Horse - Wikipedia

2/1/23, 8:39 AM Horse - Wikipedia
Contents
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1Biology
1.1Lifespan and life stages
1.2Size and measurement

1.2.1Ponies
1.3Genetics
1.4Colors and markings
1.5Reproduction and development
1.6Anatomy
1.6.1Skeletal system
1.6.2Hooves
1.6.3Teeth
1.6.4Digestion
1.6.5Senses
1.7Movement
1.8Behavior
1.8.1Intelligence and learning
1.8.2Temperament
1.8.3Sleep patterns
2Taxonomy and evolution
2.1Wild species surviving into modern times

2.2Other modern equids
3Domestication
3.1Feral populations
https://en.wikipedia.org/wiki/Horse 1/34
3.2Breeds
4Interaction with humans
4.1Sport
4.2Work
4.3Warfare
4.4Entertainment and culture
4.5Therapeutic use
4.6Products
4.7Care
5See also
6References
7Sources
8Further reading
9External links
Horse
The horse (Equus ferus caballus)[2][3] is a domesticated,
one-toed, hoofed mammal. It belongs to the taxonomic family Horse
Equidae and is one of two extant subspecies of Equus ferus. The
horse has evolved over the past 45 to 55 million years from a
small multi-toed creature, Eohippus, into the large, single-toed
animal of today. Humans began domesticating horses around
4000 BCE, and their domestication is believed to have been
widespread by 3000 BCE. Horses in the subspecies caballus are
domesticated, although some domesticated populations live in
the wild as feral horses. These feral populations are not true
wild horses, as this term is used to describe horses that have Conservation status
never been domesticated. There is an extensive, specialized
vocabulary used to describe equine-related concepts, covering Domesticated
everything from anatomy to life stages, size, colors, markings, Scientific classification
breeds, locomotion, and behavior.
Kingdom: Animalia
Horses are adapted to run, allowing them to quickly escape
Phylum: Chordata
predators, and possess an excellent sense of balance and a
strong fight-or-flight response. Related to this need to flee from Class: Mammalia
predators in the wild is an unusual trait: horses are able to sleep
Order: Perissodactyla
both standing up and lying down, with younger horses tending
to sleep significantly more than adults.[4] Female horses, called Family: Equidae
mares, carry their young for approximately 11 months and a Genus: Equus
young horse, called a foal, can stand and run shortly following
birth. Most domesticated horses begin training under a saddle Species: E. ferus
or in a harness between the ages of two and four. They reach full Subspecies: E. f. caballus
adult development by age five, and have an average lifespan of
between 25 and 30 years. Trinomial name
Equus ferus caballus
Horse breeds are loosely divided into three categories based on
general temperament: spirited "hot bloods" with speed and Linnaeus, 1758[1]
endurance; "cold bloods", such as draft horses and some ponies, Synonyms[2]
suitable for slow, heavy work; and "warmbloods", developed
from crosses between hot bloods and cold bloods, often focusing
at least 48 published
on creating breeds for specific riding purposes, particularly in
Europe. There are more than 300 breeds of horse in the world
today, developed for many different uses.
Horses and humans interact in a wide variety of sport competitions and non-competitive recreational
pursuits as well as in working activities such as police work, agriculture, entertainment, and therapy.
Horses were historically used in warfare, from which a wide variety of riding and driving techniques
developed, using many different styles of equipment and methods of control. Many products are
derived from horses, including meat, milk, hide, hair, bone, and pharmaceuticals extracted from the
urine of pregnant mares. Humans provide domesticated horses with food, water, and shelter as well as
attention from specialists such as veterinarians and farriers.
Biology
Specific terms and specialized language are used to
describe equine anatomy, different life stages, and
colors and breeds.
Lifespan and life stages
Depending on breed, management and

environment, the modern domestic horse has a life
expectancy of 25 to 30 years.[7] Uncommonly, a
few animals live into their 40s and, occasionally,
beyond.[8] The oldest verifiable record was "Old
Points of a horse[5][6]
Billy", a 19th-century horse that lived to the age of
62.[7] In modern times, Sugar Puff, who had been
listed in Guinness World Records as the world's oldest living pony, died in 2007 at age 56.[9]
Regardless of a horse or pony's actual birth date, for most competition purposes a year is added to its
age each January 1 of each year in the Northern Hemisphere[7][10] and each August 1 in the Southern
Hemisphere.[11] The exception is in endurance riding, where the minimum age to compete is based on
the animal's actual calendar age.[12]
The following terminology is used to describe horses of various ages:
Foal
A horse of either sex less than one year old. A nursing foal is sometimes called a suckling, and a
foal that has been weaned is called a weanling.[13] Most domesticated foals are weaned at five
to seven months of age, although foals can be weaned at four months with no adverse physical
effects.[14]
Yearling
A horse of either sex that is between one and two years old.[15]
Colt
A male horse under the age of four.[16] A common terminology error is to call any young horse a
"colt", when the term actually only refers to young male horses.[17]
Filly
A female horse under the age of four.[13]
Mare
A female horse four years old and older.[18]
Stallion
A non-castrated male horse four years old and older.[19] The term "horse" is sometimes used
colloquially to refer specifically to a stallion.[20]
Gelding
A castrated male horse of any age.[13]
In horse racing, these definitions may differ: For example, in the British Isles, Thoroughbred horse
racing defines colts and fillies as less than five years old.[21] However, Australian Thoroughbred racing
defines colts and fillies as less than four years old.[22]
Size and measurement
The height of horses is measured at the highest point of the withers, where the neck meets the
back.[23] This point is used because it is a stable point of the anatomy, unlike the head or neck, which
move up and down in relation to the body of the horse.
In English-speaking countries, the height of horses is

often stated in units of hands and inches: one hand is
equal to 4 inches (101.6 mm). The height is expressed
as the number of full hands, followed by a point, then
the number of additional inches, and ending with the
abbreviation "h" or "hh" (for "hands high"). Thus, a
horse described as "15.2 h" is 15 hands plus 2 inches,
for a total of 62 inches (157.5 cm) in height.[24]
The size of horses varies by breed, but also is

