl. 2, figs. 1, 6 ; Pl. 3, figs. 1-4 ; Pl. 4, figs. 1, 4, 1224; Pl. 20, figs. 2, 3; Pl. 21, fig. 2
6-7 ; Pl. 5, figs. 3-5 ; Pl. 6, figs. 3-7
1974 Lepidocyclina (Lepidocyclina) mantelli Samples. Upper Neiba Fm (Arroyo Limón 5973-III-BR
(MORTON), Frost and Langenheim, p. 15 ; Pl. EB 9047, 9049 and 9050; Bánica 5873-II-BR JS 9022
43, figs. 1-9; Pl. 44, figs. 1-6 and 9048; Jimaní 5871-III-IG JG 9007), Brecciated
Samples. Upper Neiba Fm (Bánica 5873-II-BR JS Neiba Fm (La Descubierta 5871-I-IG HH 9057, 9062,
9022), Brecciated Neiba Fm (La Descubierta 5871-I-IG 9064 and 9065), Sombrerito Fm (Bánica 5873-II-BR JS
HH 9057), Sombrerito Fm (Bánica 5873-II-BR JS 9008), 9008 and 9010) Barahona Mb (Jimaní 5871-III-IG JG
Catanamatías Fm (Bánica 5873-II-BR JS 9010), 9009).
Barahona Mb (Jimaní 5871-III-IG JG 9009). Remarks. Lepidocyclina (Eulepidina) undosa and
Remarks. Large, flat species, with poorly developed Lepidocyclina favosa, two species defined by
or without piles. The megalospheric embryo is simi- Cushman (1919, pp. 65-66), are always found
lar to that of Lepidocyclina (Lepidocyclina) canellei together and show a complete morphological gra-
(two thick-walled chambers separated by a flat thin- dation. They are considered as morphotypes of the
ner wall), but of larger dimensions, of about half a same species, Lepidocyclina (Eulepidina) undosa
millimetre. See Frost and Langenheim (1974, p.155- (Vaughan, 1928; Cole, 1952; Cole y Applin, 1961 -
160) for a detailed description of this species. who formally defined Lepidocyclina favosa as a
Distribution. Lepidocyclina (Lepidocyclina) mantelli synonym- Frost and Langenheim 1974; Caudri,
occurs in the Brecciated Neiba Fm. In the Upper 1996). The morphotype “favosa” has a more inflat-
Neiba Fm it is associated with Lepidocyclina ed test, with a higher number of lateral chamber-
(Eulepidina) undosa and Nummulites cf. panamen- lets.
sis and in the Catanamatías Fm and Barahona Mb it Similarly, Lepidocyclina gigas CUSHMAN 1919 is
is associated with Lepidocyclina (Eulepidina) found always associated to Lepidocyclina undosa and
undosa. it is considered its microspheric form (Vaughan, 1924,
Age. Some authors such as Eames et al. (1968) and 1928; Cole, 1952; Frost and Langenheim 1974; Caudri,
Cole (1957c) place it in Oligocene, while others 1996), and thus, another synonym (Sachs and
(Butterlin, 1981, 1984; Frost and Langenheim 1974) Gordon, 1962). Only few authors (e.g. Eames et al.
extend its range up to the Early Miocene. 1968) consider them three different species.
