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Study and
1MENT Description of the
Flowers
INTRODUCTION
Flower is basically ashoot havinga condensed axis called thalamus and anumber of appendages
called floral leaves. Atypical flower has four types of fAoral leaves called sepals, petals, stamens and carpels
in distinct whorls known as calyx, corolla, androecium and gynoecium respectively. The axís on which a
lower arises is called mother axis. The flower may possess a stalk called pedicel. The pedicel may be borne
in the axi of a small leaf like structure called bract. Rarely the pedicel may bear one or more leafy bract
Hike structures called bracteoles. Sometimes a whorl of bracteoles is present below the calyx. This whorls
is called epicalyx.
IMPORTANT TERMS
1. Bracteate. Aflower having bract at its base.
2. Ebracteate. A fower without a bract.
3. Pedicellate. Aflower having a stalk or pedicel.
4. Sessile. Aflower without a stalk or pedicel.
5. Complete. Aflower in which all the four whorls are present.
6. Incomplete. Aflower in which any one or more whorls is/are absent.
7. Actinomorphic. Aflower that can be divided into two equal halves through any vertical plane,
eg, Petunia.
Carpel
(Gynoecíum) Stamen
(Androecium)
Petal
(Corolla)
Sepal
(Calyx)
Thalamus
B. Zygomorphic B. Corolla
It is the second whorl of flower and is composed of variously coloured
petals.
Forms of Corolla
A. Actinomorphic C. Asymmetrical 1. Polypetalous. Corolla consists of free petals. Polypetalous corrolla has following forms :
Fig. 1.2.A. Actinomorphic flower; B. Zygomorphic flower;C. Asymmetrical. () Cruciform. Consists of four, free, clawed petals, arranged in the form
ofa cross. eg,
Mustard.
(ii) Caryophyllace ous. Consists of five clawed petals in which limbs of the petals are placed
INSERTION OF FLORAL LEAVES at right angle to the claws. e.g, Dianthus (Pinlk).
1. Hypogynous. The ovary occupies the highest position on thalamus and all other floral organs (iiil) Rosaceous. Consists of five or more sessile petals or with very short claws and their limbs
are situated below the ovary. e.g, Mustard, shoe flower. are spread outwards. e.g., Rose.
2. Gamopetalous. Corolla composed of fused petals. Following are the important forms of
gamopetalous corolla:
(i) Campanulate. Bell shaped. e.g, Campanula.
(i) Tubular. Tube like. e.g, Sunflower.
(ii) Infundibuliform. Funnel shaped. e.g, Petunia.
Thalamus - (iv) Rotate. More or less wheel shaped. e.g, Exora.
-Ovary
Thalamus -Ovule () Bilabiate. Bilipped. eg, Ocimum.
Ovary (vi) Ligulate. With narrow tube below and strap shaped structure above. e.g., Ray foret of
(Epigynous) sunflowe.
Superior ovary (Hypogynous) (Perigynous)
Superior ovary Inferior ovary
(vi) Papilionaceous. Five, free petals are so arranged as to resemble abutterfly. It consists
Fig. 1.3. Position of floral whorls on Thalamus. of an outer most, the largest vexillum or standard, two laterals or wings and two anterior
fused petals called keel, e.g, Lathyrus.
O000O0
RosacooUs Campanulato
Cruciform Caryophylaceous
C. Androecium
Infundibuliform
Tubular Rotato It is third whorl of the Aower and is composed of male reproductive organs, stamens. Each stamen
consists of a filament and anther.
Standard
Wing
Papilionaceous
Bilabiato Ligulate
Fig. 14. Shapes of Corolla. Totradynamous Diadalphous Didynamous
Aestivation. Aestivation is the arrangement of sepals or petals in the floral bud, with respect
to the members of the same whorl (calyx or corolla).
() Valvate. When sepals or petals meet by their edges without overlapping,
() Twisted. One margin of each sepal or petal overlaps the next and the other margin is
overlapped by the previous sepal or petal.
(i) Imbricate. There are five sepals or petals. One is completely overlapped by the adjacent
members and one is completely overlapping the adjacent members. The other three members
are partially outside and partially inside. eg, Cleome.