influenced by nutrition. Light-riding horses usually
range in height from 14 to 16 hands (56 to 64 inches,
Size varies greatly among horse breeds, as with
142 to 163 cm) and can weigh from 380 to 550
this full-sized horse and small pony.
kilograms (840 to 1,210 lb).[25] Larger-riding horses
usually start at about 15.2 hands (62 inches, 157 cm)
and often are as tall as 17 hands (68 inches, 173 cm), weighing from 500 to 600 kilograms (1,100 to
1,320 lb).[26] Heavy or draft horses are usually at least 16 hands (64 inches, 163 cm) high and can be
as tall as 18 hands (72 inches, 183 cm) high. They can weigh from about 700 to 1,000 kilograms (1,540
to 2,200 lb).[27]
The largest horse in recorded history was probably a Shire horse named Mammoth, who was born in
1848. He stood 21.21⁄4 hands (86.25 inches, 219 cm) high and his peak weight was estimated at 1,524
kilograms (3,360 lb).[28] The record holder for the smallest horse ever is Thumbelina, a fully mature
miniature horse affected by dwarfism. She was 43 cm (17 in) tall and weighed 26 kg (57 lb).[29][30]
Ponies
Ponies are taxonomically the same animals as horses. The distinction between a horse and pony is
commonly drawn on the basis of height, especially for competition purposes. However, height alone is
not dispositive; the difference between horses and ponies may also include aspects of phenotype,
including conformation and temperament.
The traditional standard for height of a horse or a pony at maturity is 14.2 hands (58 inches, 147 cm).
An animal 14.2 hands (58 inches, 147 cm) or over is usually considered to be a horse and one less than
14.2 hands (58 inches, 147 cm) a pony,[31] but there are many exceptions to the traditional standard.
In Australia, ponies are considered to be those under 14 hands (56 inches, 142 cm).[32] For
competition in the Western division of the United States Equestrian Federation, the cutoff is
14.1 hands (57 inches, 145 cm).[33] The International Federation for Equestrian Sports, the world
governing body for horse sport, uses metric measurements and defines a pony as being any horse
measuring less than 148 centimetres (58.27 in) at the withers without shoes, which is just over
14.2 hands (58 inches, 147 cm), and 149 centimetres (58.66 in; 14.21⁄2 hands), with shoes.[34]
Height is not the sole criterion for distinguishing horses from ponies. Breed registries for horses that
typically produce individuals both under and over 14.2 hands (58 inches, 147 cm) consider all animals
of that breed to be horses regardless of their height.[35] Conversely, some pony breeds may have
features in common with horses, and individual animals may occasionally mature at over 14.2 hands
(58 inches, 147 cm), but are still considered to be ponies.[36]
Ponies often exhibit thicker manes, tails, and overall coat. They also have proportionally shorter legs,
wider barrels, heavier bone, shorter and thicker necks, and short heads with broad foreheads. They
may have calmer temperaments than horses and also a high level of intelligence that may or may not
be used to cooperate with human handlers.[31] Small size, by itself, is not an exclusive determinant.
For example, the Shetland pony which averages 10 hands (40 inches, 102 cm), is considered a
pony.[31] Conversely, breeds such as the Falabella and other miniature horses, which can be no taller
than 76 centimetres (30 in), are classified by their registries as very small horses, not ponies.[37]
Genetics
Horses have 64 chromosomes.[38] The horse genome was sequenced in 2007. It contains 2.7 billion
DNA base pairs,[39] which is larger than the dog genome, but smaller than the human genome or the
bovine genome.[40] The map is available to researchers.[41]
Colors and markings
Horses exhibit a diverse array of coat colors and distinctive markings, described by a specialized
vocabulary. Often, a horse is classified first by its coat color, before breed or sex.[42] Horses of the
same color may be distinguished from one another by white markings,[43] which, along with various
spotting patterns, are inherited separately from coat color.[44]
Many genes that create horse coat colors and patterns

have been identified. Current genetic tests can
identify at least 13 different alleles influencing coat
color,[45] and research continues to discover new
genes linked to specific traits. The basic coat colors of
chestnut and black are determined by the gene
controlled by the Melanocortin 1 receptor,[46] also
known as the "extension gene" or "red factor,"[45] as
its recessive form is "red" (chestnut) and its dominant
form is black.[47] Additional genes control
suppression of black color to point coloration that
results in a bay, spotting patterns such as pinto or Bay (left) and chestnut (sometimes called "sorrel")
leopard, dilution genes such as palomino or dun, as are two of the most common coat colors, seen in
well as greying, and all the other factors that create almost all breeds.
the many possible coat colors found in horses.[45]
Horses that have a white coat color are often mislabeled; a horse that looks "white" is usually a
middle-aged or older gray. Grays are born a darker shade, get lighter as they age, but usually keep
black skin underneath their white hair coat (with the exception of pink skin under white markings).
The only horses properly called white are born with a predominantly white hair coat and pink skin, a
fairly rare occurrence.[47] Different and unrelated genetic factors can produce white coat colors in
horses, including several different alleles of dominant white and the sabino-1 gene.[48] However, there
are no "albino" horses, defined as having both pink skin and red eyes.[49]
Reproduction and development
Gestation lasts approximately 340 days, with an average range

320–370 days,[50] and usually results in one foal; twins are
rare.[51] Horses are a precocial species, and foals are capable of
standing and running within a short time following birth.[52] Foals
are usually born in the spring. The estrous cycle of a mare occurs
roughly every 19–22 days and occurs from early spring into
autumn. Most mares enter an anestrus period during the winter
and thus do not cycle in this period.[53] Foals are generally weaned
from their mothers between four and six months of age.[54] Mare with a foal
Horses, particularly colts, are sometimes physically capable of

reproduction at about 18 months, but domesticated horses are rarely allowed to breed before the age
of three, especially females.[55] Horses four years old are considered mature, although the skeleton
normally continues to develop until the age of six; maturation also depends on the horse's size, breed,
sex, and quality of care. Larger horses have larger bones; therefore, not only do the bones take longer
to form bone tissue, but the epiphyseal plates are larger and take longer to convert from cartilage to
bone. These plates convert after the other parts of the bones, and are crucial to development.[56]
Depending on maturity, breed, and work expected, horses are usually put under saddle and trained to
be ridden between the ages of two and four.[57] Although Thoroughbred race horses are put on the
track as young as the age of two in some countries,[58] horses specifically bred for sports such as
dressage are generally not put under saddle until they are three or four years old, because their bones
and muscles are not solidly developed.[59] For endurance riding competition, horses are not deemed
mature enough to compete until they are a full 60 calendar months (five years) old.[12]
Anatomy
Skeletal system
The horse skeleton averages 205 bones.[60] A