Frost and Langenheim (1974) described a com-
plete gradation between the two types, which they
Subgenus Eulepidina H. Douvillé, 1911 considered related to the environment. They find the
Lepidocyclina (Eulepidina) undosa CUSHMAN, 1919 “undosa” type in reef facies and the “favosa” type
Pl. 2, figs. 22, and 23 in inter-reef or back-reef facies. Following Frost and
Langenheim (1974) the “undosa” type has few (7-8)
1933 Lepidocyclina (Eulepidina) favosa CUSHMAN, layers of lateral chamberlets and reduced embryo
Vaughan, p. 37; Pl. 19, figs. 1, 2; Pl 20, figs. 1- dimensions (diameter of the deuteroconch between
3 500 and 875 µm), whereas the “favosa” type has a
1968 Eulepidina undosa (CUSHMAN), Eames et al., p. higher number of lateral layers (12-15), larger
285; Pl. 49, figs. 6, 7 (topotypes) embryos (diameter of the deuteroconch between
1968 Eulepidina favosa (CUSHMAN), Eames et al., p. 975 and 1500 µm), a larger test size, larger equatori-
296; Pl. 57, fig. 6 al chamberlets, and a selliform morphology. In their
1968 Eulepidina undosa (C USHMAN ) nov. subsp. Text-figure 24, a plot of embryo dimensions, two
laramblaensis, Eames et al., p. 296; Pl. 57, figs. groups (the inter-reef facies “favosa” type and the
7, 8 reef facies “undosa” type) are clearly differentiated,
374
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
but the supposed “complete morphologic grada- Fabiania cassis, Amphistegina parvula, Eodictyoconus
tion” is not so evident because they do not overlap, cubensis, Victoriella conoidea, Polylepidina antillea and
and the existence of two separate species cannot be Lepidocyclina (Lepidocyclina) macdonaldi. In the Upper
discarded. Neiba Fm it is associated with Fallotella cookei.
Distribution. In the Brecciated Neiba Fm Age. The age of this form is Late Eocene according to
Lepidocyclina (Eulepidina) undosa occurs with Cole (1963) Butterlin (1963, 1981), Butterlin and
Lepidocyclina (Lepidocyclina) mantelli. In the Upper Moullade (1968) and Robinson and Wright (1993), also
Neiba Fm it is associated with Lepidocyclina latest Middle Eocene according to Frost and
(Lepidocyclina) mantelli and Nummulites cf. pana- Langenheim (1974) and De Mello e Sousa et al. (2003).
mensis. In the Barahona Mb it is associated with
Lepidocyclina (Lepidocyclina) mantelli.
Age. Authors such as Vaughan (1933) Butterlin (1963) Lepidocyclina (Nephrolepidina) tournoueri LEMOINE
and Caudri (1996) place it in Oligocene. Others as and R. DOUVILLÉ, 1904
Cole and Applin (1961), Eames et al. (1968), Butterlin
(1981, 1984), Frost and Langenheim (1974), Robinson 1933 Lepidocyclina (Nephrolepidina) tournoueri
and Wright (1993) and De Mello e Sousa et al. (2003) LEMOINE and R. DOUVILLÉ, Vaughan, p. 25; Pl. 13,
extend its range up to the Early Miocene. figs. 1, 2
1961a Lepidocyclina (Eulepidina) tournoueri LEMOINE
and R. DOUVILLÉ, Cole, Pl. 32, fig. 5; Pl. 34, fig.
Subgenus Nephrolepidina H. Douvillé, 1911 9; Pl. 35, fig. 3
Lepidocyclina (Nephrolepidina) cf. chaperi LEMOINE 1981 Lepidocyclina (Nephrolepidina) tournoueri
and R. DOUVILLÉ, 1904 Lemoine and R. Douvillé, Butterlin, p. 73-75 ;
Pl. 2, figs. 24 and 25 Pl. 51, figs. 1, 2, 4
1952 Lepidocyclina (Nephrolepidina) chaperi Samples. Upper Neiba Fm (Arroyo Limón 5973-III-BR
LEMOINE and R. DOUVILLÉ, Cole, Pl. 8, figs. 5-8;
EB 9054), Sombrerito Fm (Bánica 5873-II-BR JS 9010).
Pl. 9, figs. 3-19; Pl. 10, figs. 1-10; Pl. 11, figs. 1-
Remarks. It is similar to small morphotypes of
8; Pl. 12, figs 3-10; Pl. 20, figs. 8-10; Pl. 23, figs.