(iv) Ascending imbricate. The posterior member is held inside by the upper margins of two Monoadalphous Polyadolphous Synandrous
Syngenesious
(Synantherous)
laterals. They are in turn overlapped by the upper margin of two anterio-lateral members.
eg, Cassia. Fig. 1.6.Forms of Androecium.
() Descending imbricate (Veillary). The posterior petal is large and overlaps the two
lateral petals. The later overlap the two anterior petals. e.g, Lathyrus.
26
Canroteaw Laboratoy Manval in Biology y Core Experiment-1 27
Forms of Androecium 2. Axile. Placentae arise from the axils of polycarpellary, multilocular, ovary projecting towards
1. Polyandrens Stamens free, not attache to any whorl. periphery, eg China-rose, Petunla, Onion.
2. Ipipetalous, Stamens fused with the petals, eg. Petunia, Slamm.
3. DidynamOns. Two short and two long stamens in a flower. eg., Ocimum.
4. Tetradynamous. Six stansens in two whorls, two short stanmens in outer whorl and four lon
stamens in the inner whorl eg Brassia. A
Cohesion of Stamens
Stamens are variously fused among themselves
1. Monadelphous. Anthers free but all the filaments are fused into one bundle, eg. Hiblscus.
2. Diadelpheus. Anthers are free but filaments are fused in two bundles eg, Lathyrus.
3. Polyandrous. Filaments are fused in many bundles with free anthers, eg. Citrus.
4. Syngenesious. The anthers are fused but filaments are free, eg, Sunflower. E F
5. Synandroas. Stamens are fused both in the regions of filament and anther, eg. Cucurbita:
Fig. 1.8. Various types of Placentation. A. Marginal; B. Axile; C. Parietal:
D. Free-central; E. Basal ; F. Superficial.
D. Gynoecium
This is the innermost whotl consisting of female reproductive organs-carpels. Acarpel consists of 9. Parietal. Two or more longitudinal placentae develop along the wall of asyncarpous unilocular
stigma, style and ovary. Folowing arethe forms of gynoecium: ovary, eg, Poppy, Mustard.
1. Monocarpellary. Gynoecium with one carpel, eg. Sweet Pea. 4. Free central. Placenta arise from central axis of one-chambered, polycarpellary, syncarpous
2. Bicarpellary. Gynoecium with two carpels, eg, Petunia. ovary, e.g., Dianthus.
3. Polycarpellary. Gynoecium with many carpels, eg. China-rose. 5. Basal. Monocarpellary ovary has single locule with one placenta attached to the base of the
ovary, e.g., Sunflower.
Carpels are of two type, depending upon fusion: 6. Superficial. Multicarpellary, syncarpous, multiocular ovary, placentae are scattered all around
1. Apocarpous.Carpels free from each other.
the partition wall, eg. Water lily.
2. Syncarpous. Carpels fused with each other, eg, China-rose.
Locules in Ovary Floral Fomula
Chambers in the ovary are called locules. They may be one, two or many, hence the ovaries are called It represents the floral characters of aflower by abbreviations and symbols. The symbols used
unilocular, bilocular or multilocular ovaries respectively. for representing the floral characters are as follows:
Br-Bracteate Ebr-Ebracteate
-Actinomorphic t-Zygomorphic
-Hermaphrodite or bisexual o-Male or staminate
Q-Female or pistillate N-Neuter
Undocular Bilocular Trilocular Tetralocular Pentalocular
Epi--Epicalyx K-Calyx, sepals
C-Corolla, petals A-Androecium, stamens
Fig. 1.7.One to five chambered ovaries.
G-Gynoecium, carpels G-Superior ovary
Placentation G-Inferior ovary CA-Epipetalous stamen.