significant difference between the horse skeleton and
that of a human is the lack of a collarbone—the
horse's forelimbs are attached to the spinal column by
a powerful set of muscles, tendons, and ligaments
that attach the shoulder blade to the torso. The
horse's four legs and hooves are also unique
structures. Their leg bones are proportioned
differently from those of a human. For example, the
body part that is called a horse's "knee" is actually
made up of the carpal bones that correspond to the
human wrist. Similarly, the hock contains bones
equivalent to those in the human ankle and heel. The The skeletal system of a modern horse
lower leg bones of a horse correspond to the bones of
the human hand or foot, and the fetlock (incorrectly
called the "ankle") is actually the proximal sesamoid bones between the cannon bones (a single
equivalent to the human metacarpal or metatarsal bones) and the proximal phalanges, located where
one finds the "knuckles" of a human. A horse also has no muscles in its legs below the knees and
hocks, only skin, hair, bone, tendons, ligaments, cartilage, and the assorted specialized tissues that
make up the hoof.[61]
Hooves
The critical importance of the feet and legs is summed up by the traditional adage, "no foot, no
horse".[62] The horse hoof begins with the distal phalanges, the equivalent of the human fingertip or
tip of the toe, surrounded by cartilage and other specialized, blood-rich soft tissues such as the
laminae. The exterior hoof wall and horn of the sole is made of keratin, the same material as a human
fingernail.[63] The result is that a horse, weighing on average 500 kilograms (1,100 lb),[64] travels on
the same bones as would a human on tiptoe.[65] For the protection of the hoof under certain
conditions, some horses have horseshoes placed on their feet by a professional farrier. The hoof
continually grows, and in most domesticated horses needs to be trimmed (and horseshoes reset, if
used) every five to eight weeks,[66] though the hooves of horses in the wild wear down and regrow at a
rate suitable for their terrain.
Teeth
Horses are adapted to grazing. In an adult horse, there are 12 incisors at the front of the mouth,
adapted to biting off the grass or other vegetation. There are 24 teeth adapted for chewing, the
premolars and molars, at the back of the mouth. Stallions and geldings have four additional teeth just
behind the incisors, a type of canine teeth called "tushes". Some horses, both male and female, will
also develop one to four very small vestigial teeth in front of the molars, known as "wolf" teeth, which
are generally removed because they can interfere with the bit. There is an empty interdental space
between the incisors and the molars where the bit rests directly on the gums, or "bars" of the horse's
mouth when the horse is bridled.[67]
An estimate of a horse's age can be made from looking at its teeth. The teeth continue to erupt
throughout life and are worn down by grazing. Therefore, the incisors show changes as the horse ages;
they develop a distinct wear pattern, changes in tooth shape, and changes in the angle at which the
chewing surfaces meet. This allows a very rough estimate of a horse's age, although diet and veterinary
care can also affect the rate of tooth wear.[7]
Digestion
Horses are herbivores with a digestive system adapted to a forage diet of grasses and other plant
material, consumed steadily throughout the day. Therefore, compared to humans, they have a
relatively small stomach but very long intestines to facilitate a steady flow of nutrients. A 450-
kilogram (990 lb) horse will eat 7 to 11 kilograms (15 to 24 lb) of food per day and, under normal use,
drink 38 to 45 litres (8.4 to 9.9 imp gal; 10 to 12 US gal) of water. Horses are not ruminants, they have
only one stomach, like humans, but unlike humans, they can use cellulose, a major component of
grass. Horses are hindgut fermenters. Cellulose fermentation by symbiotic bacteria occurs in the
cecum, or "water gut", which food goes through before reaching the large intestine. Horses cannot
vomit, so digestion problems can quickly cause colic, a leading cause of death.[68] Horses do not have
a gallbladder; however, they seem to tolerate high amounts of fat in their diet despite lack of a
gallbladder.[69][70]
Senses
The horses' senses are based on their status as prey animals, where
they must be aware of their surroundings at all times.[71] They
have the largest eyes of any land mammal,[72] and are lateral-eyed,
meaning that their eyes are positioned on the sides of their
heads.[73] This means that horses have a range of vision of more
than 350°, with approximately 65° of this being binocular vision
and the remaining 285° monocular vision.[72] Horses have
excellent day and night vision, but they have two-color, or A horse's eye
dichromatic vision; their color vision is somewhat like red-green
color blindness in humans, where certain colors, especially red
and related colors, appear as a shade of green.[74]
Their sense of smell, while much better than that of humans, is not quite as good as that of a dog. It is
believed to play a key role in the social interactions of horses as well as detecting other key scents in
the environment. Horses have two olfactory centers. The first system is in the nostrils and nasal
cavity, which analyze a wide range of odors. The second, located under the nasal cavity, are the
vomeronasal organs, also called Jacobson's organs. These have a separate nerve pathway to the brain
and appear to primarily analyze pheromones.[75]
A horse's hearing is good,[71] and the pinna of each ear can rotate up to 180°, giving the potential for
360° hearing without having to move the head.[76] Noise impacts the behavior of horses and certain
kinds of noise may contribute to stress: A 2013 study in the UK indicated that stabled horses were
calmest in a quiet setting, or if listening to country or classical music, but displayed signs of
nervousness when listening to jazz or rock music. This study also recommended keeping music under
a volume of 21 decibels.[77] An Australian study found that stabled racehorses listening to talk radio
had a higher rate of gastric ulcers than horses listening to music, and racehorses stabled where a radio
was played had a higher overall rate of ulceration than horses stabled where there was no radio
playing.[78]
Horses have a great sense of balance, due partly to their ability to feel their footing and partly to highly
developed proprioception—the unconscious sense of where the body and limbs are at all times.[79] A
horse's sense of touch is well-developed. The most sensitive areas are around the eyes, ears, and
nose.[80] Horses are able to sense contact as subtle as an insect landing anywhere on the body.[81]
Horses have an advanced sense of taste, which allows them to sort through fodder and choose what
they would most like to eat,[82] and their prehensile lips can easily sort even small grains. Horses
generally will not eat poisonous plants, however, there are exceptions; horses will occasionally eat
toxic amounts of poisonous plants even when there is adequate healthy food.[83]
Movement
Walk 5–8 km/h (3.1– Trot 8–13 km/h Pace 8–13 km/h Canter 16–27 km/h
5.0 mph) (5.0–8.1 mph) (5.0–8.1 mph) (9.9–16.8 mph)
Gallop 40–48 km/h