Lepidocyclina (Lepidocyclina) canellei, because it has
11-12
similar morphology, equatorial section and an embryo
1974 Lepidocyclina (Nephrolepidina) chaperi
of similar size. It can be distinguished form this species
LEMOINE and R. DOUVILLÉ, Frost and
by the embryo, of nephrolepidine configuration, with a
Langenheim, p. 161-165; Pl.45, figs. 1-6; Pl. 46,
spherical protoconch embraced by a kidney-shaped
figs. 3, 4
1981 Lepidocyclina (Nephrolepidina) chaperi deuteroconch, whereas Lepidocyclina (Lepidocyclina)
LEMOINE and R. DOUVILLÉ, Butterlin, p. 73, 75; Pl. canellei has two embryonic chambers of a similar size
50, figs. 1-5 separated by a flat wall. Another characteristic of
1993 Lepidocyclina (Nephrolepidina) chaperi Lepidocyclina (Nephrolepidina) tournoueri is the ten-
LEMOINE and R. DOUVILLÉ, Robinson and Wright, dency to form wavy to stellate annuli. According to
p. 321; Figs. 22.5; 25.1-5 Cole (1961a, p. 388) the stellate pattern of annuli is an
intraspecific species, and as long it is the only charac-
Samples. Upper Neiba Fm (Arroyo Limón 5973-III-BR ter that differentiates “Lepidocyclina (Eulepidina)
EB 9052), Catanamatías Fm (Arroyo Limón 5973-III-BR tournoueri” from Lepidocyclina (Eulepidina) dartoni
EB 9048; Bánica 5873-II-BR JS 9009). VAUGHAN 1933, this latter species must be considered
Remarks. This species has been included in the sub- as a junior synonym of the first.
genera Eulepidina (e.g. Cole, 1963) and Distribution. Lepidocyclina (Nephrolepidina) tournoueri
Nephrolepidina (e.g. Butterlin, 1981). See Cole (1952), occurs in the Upper Neiba. In the Sombrerito Fm it is
Frost and Langenheim (1974), and Robinson and associated with Lepidocyclina (Eulepidina) undosa and
Wright (1993) for detailed description, figuration and Lepidocyclina (Lepidocyclina) mantelli.
discussion. The specimens from Sombrerito Fm Age. According to Vaughan (1933), Van de Geyn and
(Bánica 9009) are larger than those from Neiba van der Vlerk (1935), Cole (1952), and Eames et al.
Superior Fm (Arroyo Limón 9052). (1968) this form is Oligocene in age. Other authors
Distribution. Lepidocyclina (Nephrolepidina) cf. chaperi such as Cole (1961a) and Butterlin (1984) extend its
occurs in the breccias of the Cantanamatías Fm associ- range up to the Early Miocene, or also to the Middle
ated with Fallotella cookei, Cushmania americana, Miocene (Butterlin, 1981).
375
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Subgenus Pliolepidina H. Douvillé, 1915 Cushman, Frost and Langenheim, p. 121, Pl.
Lepidocyclina (Pliolepidina) cf. peruviana CUSHMAN, 29, figs. 1-13; Pl 30, figs. 1-6; Pl. 31, figs. 1-6
1922
Pl. 2, figs. 26 Samples. Upper Neiba Fm (Bánica 5873-II-BR JS
9029), Catanamatías Fm (Bánica 5873-II-BR JS 9009).
1937 Lepidocyclina (Pliolepidina) peruviana Remarks. Cole (1944) considers as synonyms of
CUSHMAN, Vaughan, Pl. 117, figs. 1-3; Pl. 118, Lepidocyclina (Polylepidina) antillea (defined in St
figs. 2-8; Pl. 119, figs. 1-6; Pl. 120, figs. 1-4 Bartholomew) the following species: adkinsi, chia-
1944 Lepidocyclina (Pliolepidina) peruviana pasensis, chiapasensis var. subplana (from Mexico),
CUSHMAN, Cole, p. 62; Pl. 15, figs. 11-10 kinlossensis (from Jamaica), and gardnerae (from
1975 Lepidocyclina peruviana CUSHMAN, Caudri, p. Texas, Louisiana, Mississippi and Alabama), to which
573-574; Pl. 25, figs. 5-14; Pl. 26, figs. 1-14 he adds later (Cole 1960a) Eulinderina guayabalensis
1996 Lepidocyclina peruviana CUSHMAN, Caudri, p. (NUTALL 1930) and Eulinderina semiradiata BARKER and
1217; Pl. 6, fig. 12; Pl. 15, figs. 4, 6 GRIMSDALE 1936. Robinson and Wright (1993) consider
Samples. Upper Neiba Fm (Bánica 5873-II-BR JS Eulinderina antillea (CUSHMAN 1919) and Polylepidina
9030). chiapasensis VAUGHAN 1924, to be two different
Remarks. This species is difficult to distinguish from species, differentiated by the kind of initial growth.