The arrangement of placentae bearing ovules inside the ovary ís called placentation. It is of following The number of individual member present in the whorl is indicated by writing appropriate figure
types : at the foot of the symbol. If the members of the whorl are fused among themseves, their number is
1. Marginal, Single placenta is attached to the marginal suture of monocarpellary ovary, eg, Pea. put in a bracket.
Cnprekenstne Laboratory Manual iin Biology--X1
28 Core Experiment-1 29
Floral Diagram the floral lbud in diagram by T.S. of the ovary. The nectary glands are shown by dots in their respective position. The
representation of cross-section of relation to bract is shown outside the sepals usually opposite to the mother ais.
Floral diagram iis a diagrammaticcharacters of the ffower which are not representedlby its floral
mother axis, and it shows manysuch characters of the flower:
its tells us about the following
formula. The Aoral diagram of a flower
absence of bract and bracteoles.
1. Presence or the mother axis). Normallyythe odd sepal
(the sepal which is in line of
2. The position of odd sepal it is anterior.
family Lcguminasac where
is posterior except in relation to one another.
arrangement of fforal leaves in
3. The number and and adhesion (fusion of
foral leavesamongst themselves) Aoral leaves
4. The cohesion (fusion of whorl) of fAoral leaves.
with fAoralleaves of other
A
S. Symmetry of the Bower.
Fig. 1.9. Floral diagrams. A Petunia alba ; B. Brassica campestris.
6. Aestivation of sepals and petals.
bithecous nature of anther.
7. The monothecous and Floral formula and floral diagram of a lower help us in identification of a flower. Therefore, for
placentation.
8. Number of locules in the ovary and acomplete description and correct identification of fower, both Aoral formula and loral diagram are
9. Nectary glands and their position. necessary.
)X((oesg*o
3. Epicalyz
4 Calyx or sepals
AIM: To study and describe the given flowering plant (1. Petunia, 2. Lathyrus, 3. Asparagus, 4. Allim) and
display of whorls and anther and ovary to show number of chambers.
REQUIREMENTS
5. Corolla or petals Flower, forceps, razor or blade, slides, needles-2, coverslip, microscope, white paper.
6. Stamens (bithecous)
PROCEDURE
7. Stamens (monothecous)
1. Take one suggested fAowering plants and make the following observations
8. Nectar secreting disc around carpel 2. Note down the type of root (tap or adventitious), stem, herbaceous or woody) and leaf
9. Monocarpellary gynoecium
(arrangement, shape, variation, simple or compound) of the plant.
3. Note down, whether its Aower has a pedicel, bracts, epicalyx or not.
10. Bicarpellary syncarpous gynoecium 4. Observe the presence of different whorls of foral leaves (ie, calyx, corolla, androecium and
gynoecium) and insertion of foral leaves.
11 Palycarpelary syncarpous gynoecium 5. Carefully take out floral leaves of each whorl and place them whorl wise on a white paper ie,
12 Monoadelphous androecium epicalyx (if present), sepals, petals, stamens and carpels.
6. Count the number of foral leaves of each whorl and note down their cohesion (union among
13. Syngenecious androecium. the members of the same whorl) or adhesion (union with the Roral leaves of other whor).
In the foral diagram the mother aris is shown by acircular dot in the posterior position. Ihe 7. With the help of a blade or razer, cut thin sections of anther and ovary, Mount the secticn on
epicalyz, sepals, petals or tepals, stamens and carpels are drawm by their respective symbols, in their aslide in a drop of water and examine under the dissecting microscope. Count the number of
respective position. The aestivation of sepals and petals is shown by overlapping their margins and chambers in anther and ovary and number of ovules in each chamber locule.
cohesion by joining their marzins. The epipetalous or epiphyllous starnen is shown byjoiningthe middle 8. Find out the nature of ovary, style, stigma and placentation.
of the symbol of the anther to the petal or tepal by aline. Gynoecium is shown in the centre offoral 9. Observe the floral characters. Try to write foral formula and draw the Rloral diagram of the Roer.