(25–30 mph),
record: 70.76 km/h
(43.97 mph)
All horses move naturally with four basic gaits:[84]
the four-beat walk, which averages 6.4 kilometres per hour (4.0 mph);
the two-beat trot or jog at 13 to 19 kilometres per hour (8.1 to 11.8 mph) (faster for harness racing
horses);
the canter or lope, a three-beat gait that is 19 to 24 kilometres per hour (12 to 15 mph);
the gallop, which averages 40 to 48 kilometres per hour (25 to 30 mph),[85] but the world record for
a horse galloping over a short, sprint distance is 70.76 kilometres per hour (43.97 mph).[86]
Besides these basic gaits, some horses perform a two-beat pace, instead of the trot.[87] There also are
several four-beat 'ambling' gaits that are approximately the speed of a trot or pace, though smoother
to ride. These include the lateral rack, running walk, and tölt as well as the diagonal fox trot.[88]
Ambling gaits are often genetic in some breeds, known collectively as gaited horses.[89] These horses
replace the trot with one of the ambling gaits.[90]
Behavior
Horses are prey animals with a strong fight-or-flight response. 0:03

Their first reaction to a threat is to startle and usually flee,
Horse neigh
although they will stand their ground and defend themselves when
flight is impossible or if their young are threatened.[91] They also
tend to be curious; when startled, they will often hesitate an instant to ascertain the cause of their
fright, and may not always flee from something that they perceive as non-threatening. Most light
horse riding breeds were developed for speed, agility, alertness and endurance; natural qualities that
extend from their wild ancestors. However, through selective breeding, some breeds of horses are
quite docile, particularly certain draft horses.[92]
Horses are herd animals, with a clear hierarchy of rank, led by a dominant individual, usually a mare.
They are also social creatures that are able to form companionship attachments to their own species
and to other animals, including humans. They communicate in various ways, including vocalizations
such as nickering or whinnying, mutual grooming, and body language. Many horses will become
difficult to manage if they are isolated, but with training, horses can learn to accept a human as a
companion, and thus be comfortable away from other horses.[93] However, when confined with
insufficient companionship, exercise, or stimulation, individuals may develop stable vices, an
assortment of bad habits, mostly stereotypies of psychological origin, that include wood chewing, wall
kicking, "weaving" (rocking back and forth), and other problems.[94]
Intelligence and learning
Studies have indicated that horses perform a number of cognitive tasks on a daily basis, meeting
mental challenges that include food procurement and identification of individuals within a social
system. They also have good spatial discrimination abilities.[95] They are naturally curious and apt to
investigate things they have not seen before.[96] Studies have assessed equine intelligence in areas
such as problem solving, speed of learning, and memory. Horses excel at simple learning, but also are
able to use more advanced cognitive abilities that involve categorization and concept learning. They
can learn using habituation, desensitization, classical conditioning, and operant conditioning, and
positive and negative reinforcement.[95] One study has indicated that horses can differentiate between
"more or less" if the quantity involved is less than four.[97]
Domesticated horses may face greater mental challenges than wild horses, because they live in
artificial environments that prevent instinctive behavior whilst also learning tasks that are not
natural.[95] Horses are animals of habit that respond well to regimentation, and respond best when
the same routines and techniques are used consistently. One trainer believes that "intelligent" horses
are reflections of intelligent trainers who effectively use response conditioning techniques and positive
reinforcement to train in the style that best fits with an individual animal's natural inclinations.[98]
Temperament
Horses are mammals, and as such are warm-blooded, or endothermic creatures, as opposed to cold-
blooded, or poikilothermic animals. However, these words have developed a separate meaning in the
context of equine terminology, used to describe temperament, not body temperature. For example, the
"hot-bloods", such as many race horses, exhibit more sensitivity and energy,[99] while the "cold-
bloods", such as most draft breeds, are quieter and calmer.[100] Sometimes "hot-bloods" are classified
as "light horses" or "riding horses",[101] with the "cold-bloods" classified as "draft horses" or "work
horses".[102]
"Hot blooded" breeds include "oriental horses"

such as the Akhal-Teke, Arabian horse, Barb, and
now-extinct Turkoman horse, as well as the
Thoroughbred, a breed developed in England from
the older oriental breeds.[99] Hot bloods tend to be
spirited, bold, and learn quickly. They are bred for
agility and speed.[103] They tend to be physically
refined—thin-skinned, slim, and long-legged.[104]
The original oriental breeds were brought to
Europe from the Middle East and North Africa
when European breeders wished to infuse these
traits into racing and light cavalry horses.[105][106]
Muscular, heavy draft horses are known as "cold

bloods", as they are bred not only for strength, but Illustration of assorted breeds; slim, light hotbloods,
also to have the calm, patient temperament needed medium-sized warmbloods and draft and pony-type
to pull a plow or a heavy carriage full of coldblood breeds
people.[100] They are sometimes nicknamed
"gentle giants".[107] Well-known draft breeds
include the Belgian and the Clydesdale.[107] Some, like the Percheron, are lighter and livelier,
developed to pull carriages or to plow large fields in drier climates.[108] Others, such as the Shire, are
slower and more powerful, bred to plow fields with heavy, clay-based soils.[109] The cold-blooded
group also includes some pony breeds.[110]
"Warmblood" breeds, such as the Trakehner or Hanoverian, developed when European carriage and
war horses were crossed with Arabians or Thoroughbreds, producing a riding horse with more
refinement than a draft horse, but greater size and milder temperament than a lighter breed.[111]
Certain pony breeds with warmblood characteristics have been developed for smaller riders.[112]
Warmbloods are considered a "light horse" or "riding horse".[101]
Today, the term "Warmblood" refers to a specific subset of sport horse breeds that are used for
competition in dressage and show jumping.[113] Strictly speaking, the term "warm blood" refers to any
cross between cold-blooded and hot-blooded breeds.[114] Examples include breeds such as the Irish
Draught or the Cleveland Bay. The term was once used to refer to breeds of light riding horse other
than Thoroughbreds or Arabians, such as the Morgan horse.[103]
Sleep patterns
Horses are able to sleep both standing up and lying down. In an