Lepidocyclina (Lepidocyclina) pustulosa in axial sec- The forms figured by Frost and Langenheim (1974)
tion, both species have the same stratigraphic range are compressed with few lateral chamberlets.
and have been put in synonymy by some authors However, studying topotypes of Lepidocyclina
(e.g. Cole, 1963). This species has been assigned to (Polylepidina) antillea and their variability in a single
the subgenus Pliolepidina by some authors (e.g. population (“in the number of periembryonic cham-
Vaughan, 1937; Cole, 1944). For a discussion on the bers, the size of these chambers, the length of the
genus Pliolepidina see Remarks in Lepidocyclina periembryonic coil and the thickness of the revolving
(Lepidocyclina) macdonaldi. wall adjacent to the embryonic chambers”), Cole
Distribution. This form occurs in the Upper Neiba Fm (1960a) concluded that the genus Eulinderina (Barker
associated with Cushmania americana, Fallotella and Grimsdale 1936) is a synonym of Polylepidina.
cookei, Amphistegina parvula, Fabiania cassis and Distribution. Polylepidina antillea occurs in the Upper
Lepidocyclina (Lepidocyclina) macdonaldi. Neiba Fm associated with Fallotella cookei,
Age. According to Cole (1944), Klugler and Caudri Eodictyoconus cubensis, Fabiania cassis and
(1975), Caudri (1975) this species occurs in the Late Polylepidina antillea. In the breccias of the
Eocene. Caudri (1996) extends its range to the latest Cantanamatías Fm it is associated with Eodictyoconus
Middle Eocene. cubensis, Victoriella conoidea, Lepidocyclina
(Nephrolepidina) cf. chaperi, Lepidocyclina
(Lepidocyclina) macdonaldi and Polylepidina antillea.
Genus Polylepidina Vaughan, 1924 Age. According to Cole (1944, 1956), Butterlin and Moullade
Polylepidina antillea VAUGHAN, 1924 (1968), Butterlin (1981) and Frost and Langenheim (1974)
Pl. 3, fig. 1 this species occurs in the Middle Eocene.
376
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
diameter the proloculus is of about 0.60 mm, and the 1966 Camerina catenula (CUSHMAN and JARVIS),
diameter of the test is about 1 mm for three whorls. Cole, Pl. 23, fig. 5; Pl. 26, figs. 1, 6
Distribution. Penoperculinoides cubensis occurs in 1981 Ranikothalia bermudezi (PALMER), Butterlin, p.
the Brecciated Neiba Fm. 30, 33; Pl. 9, figs. 1-3
Age. According to Cole and Gravell (1952) and 1993 Ranikothalia catenula (CUSHMAN and JARVIS),
Butterlin (1981) this form occurs in the Middle Eocene. Robinson and Wright, p. 329-331; Figs 28.1-7
1997 Ranikothalia bermudezi (PALMER), Berlanga, p.
136-144; Pl. 28, figs. 1-6; Pl. 29, figs. 1-5
Family MIOGYPSINIIDAE Vaughan, 1928 1999 Ranikothalia catenula (CUSHMAN and JARVIS),
Genus Miogypsina Sacco, 1893 Blanco-Bustamante et al., Pl. 1, figs. 1-3
Miogypsina cf. antillea (CUSHMAN, 1919) Samples. Don Juan Fm (Monte Plata 6272-III-IG HH 9008),
Pl. 3, fig. 4 Loma Peñón Fm (Rincón Chavón 6472-III-IG JG 9002).
Remarks. Following Berlanga (1997), Ranikothalia
1957b Miogypsina (Miogypsina) antillea CUSHMAN,
bermudezi is characterized by a lenticular robust test
Cole, p. 320-321; Pl. 26, figs. 6-7; Pl. 28, figs. 1-
with a great marginal cord and some piles in the
9; Pl. 29, figs. 1-9
umbonal zone. The diameter of the megalospheric
1961 Miogypsina (Miogypsina) antillea CUSHMAN,
test ranges from 1.90 to 2.75 mm and its height
Cole and Applin, p. 133; Pl. 7, figs. 1-7
1961b Miogypsina (Miogypsina) antillea CUSHMAN, ranges from 1.25 to 1.35 mm. The diameter the pro-
Cole, p. 145; Pl. 8, figs. 7-11; Pl. 9, figs. 3, 4; Pl. 10, fig.1 loculus ranges between 0.250-0.350 mm.