ComprckeasteLaboratory Manuallin
30 Biology-0 Core Experiment-1 31
ALBA-PETUNIA
TYPE 1. PETUNIA Anthor lobe
Classification
-Angiospermae
Division Stigma
- Dicotyledonae Flower Potal
Connective
Class -Stamen -Filament
-Solanaceae
Eamily Slamen
-Petunia Style
Genus
-alba Flower
Species bud Ovary -Sepal
-Tap root, branched
Root Podicel -Thalamus
- Herbaceous, erect, hairy Stigma LS. flower
Stem
-Alternate, opposite in the florallregion, petiolate,,
Leaf ovate, hairy.
simple, Flowering twig
-Stylo ovary wal.
-Solitary, axillary. Ovule
Inflorescence
- Ebracteate, pedicellte, complete, bisexual, actinomorphi: Placenta
Flower
pentamerous, hypogynous.
Ovary. T.S. of ovary
Calyx
- Sepals 5, gamosepalous, valvate aestivation, persistent, Floral diagram
inferior, green, hairy.
Gynoecium
Corolla
-Petals 5, gamopetalous, infundibuliform, valvate
aestivation, white or purplish in colour. Fig. 1.10. Aflowering twig, parts of flower and floral diagram of Petunia.
Androecium -Stamens 5, alternipetalous, epipetalous, anther
bithecous, basifixed.
1. Persistent sepals.
2. Infundibuliform corolla.
3. Stamen epipetalous.
4. Ovary obliquely placed, placentation axile with swollen placenta.
Some more similar Aowers are-Solanum tuberosum (potato), Solanum nigrum(Makoy).
32 Cmrelcmsne Laboratory Manual in Biology Core Experiment-1 33
.vule
Ovary wal
TS. of ovary
Fig. 1.11. Aflowering twig. parts of aflower and floral diagram of Lathy rsodorats
realme
Comtrckenslue Laboratory Manual ini
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OFFICINALIS-SATAVARI
Biology- Core Experiment-1 35
TYPE 3. ASPARAGUS
Classification
-Angiospermae
Division
-Monocotyledonae -Cladodes
Class
Liliaceae MMe Flowers
Family
-Asparagus
Genus
-officinalis -Peduncle
Species -Adventitious, tuberous to store food.
Root Bract
-Erect, aerial, woody.
Stem
Leaf
- Scaly or modifed into spines bearing axillary cadol <Leaf spine
- Racemose.
Infßorescence
pedicellate, AFlowering Branch
Flower
-Bracteate, incomplete, bisenual
actinomorphic, hypogynous, trimerous.
-Tepals 6, arranged in two whorls of 3
Perianth each, ightyi
camophyllous at the base, petaloid, white, eachsltenal
marked with a brown midrib.
Floral Formula
V.S.Flower
Floral diagram
Reasons for Identification
1 Flower trimerous.
Ovary wall
2. Petaloid perianth in two whorls of 3each.
3. Stamens 6in two whorls of 3each, epiphyllous. Ovules
-Locule
4. Ovary tricarpellary, trilocular with ale placentation. -Placentae
Some similar Aowers-A_phodelus tenuifolius (Piazi).
T.S. of ovary
Classification Inflorescence
Divisson -Angiospermae
Class -Monocotyledonae
Family -Liliaceae
Flower bud -Bract
Genus -Allium
-Scape
Specdes -cepa
Root
-Adventitious, fibrous
Inflorescene
Stem - Underground, bulb, inner scales feshy outer dry
membranous and brown.
Stigma
Leaf Radical, cylindrical, sheathing,
Umbellate cyne. -Leaf
Infloress cence Perianth
Flower -Bracteate, pedicillate, incomplete, bisexual,
actinomorphic, hypogynous, trimerous.
Anther
Perianth - Tepals 6, arranged in two whorls of 3 each, slightly A fiower
gamophyllous at the base, petaloid, white in colour, each Tunicated bulb Anther Stigma
tepal is marked with a brown midrib.