adaptation from life in the wild, horses are able to enter light sleep
by using a "stay apparatus" in their legs, allowing them to doze
without collapsing.[115] Horses sleep better when in groups
because some animals will sleep while others stand guard to watch
for predators. A horse kept alone will not sleep well because its
instincts are to keep a constant eye out for danger.[116]
When horses lie down to sleep, Unlike humans, horses do not sleep in a solid, unbroken period of
others in the herd remain standing,time, but take many short periods of rest. Horses spend four to
awake, or in a light doze, keeping fifteen hours a day in standing rest, and from a few minutes to
watch.
several hours lying down. Total sleep time in a 24-hour period may
range from several minutes to a couple of hours,[116] mostly in
short intervals of about 15 minutes each.[117] The average sleep
time of a domestic horse is said to be 2.9 hours per day.[118]
Horses must lie down to reach REM sleep. They only have to lie down for an hour or two every few
days to meet their minimum REM sleep requirements.[116] However, if a horse is never allowed to lie
down, after several days it will become sleep-deprived, and in rare cases may suddenly collapse as it
involuntarily slips into REM sleep while still standing.[119] This condition differs from narcolepsy,
although horses may also suffer from that disorder.[120]
Taxonomy and evolution

The horse adapted to survive in areas of wide-open terrain with sparse vegetation, surviving in an
ecosystem where other large grazing animals, especially ruminants, could not.[121] Horses and other
equids are odd-toed ungulates of the order Perissodactyla, a group of mammals dominant during the
Tertiary period. In the past, this order contained 14 families, but only three—Equidae (the horse and
related species), Tapiridae (the tapir), and Rhinocerotidae (the rhinoceroses)—have survived to the
present day.[122]
The earliest known member of the family Equidae was the Hyracotherium, which lived between 45
and 55 million years ago, during the Eocene period. It had 4 toes on each front foot, and 3 toes on each
back foot.[123] The extra toe on the front feet soon disappeared with the Mesohippus, which lived 32 to
37 million years ago.[124] Over time, the extra side toes shrank in size until they vanished. All that
remains of them in modern horses is a set of small vestigial bones on the leg below the knee,[125]
known informally as splint bones.[126] Their legs also lengthened as their toes disappeared until they
were a hooved animal capable of running at great speed.[125] By about 5 million years ago, the modern
Equus had evolved.[127] Equid teeth also evolved from browsing on soft, tropical plants to adapt to
browsing of drier plant material, then to grazing of

tougher plains grasses. Thus proto-horses changed
from leaf-eating forest-dwellers to grass-eating
inhabitants of semi-arid regions worldwide,
including the steppes of Eurasia and the Great
Plains of North America.
By about 15,000 years ago, Equus ferus was a

widespread holarctic species. Horse bones from
this time period, the late Pleistocene, are found in
Europe, Eurasia, Beringia, and North America.[128]
Yet between 10,000 and 7,600 years ago, the horse
became extinct in North America and rare
elsewhere.[129][130][131] The reasons for this
extinction are not fully known, but one theory
notes that extinction in North America paralleled Diagram of evolution in horses showing size
human arrival.[132] Another theory points to development, biometrical changes in the cranium and
climate change, noting that approximately 12,500 reduction of toes (left forefoot)
years ago, the grasses characteristic of a steppe
ecosystem gave way to shrub tundra, which was
covered with unpalatable plants.[133]
Wild species surviving into modern times
A truly wild horse is a species or subspecies with no ancestors that

were ever domesticated. Therefore, most "wild" horses today are
actually feral horses, animals that escaped or were turned loose
from domestic herds and the descendants of those animals.[134]
Only two wild subspecies, the tarpan and the Przewalski's horse,
survived into recorded history and only the latter survives today.
The Przewalski's horse (Equus ferus przewalskii), named after the