1967 Miogypsina antillea CUSHMAN, Cole, Pl. 8, fig. Distribution. Ranikothalia bermudezi occurs in the
4; Pl. 9, figs. 6, 9 Don Juan Fm associated with Neodiscocyclina grims-
1981 Miogypsina antillea CUSHMAN, Butterlin, p. 43; dale. In the Loma Peñón Fm this form is associated
Pl. 21, figs. 3-5 with reworked Paleocene larger foraminifer as
Fallotella sp., and with Eocene larger foraminifers as
Samples. Barahona Mb (La Descubierta 5871-I-IG HH Eoconuloides lopeztrigoi and Discocyclina marginata.
9009). Age. According to Berlanga (1997), Robinson and
Remarks. Species with embryo in marginal position, Wright (1993) and Serra-Kiel et al. (1998) Ranikothalia
with well-developed lateral chamberlets, in tiers of up bermudezi is Late Paleocene.
to 5 superposed chamberlets. Caudri (1996) reported
a similar form from the Lower Miocene of Trinidad
that she name Miogypsina (Miogypsina) bramletti Genus Nummulites Lamarck, 1801
GRAVELL 1933. Nummulites cf. willcoxi (HEILPRIN, 1882)
Distribution. This form occurs in the Barahona Mb Pl. 3, figs. 8 and 9
associated with Operculinoides sp.
Age. According to Cole (1961b), Drooger (1952), Cole 1953 Operculinoides willcoxi (HEILPRIN), Cole, Pl. 1,
and Applin (1961), Butterlin (1963) this form occurs in figs. 8-10; Pl. 2, fig. 2
the Early Miocene, Butterlin (1981) and De Mello e 1958a Operculinoides willcoxi (HEILPRIN), Cole, p.273-
Sousa et al. (2003) extends its range up to Middle 276; Pl. 33, figs. 1, 3-12
Miocene.
1961b Camerina willcoxi (HEILPRIN), Cole, Pl. 17, figs.
1, 10
1964 Camerina willcoxi (HEILPRIN), Cole and Applin,
Family NUMMULITIDAE de Blainville, 1827
p. 24; Pl. 4, figs. 9, 14-16
Genus Ranikothalia Caudri, 1944
Ranikothalia bermudezi (PALMER, 1934) 1966 Camerina willcoxi (HEILPRIN), Cole, Pl. 20, fig.
Pl. 3, figs. 5, 6 and 7 11; Pl. 22, fig. 3
1974 Nummulites (Paleonummulites) willcoxi
1934 Operculina bermudezi n. sp., Palmer, p. 238- HEILPRIN, Frost and Langenheim, p. 79-83; Pl.
240; Pl. 12, figs. 3, 6-9 13; Pl. 14
1947 Miscellanea antillea (HANZAWA), Cole and 1981 Nummulites willcoxi HEILPRIN, Butterlin, p. 32;
Bermúdez, p. 195-196; Pl. 2, figs. 10-11 Pl. 11
1953 Operculinoides bermudezi (PALMER), Cole, p. 1993 Paleonummulites willcoxi (HEILPRIN), Robinson
35-37; Pl. 1, fig. 5-7; Pl. 2, fig. 4; Pl. 3, figs. 2-12 and Wright, p. 333, 335; Figs. 29.6, 30.4-5
1960b Camerina catenula (CUSHMAN and JARVIS), 1997 Nummulites cf. willcoxi HEILPRIN, Berlanga, p.
Cole, Pl. 25, fig. 6 165; Pl. 35, fig. 8
377
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378
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
Samples. Upper Neiba Fm (Arroyo Limón 5973-III-BR the Upper Neiba Fm it is associated with Lepidocyclina
EB 9053). (Eulepidina) undosa, Lepidocyclina (Lepidocyclina)
Remarks. We have only axial and oblique sections. mantelli and Lepidocyclina (Lepidocyclina) canellei.