Filament
Androecium -Stamens 6,polyandrous, arranged in two whorls of3each,
antiphyllous, epiphyllous, anther bithecous, basifixed. Style
Gynoecium -Tricarpellary, syncarpous, ovary superior, trilocular each Perlanth
locule with two ovule, axile placentation, style short, -Reduced stem Thalamus
stigma trilobed. Pistil Ovary
Roots Pedicel
Floral Formula Mother axis LS. of flower
Flowering twig
Reasons for Identification
1 Flower trimerous.
-Ovary wall
2 Petaloid perianth in two whorls of 3 each.
Locule
3. Stamens six in two whorls of 3 each, epiphyllous.
4. Ovary tricarpellary, trilocular with adile placentation. Ovule
T. S, of ovary
Floral diagram
Fig. 1.13. Flowering twig, parts of flower and floral diagram of Allium cepa (Onion).
38 Comtrekemsie Laboratory Manual in Biology 39
Core Experiment--1
TYPE 5. BRASSICA CAMPESTRIS-MUSTARD
Flowers
Classifcation Long Stigma
Stigma starmen Petal
Division -Angiospermae
Class -Dicotyledonae
Family -Bassicaceae (Cruciferae)
Pedicel Short
Gemus -Brassica style
-Ovary Lateral
-pestris Peduncle Replum
Species stamen
1 Calyx arranged in two whorls of two each. Fig. 1.14. Brassica campestris. A. a flowering branch; B. VS. flower (antero-posterior
2. Corolla 4, polypetalous cruciform
plane); C. V.S. flower (lateral plane) ; D. petal ;E. flower with non-essential
parts removed; F. floral diagram.
3. Androecium 6, tetradynamous.
4. Gynoecium bicarpellary, syncarpous, superior.
5. Ovary bilocular due to the presence of false septum/replum, placentation parietal.
realmeShot on realme 7
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40 Compreheaste Laboratory Manual in Biology-y Core Experiment-1
TYPE 6. HELIANTHUS ANNUUS SUNFLOWER
Classifcation
Division -Angiospermae
Class Dicotyledonae
Family
Asteraceae (Compositae)
Genus Helianthus
annuus
Species
Root Tap root
Stem Erect, herbaceous, hairy
Leaf -Alternate, simple, hairy, reticulate venation
Infiorescence Racemose head or capitulum. It consists of central tubular dise Corolla limb
Corolla tube
forets and the peripheral ligulate ray florets. The capitulurn
is usually surrounded by whorls of bracts collectively called Calyx
Thalamus
involucre.
C D
Ray Florets Flower -Bracteate, sessile, incomplete, either pistillate or neuter,
zygomorphic, epigynous. -Stlgma
Calyx -Represented by 2-3 small free white scales.
Corolla -Sepals 3-5, gamopetalous, ligulate (formed of asmalbasal Corolla
Fused
anthers
hairy tube and a large strap shaped limb) tube
Androecium -Absent Sepal -Style
Androecium Stamens 5, syngenesious or synanthrous (filament freeanthers Fig. 1.15. Helianthus annuus. A. a flowering branch; B. V.S. capitulum; C. ray floret
are fused), epipetalous D.V.S. ray floret; E. disc floret; F. V.S. disc floret; G. floral diagram of ray floret
Gynoecium - Bicarpellary syncarpous, ovary inferior unilocular, basal H. floral diagram of disc floret.
placentation.
Floral Formula Ray Florets: Br %Qor Neuter K,-3 scales Ca-5A, G2) or0
Disc Florets: Br ØK,_3scales C(5) A(5) G(2)
Reasons for Identification
1. Inflorescence capitulum
2. Flowers are small called Alorets, which are ligulate or tubular, epigynous.
3. Calyx reduced and represented by scales.
4. Androecium 5, syngenesious and epipetalous.
5. Gynoecium bicarpellary, syncarpous, ovary inferior, unilocular with basal placentation.
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Floral Formula Br Brl %O P,lodicules A, G Fig. 1.16. Triticum aestivum. A. inflorescence; B. opened spikelet C lemma;
D. flower with lemma removed; E. VS. flower or floret F. floral digram.
Reasons for Iåentification
L Inforescence spike of spikelets.
2 Each spike is surrounded by two scales called glumes.
3. Each fower is covered over by two scales, lernna and pale.
4. Flower is incomplete and zygomorphic.
5. Perianth represented by two lodicules.
6. Orary monocapellary with unilocular, uniovuled ovary, having basal placentation.