Russian explorer Nikolai Przhevalsky, is a rare Asian animal. It is
A small herd of Przewalski's Horses
also known as the Mongolian wild horse; Mongolian people know
it as the taki, and the Kyrgyz people call it a kirtag. The subspecies
was presumed extinct in the wild between 1969 and 1992, while a small breeding population survived
in zoos around the world. In 1992, it was reestablished in the wild by the conservation efforts of
numerous zoos.[135] Today, a small wild breeding population exists in Mongolia.[136][137] There are
additional animals still maintained at zoos throughout the world.
The question of whether the Przewalski's horse was ever domesticated was challenged in 2018 when
DNA studies of horses found at Botai culture sites revealed captured animals with DNA markers of an
ancestor to the Przewalski's horse. The study concluded that the Botai animals appear to have been an
independent domestication attempt involving a different wild population from all other domesticated
horses. However, the question of whether all Przewalski's horses descend from this domesticated
population is unresolved, as only one of seven modern Przewalski’s horses in the study shared this
ancestry.[138][139][140]
The tarpan or European wild horse (Equus ferus ferus) was found in Europe and much of Asia. It
survived into the historical era, but became extinct in 1909, when the last captive died in a Russian
zoo.[141] Thus, the genetic line was lost. Attempts have been made to recreate the tarpan,[141][142][143]
which resulted in horses with outward physical similarities, but nonetheless descended from
domesticated ancestors and not true wild horses.
Periodically, populations of horses in isolated areas are speculated to be relict populations of wild
horses, but generally have been proven to be feral or domestic. For example, the Riwoche horse of
Tibet was proposed as such,[137] but testing did not reveal genetic differences from domesticated
horses.[144] Similarly, the Sorraia of Portugal was proposed as a direct descendant of the Tarpan on
the basis of shared characteristics,[145][146] but genetic studies have shown that the Sorraia is more
closely related to other horse breeds, and that the outward similarity is an unreliable measure of
relatedness.[145][147]
Other modern equids
Besides the horse, there are six other species of genus Equus in the Equidae family. These are the ass
or donkey, Equus asinus; the mountain zebra, Equus zebra; plains zebra, Equus quagga; Grévy's
Zebra, Equus grevyi; the kiang, Equus kiang; and the onager, Equus hemionus.[148]
Horses can crossbreed with other members of their genus. The most common hybrid is the mule, a
cross between a "jack" (male donkey) and a mare. A related hybrid, a hinny, is a cross between a
stallion and a "jenny" (female donkey).[149] Other hybrids include the zorse, a cross between a zebra
and a horse.[150] With rare exceptions, most hybrids are sterile and cannot reproduce.[151]
Domestication
Domestication of the horse most likely took place in central Asia
prior to 3500 BCE. Two major sources of information are used to
determine where and when the horse was first domesticated and
how the domesticated horse spread around the world. The first
source is based on palaeological and archaeological discoveries;
the second source is a comparison of DNA obtained from modern
horses to that from bones and teeth of ancient horse remains.
Bhimbetka rock painting showing a
The earliest archaeological evidence for the domestication of the
man riding on a horse, India
horse comes from sites in Ukraine and Kazakhstan, dating to
approximately 4000–3500 BCE.[152][153][154] By 3000 BCE, the
horse was completely domesticated and by 2000 BCE there was a sharp increase in the number of
horse bones found in human settlements in northwestern Europe, indicating the spread of
domesticated horses throughout the continent.[155] The most recent, but most irrefutable evidence of
domestication comes from sites where horse remains were interred with chariots in graves of the
Sintashta and Petrovka cultures c. 2100 BCE.[156]
A 2021 genetic study suggested that most modern domestic horses descend from the lower Volga-Don
region. Ancient horse genomes indicate that these populations influenced almost all local populations
as they expanded rapidly throughout Eurasia, beginning about 4,200 years ago. It also shows that
certain adaptations were strongly selected due to riding, and that equestrian material culture,
including Sintashta spoke-wheeled chariots spread with the horse itself.[157][158]
Domestication is also studied by using the genetic material of present-day horses and comparing it
with the genetic material present in the bones and teeth of horse remains found in archaeological and
palaeological excavations. The variation in the genetic material shows that very few wild stallions
contributed to the domestic horse,[159][160] while many mares were part of early domesticated
herds.[147][161][162] This is reflected in the difference in genetic variation between the DNA that is
passed on along the paternal, or sire line (Y-chromosome) versus that passed on along the maternal,
or dam line (mitochondrial DNA). There are very low levels of Y-chromosome variability,[159][160] but
a great deal of genetic variation in mitochondrial DNA.[147][161][162] There is also regional variation in
mitochondrial DNA due to the inclusion of wild mares in domestic herds.[147][161][162][163] Another
characteristic of domestication is an increase in coat color variation.[164] In horses, this increased
dramatically between 5000 and 3000 BCE.[165]
Before the availability of DNA techniques to resolve the questions related to the domestication of the
horse, various hypotheses were proposed. One classification was based on body types and
conformation, suggesting the presence of four basic prototypes that had adapted to their environment
prior to domestication.[110] Another hypothesis held that the four prototypes originated from a single
wild species and that all different body types were entirely a result of selective breeding after
domestication.[166] However, the lack of a detectable substructure in the horse has resulted in a
rejection of both hypotheses.
Feral populations
Feral horses are born and live in the wild, but are descended from domesticated animals.[134] Many
populations of feral horses exist throughout the world.[167][168] Studies of feral herds have provided
useful insights into the behavior of prehistoric horses,[169] as well as greater understanding of the
instincts and behaviors that drive horses that live in domesticated conditions.[170]
There are also semi-feral horses in many parts of the world, such as Dartmoor and the New Forest in
the UK, where the animals are all privately owned but live for significant amounts of time in "wild"
conditions on undeveloped, often public, lands. Owners of such animals often pay a fee for grazing
rights.[171][172]
Breeds
The concept of purebred bloodstock and a controlled, written breed registry has come to be
particularly significant and important in modern times. Sometimes purebred horses are incorrectly or
inaccurately called "thoroughbreds". Thoroughbred is a specific breed of horse, while a "purebred" is a
horse (or any other animal) with a defined pedigree recognized by a breed registry.[173] Horse breeds
are groups of horses with distinctive characteristics that are transmitted consistently to their
offspring, such as conformation, color, performance ability, or disposition. These inherited traits
result from a combination of natural crosses and artificial selection methods. Horses have been
selectively bred since their domestication. An early example of people who practiced selective horse
breeding were the Bedouin, who had a reputation for careful practices, keeping extensive pedigrees of
their Arabian horses and placing great value upon pure bloodlines.[174] These pedigrees were
originally transmitted via an oral tradition.[175] In the 14th century, Carthusian monks of southern
Spain kept meticulous pedigrees of bloodstock lineages still found today in the Andalusian horse.[176]
Breeds developed due to a need for "form to function", the necessity to develop certain characteristics
in order to perform a particular type of work.[177] Thus, a powerful but refined breed such as the
Andalusian developed as riding horses with an aptitude for dressage.[177] Heavy draft horses were
developed out of a need to perform demanding farm work and pull heavy wagons.[178] Other horse
breeds had been developed specifically for light agricultural work, carriage and road work, various
sport disciplines, or simply as pets.[179] Some breeds developed through centuries of crossing other
breeds, while others descended from a single foundation sire, or other limited or restricted foundation
bloodstock. One of the earliest formal registries was General Stud Book for Thoroughbreds, which
began in 1791 and traced back to the foundation bloodstock for the breed.[180] There are more than
300 horse breeds in the world today.[181]
Interaction with humans