The axial section is lenticular, slightly rhomboidal in Age. According to Cole (1966), Butterlin (1981), Frost
outline and shows a marked marginal cord and some and Langenheim (1974) and De Mello e Sousa et al.
piles in the polar zone (2003) this species occurs in the Oligocene.
Distribution. This form occurs in the Upper Neiba Fm
associated with Amphistegina parvula and
Lepidocyclina (Lepidocyclina) macdonaldi. Genus Heterostegina d’Orbigny, 1826
Age. According to Cole (1964, 1966), Butterlin (1971, Heterostegina cf. antillea CUSHMAN, 1919
1981), Frost and Langenheim (1974) and Robinson Pl. 3, figs. 12 and 13
and Wright (1993) this form ranges from the Middle to
the Late Eocene. 1957b Heterostegina antillea CUSHMAN, Cole, p. 327;
Nummulites cf. panamensis CUSHMAN, 1918 Pl. 25, figs. 3-5
Pl. 3, figs. 10 and 11 1974 Heterostegina antillea CUSHMAN, Frost and
Langenheim, p. 90-92; Pl. 18
1958b Operculinoides panamensis (CUSHMAN), Cole, 1971 Heterostegina antillea CUSHMAN, Butterlin, Pl.
Pl. 25, figs. 1-2, 14-15 5, figs. 1-4
1966 Camerina panamensis (CUSHMAN), Cole, Pl. 20, 1981 Heterostegina antillea CUSHMAN, Butterlin, p.
figs. 1-10, 12; Pl. 25, fig. 5 32; Pl. 12, figs. 1-2
1974 Nummulites (Paleonummulites) panamensis 1993 Heterostegina (Vlerkina) antillea CUSHMAN,
CUSHMAN, Frost and Langenheim, p. 84-89; Pls. Robinson and Wright, p. 337; Figs. 31.3, 31.5
15, 16, 17 1996 Heterostegina antillea CUSHMAN, Caudri, Pl. 8,
1981 Nummulites panamensis CUSHMAN, Butterlin, fig. 6; Pl. 11, fig. 8
p. 12; Pl. 10, figs. 11-12 2003 Heterostegina antillea CUSHMAN, De Mello e
Sousa et al., Pl. 1, figs. 8-9
Samples. Brecciated Neiba Fm (Galván 5971-IV-IG HH
9034); Upper Neiba Fm (Bánica 5873-II-BR JS 9022 Samples. Brecciated Neiba Fm (Galván 5971-IV-IG HH
and 9048). 9034), Upper Neiba Fm (Bánica 5873-II-BR JS 9001;
Remarks. Planispiral-involute forms with slightly Galván 5971-IV-IG HH 9012; La Descubierta 5871-I-IG
operculiniform growth. The chambers are higher that HH 9023).
wide. The septa are straight in the bottom and middle Remarks. Nummulitid that shows chambers subdivid-
part and strongly oblique in the top of the chamber. ed into chamberlets in equatorial section. Because of
The specific determination is not confirmed because the lack of centred equatorial sections it could not be
of the absence of centred equatorial sections. determined at the specific level.
Distribution. This species occurs in the Brecciated Distribution. Heterostegina cf. antillea occurs in the
Neiba Fm associated with Heterostegina cf. antillea. In Brecciated Neiba Fm associated with Nummulites cf.
379
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380
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
and Cole (1941), Cole and Bermúdez (1947), Butterlin Lower Neiba Fm
(1981) and Blanco-Bustamante et al. (1999, no fig.)
The association of Eoconuloides wellsi, Eoconuloides
lopeztrigoi and Fabiania cassis characterizes the
Neodiscocyclina grimsdalei VAUGHAN and COLE, 1941 Middle Eocene
Pl. 3, fig. 24 Upper Neiba Fm and the Brecciated Neiba Fm
1945 Discocyclina grimsdalei VAUGHAN and COLE, The association of Coleiconus elongatus, Fallotella
Vaughan 1945, p. 39-42; Pl. 6, figs. 11-12; Pl. cookei, Cushmania puilboreauensis, Cushmania
13, Figs 1-6 americana, Fabiania cassis, Eoconuloides lopeztrigoi,
1975 Neodiscocyclina grimsdalei (VAUGHAN and Amphistegina parvula, Discocyclina marginata,
COLE), Caudri, p. 552-553; Pl. 3, figs. 8; Pl. 15; Polylepidina antillea and Penoperculinoides cubensis
Pl. 23, fig. 2 characterizes the Middle Eocene in the lower part of
1996 Neodiscocyclina grimsdalei (VAUGHAN and these units.
COLE), Caudri, p. 1203, Pl. 12, fig. 7 The association of Fallotella cookei, Cushmania
Samples. La Luisa Fm (Monte Plata 6272-III-IG HH americana, Fabiania cassis, Amphistegina parvula,
9008). Lepidocyclina (Lepidocyclina) macdonaldi,
Remarks: This species is similar, in equatorial section, Lepidocyclina (Nephrolepidina) cf. chaperi,
to the also Paleocene species Discocyclina weaveri Lepidocyclina (Pliolepidina) cf. peruviana and N. cf.
VAUGHAN 1929, from which it differs in the number of willcoxi characterizes the Middle-Late Eocene in the
chamberlets of the first annulus, higher in middle part of these units.
Neodiscocyclina grimsdalei, and in the morphology The upper part of these units renders an associa-
of the equatorial chamberlets, rectangular in tion composed of Nummulites cf. panamensis,
Neodiscocyclina grimsdalei, and quadrangular in Heterostegina cf. antillea, Lepidocyclina
Discocyclina weaveri VAUGHAN 1929. (Lepidocyclina) mantelli, Lepidocyclina
Distribution. This form occurs in the La Luisa Fm
(Lepidocyclina) canellei, Lepidocyclina (Eulepidina)
associated with Ranikothalia bermudezi.
undosa and Lepidocyclina (Nephrolepidina)
Age. According to Vaughan (1945), Caudri (1975,
tournoueri, typical of the Oligocene in middle shelf
1996) and Kugler and Caudri (1975) this species
facies.
occurs in the Paleocene
In the shallow carbonate environments, the same
lithostratigraphic units are characterized by archa-
iasinids, Praerhapydionina cf. delicata and
Conclusions
Discorinopsis sp., which indicate an Oligocene age.
The association of Lepidocyclina (Lepidocyclina)
The systematic and biostratigraphic study of the
mantelli and Lepidocyclina (Eulepidina) undosa indi-
Cainozoic larger foraminifers from the Dominican
cates an Oligocene-Early Miocene age in middle-
Republic allowed to recognize associations and to
outer shelf.
define chronostratigraphic intervals.
In the Don Juan basin (La Luisa Fm, Monte Plata
sheet, Fig. 1) the presence of Ranikothalia bermudezi
and Neodiscocyclina grimsdalei permits to identify Catanamatías Fm and Sombrerito Fm
the Late Paleocene (Thanetian). The same association
is found reworked within Middle Eocene rocks in the Reworked casts and blocks within the Catanamatías
Bejucal basin (Loma Peñón Fm, Rincón Chavón sheet, Fm contain Fallotella cookei, Cushmania americana,
Fig. 1). The Loma Peñón Fm also contains Fabiania cassis, Eodictyoconus cubensis,
Eoconuloides wellsi, Eoconuloides lopeztrigoi and Amphistegina parvula, Victoriella conoidea,
Neodiscocyclina barkeri, an association typical of the Polylepidina antillea, Lepidocyclina (Lepidocyclina)
Early Eocene. However, the presence of Discocyclina macdonaldi and Lepidocyclina (Nephrolepidina) cf.
marginata in the same samples indicates that this chaperi, which indicate a Middle-Late Eocene age.
association is reworked within the Middle Eocene. The presence of Lepidocyclina (Nephrolepidina)
A chronostratigraphic framework for the western tournoueri in turbidites of the Catanamatías Fm and
Dominican Republic has been elaborated from the Sombrerito Fm indicates an Oligocene-earliest
following larger foraminifer associations (Fig. 2): Middle Miocene age.
381
Serra-Kiel, J. et al., 2007. Cainozoic larger foraminifers from Dominican Republic. Boletín Geológico y Minero, 118 (2): 359-384
382
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