Worldwide, horses play a role within human cultures and have
done so for millennia. Horses are used for leisure activities, sports,
and working purposes. The Food and Agriculture Organization
(FAO) estimates that in 2008, there were almost 59,000,000
horses in the world, with around 33,500,000 in the Americas,
13,800,000 in Asia and 6,300,000 in Europe and smaller portions
in Africa and Oceania. There are estimated to be 9,500,000 horses
in the United States alone.[182] The American Horse Council
estimates that horse-related activities have a direct impact on the
economy of the United States of over $39 billion, and when
indirect spending is considered, the impact is over $102 Finnhorse pulling a heavy wagon.
billion.[183] In a 2004 "poll" conducted by Animal Planet, more
than 50,000 viewers from 73 countries voted for the horse as the
world's 4th favorite animal.[184]
Communication between human and horse is paramount in any equestrian activity;[185] to aid this
process horses are usually ridden with a saddle on their backs to assist the rider with balance and
positioning, and a bridle or related headgear to assist the rider in maintaining control.[186] Sometimes
horses are ridden without a saddle,[187] and occasionally, horses are trained to perform without a
bridle or other headgear.[188] Many horses are also driven, which requires a harness, bridle, and some
type of vehicle.[189]
Sport
Historically, equestrians honed their skills through games and races. Equestrian sports provided
entertainment for crowds and honed the excellent horsemanship that was needed in battle. Many
sports, such as dressage, eventing, and show jumping, have origins in military training, which were
focused on control and balance of both horse and rider. Other sports, such as rodeo, developed from
practical skills such as those needed on working ranches and stations. Sport hunting from horseback
evolved from earlier practical hunting techniques.[185] Horse racing of all types evolved from
impromptu competitions between riders or drivers. All forms of competition, requiring demanding
and specialized skills from both horse and rider, resulted in the systematic development of specialized
breeds and equipment for each sport. The popularity of equestrian sports through the centuries has
resulted in the preservation of skills that would otherwise have disappeared after horses stopped being
used in combat.[185]
Horses are trained to be ridden or driven in a variety of

sporting competitions. Examples include show jumping,
dressage, three-day eventing, competitive driving, endurance
riding, gymkhana, rodeos, and fox hunting.[190] Horse
shows, which have their origins in medieval European fairs,
are held around the world. They host a huge range of classes,
covering all of the mounted and harness disciplines, as well
as "In-hand" classes where the horses are led, rather than
ridden, to be evaluated on their conformation. The method
of judging varies with the discipline, but winning usually
depends on style and ability of both horse and rider.[191]
Sports such as polo do not judge the horse itself, but rather A horse and rider in dressage competition
use the horse as a partner for human competitors as a at the Olympics
necessary part of the game. Although the horse requires
specialized training to participate, the details of its
performance are not judged, only the result of the rider's actions—be it getting a ball through a goal or
some other task.[192] Examples of these sports of partnership between human and horse include
jousting, in which the main goal is for one rider to unseat the other,[193] and buzkashi, a team game
played throughout Central Asia, the aim being to capture a goat carcass while on horseback.[192]
Horse racing is an equestrian sport and major international industry, watched in almost every nation
of the world. There are three types: "flat" racing; steeplechasing, i.e. racing over jumps; and harness
racing, where horses trot or pace while pulling a driver in a small, light cart known as a sulky.[194] A
major part of horse racing's economic importance lies in the gambling associated with it.[195]
Work
There are certain jobs that horses do very

well, and no technology has yet developed
to fully replace them. For example,
mounted police horses are still effective
for certain types of patrol duties and
crowd control.[196] Cattle ranches still
require riders on horseback to round up
cattle that are scattered across remote,
rugged terrain.[197] Search and rescue Horse pulling a cart A mounted police
organizations in some countries depend officer in Poland
upon mounted teams to locate people,
particularly hikers and children, and to
provide disaster relief assistance.[198] Horses can also be used in areas where it is necessary to avoid
vehicular disruption to delicate soil, such as nature reserves. They may also be the only form of
transport allowed in wilderness areas. Horses are quieter than motorized vehicles. Law enforcement
officers such as park rangers or game wardens may use horses for patrols, and horses or mules may
also be used for clearing trails or other work in areas of rough terrain where vehicles are less
effective.[199]
Although machinery has replaced horses in many parts of the world, an estimated 100 million horses,
donkeys and mules are still used for agriculture and transportation in less developed areas. This
number includes around 27 million working animals in Africa alone.[200] Some land management
practices such as cultivating and logging can be efficiently performed with horses. In agriculture, less
fossil fuel is used and increased environmental conservation occurs over time with the use of draft
animals such as horses.[201][202] Logging with horses can result in reduced damage to soil structure
and less damage to trees due to more selective logging.[203]
Warfare
Horses have been used in warfare for most of recorded history.

The first archaeological evidence of horses used in warfare dates to
between 4000 and 3000 BCE,[204] and the use of horses in
warfare was widespread by the end of the Bronze Age.[205][206]
Although mechanization has largely replaced the horse as a
weapon of war, horses are still seen today in limited military uses,
mostly for ceremonial purposes, or for reconnaissance and
transport activities in areas of rough terrain where motorized
vehicles are ineffective. Horses have been used in the 21st century Ottoman cavalry, 1917
by the Janjaweed militias in the War in Darfur.[207]
Entertainment and culture
Modern horses are often used to reenact many of their historical work
purposes. Horses are used, complete with equipment that is authentic or
a meticulously recreated replica, in various live action historical
reenactments of specific periods of history, especially recreations of
famous battles.[208] Horses are also used to preserve cultural traditions
and for ceremonial purposes. Countries such as the United Kingdom still
use horse-drawn carriages to convey royalty and other VIPs to and from
certain culturally significant events.[209] Public exhibitions are another
example, such as the Budweiser Clydesdales, seen in parades and other
public settings, a team of draft horses that pull a beer wagon similar to
that used before the invention of the modern motorized truck.[210]
Horses are frequently used in television, films and literature. They are The horse-headed deity in
sometimes featured as a major character in films about particular Hinduism, Hayagriva
animals, but also used as visual elements that assure the accuracy of
historical stories.[211] Both live horses and iconic images of horses are
used in advertising to promote a variety of products.[212] The horse frequently appears in coats of
arms in heraldry, in a variety of poses and equipment.[213] The mythologies of many cultures,
including Greco-Roman, Hindu, Islamic, and Norse, include references to both normal horses and
those with wings or additional limbs, and multiple myths also call upon the horse to draw the chariots
of the Moon and Sun.[214] The horse also appears in the 12-year cycle of animals in the Chinese zodiac
related to the Chinese calendar.[215] Horses serve as the inspiration for many modern automobile
names and logos, including the Ford Pinto, Ford Bronco, Ford Mustang, Hyundai Equus, Hyundai
Pony, Mitsubishi Starion, Subaru Brumby, Mitsubishi Colt/Dodge Colt, Volkswagen Polo, Pinzgauer,
Steyr-Puch Haflinger, Pegaso, Porsche, Rolls-Royce Camargue, Ferrari, Carlsson, Kamaz, Corre La
Licorne, Iran Khodro, Eicher, and Baojun.[216][217][218]
Therapeutic use
People of all ages with physical and mental disabilities obtain beneficial results from an association
with horses. Therapeutic riding is used to mentally and physically stimulate disabled persons and help
them improve their lives through improved balance and coordination, increased self-confidence, and a
greater feeling of freedom and independence.[219] The benefits of equestrian activity for people with
disabilities has also been recognized with the addition of equestrian events to the Paralympic Games
and recognition of para-equestrian events by the International Federation for Equestrian Sports
(FEI).[220] Hippotherapy and therapeutic horseback riding are names for different physical,
occupational, and speech therapy treatment strategies that use equine movement. In hippotherapy, a
therapist uses the horse's movement to improve their patient's cognitive, coordination, balance, and
fine motor skills, whereas therapeutic horseback riding uses specific riding skills.[221]
Horses also provide psychological benefits to people whether they actually ride or not. "Equine-
assisted" or "equine-facilitated" therapy is a form of experiential psychotherapy that uses horses as
companion animals to assist people with mental illness, including anxiety disorders, psychotic
disorders, mood disorders, behavioral difficulties, and those who are going through major life
changes.[222] There are also experimental programs using horses in prison settings. Exposure to
horses appears to improve the behavior of inmates and help reduce recidivism when they leave.[223]
Products
Horses are raw material for many products made by humans throughout history, including
byproducts from the slaughter of horses as well as materials collected from living horses.
Products collected from living horses include mare's milk, used by people with large horse herds, such
as the Mongols, who let it ferment to produce kumis.[224] Horse blood was once used as food by the
Mongols and other nomadic tribes, who found it a convenient source of nutrition when traveling.
Drinking their own horses' blood allowed the Mongols to ride for extended periods of time without
stopping to eat.[224] The drug Premarin is a mixture of estrogens extracted from the urine of pregnant
mares (pregnant mares' urine), and was previously a widely used drug for hormone replacement
therapy.[225] The tail hair of horses can be used for making bows for string instruments such as the
violin, viola, cello, and double bass.[226]
Horse meat has been used as food for humans and carnivorous animals throughout the ages.
Approximately 5 million horses are slaughtered each year for meat worldwide.[227] It is eaten in many
parts of the world, though consumption is taboo in some cultures,[228] and a subject of political
controversy in others.[229] Horsehide leather has been used for boots, gloves, jackets,[230]
baseballs,[231] and baseball gloves. Horse hooves can also be used to produce animal glue.[232] Horse
bones can be used to make implements.[233] Specifically, in Italian cuisine, the horse tibia is
sharpened into a probe called a spinto, which is used to test the readiness of a (pig) ham as it
cures.[234] In Asia, the saba is a horsehide vessel used in the production of kumis.[235]
Care
Horses are grazing animals, and their major source of nutrients is

good-quality forage from hay or pasture.[236] They can consume
approximately 2% to 2.5% of their body weight in dry feed each
day. Therefore, a 450-kilogram (990 lb) adult horse could eat up
to 11 kilograms (24 lb) of food.[237] Sometimes, concentrated feed
such as grain is fed in addition to pasture or hay, especially when
the animal is very active.[238] When grain is fed, equine
Checking teeth and other physical nutritionists recommend that 50% or more of the animal's diet by
examinations are an important part weight should still be forage.[239]
of horse care.
Horses require a plentiful supply of clean water, a minimum of 38
to 45 litres (10 to 12 US gal) per day.[240] Although horses are
adapted to live outside, they require shelter from the wind and precipitation, which can range from a
simple shed or shelter to an elaborate stable.[241]
Horses require routine hoof care from a farrier, as well as vaccinations to protect against various
diseases, and dental examinations from a veterinarian or a specialized equine dentist.[242] If horses
are kept inside in a barn, they require regular daily exercise for their physical health and mental well-
being.[243] When turned outside, they require well-maintained, sturdy fences to be safely
contained.[244] Regular grooming is also helpful to help the horse maintain good health of the hair
coat and underlying skin.[245]
See also
Glossary of equestrian terms
Lists of horse-related topics
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2/1/23, 8:39 AM Horse - Wikipedia

1.1Lifespan and life stages

1.2Size and measurement

2Taxonomy and evolution

2.1Wild species surviving into modern times

4Interaction with humans

Lifespan and life stages

Depending on breed, management and

The following terminology is used to describe horses of various ages:

Size and measurement

In English-speaking countries, the height of horses is

The size of horses varies by breed, but also is

Colors and markings

Many genes that create horse coat colors and patterns

Reproduction and development

Gestation lasts approximately 340 days, with an average range

Horses, particularly colts, are sometimes physically capable of

The horse skeleton averages 205 bones.[60] A

Gallop 40–48 km/h

All horses move naturally with four basic gaits:[84]

Horses are prey animals with a strong fight-or-flight response. 0:03

Intelligence and learning

"Hot blooded" breeds include "oriental horses"

Muscular, heavy draft horses are known as "cold

Horses are able to sleep both standing up and lying down. In an

Taxonomy and evolution

browsing of drier plant material, then to grazing of

By about 15,000 years ago, Equus ferus was a

Wild species surviving into modern times

A truly wild horse is a species or subspecies with no ancestors that

The Przewalski's horse (Equus ferus przewalskii), named after the

Other modern equids

Interaction with humans

Horses are trained to be ridden or driven in a variety of

There are certain jobs that horses do very

Horses have been used in warfare for most of recorded history.

Entertainment and culture

Horses are grazing animals, and their major source of nutrients is

3. International Commission on Zoological Nomenclature (2003). "Usage of 17 specific names based

27. Bongianni, entries 86, 96, 97

99. Belknap, p. 255

125. "The Evolution of Horses" (http://www.amnh.org/exhibitions/horse/?section=evolution&page=evolu

182. "FAO Stat – Live Animals" (http://faostat.fao.org/site/573/DesktopDefault.aspx?PageID=573#anco

235. Diseases of the Stomach and Intestines (https://archive.org/details/diseasesstomach00dujagoog).

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