Evolution of Stem Driven Computer Science Education The Perspective of Big Concepts Vytautas Stuikys Online Ebook Texxtbook Full Chapter PDF
Evolution of Stem Driven Computer Science Education The Perspective of Big Concepts Vytautas Stuikys Online Ebook Texxtbook Full Chapter PDF
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Vytautas Štuikys and Renata Burbaitė
Renata Burbaitė
Department of Software Engineering, Kaunas University of Technology,
Kaunas, Lithuania
Juozas Balčikonis Gymnasium, Panevėžys, Lithuania
This work is subject to copyright. All rights are solely and exclusively
licensed by the Publisher, whether the whole or part of the material is
concerned, specifically the rights of translation, reprinting, reuse of
illustrations, recitation, broadcasting, reproduction on microfilms or in
any other physical way, and transmission or information storage and
retrieval, electronic adaptation, computer software, or by similar or
dissimilar methodology now known or hereafter developed.
The publisher, the authors, and the editors are safe to assume that the
advice and information in this book are believed to be true and accurate
at the date of publication. Neither the publisher nor the authors or the
editors give a warranty, expressed or implied, with respect to the
material contained herein or for any errors or omissions that may have
been made. The publisher remains neutral with regard to jurisdictional
claims in published maps and institutional affiliations.
Vytautas Štuikys
Renata Burbaitė
Kaunas, Lithuania
Kaunas/Panevėžys, Lithuania
Abbreviations
AI Artificial Intelligence
ANN Artificial Neural Network
BC Big Concept
BD Big Data
CB LO Component-Based Learning Object
CBL Contest (Competition)-Based Learning
CL Collaborative Learning
CS Computer Science
CT Computational Thinking
DB Database
DL Deep Learning
DS Data Science
DT Design Thinking
ESLE Extended Smart Learning Environment
GLO Generative Learning Object
IoT Internet of Things
ISTEM Integrated STEM
ISTEM SM Integrated STEM Skills Model
LKSAM Learner’s Knowledge and Skills Assessment Module
LO Learning Object
LO LG Learning Object List Generator
MDG Metadata Generator
ML Machine Learning
NLP Natural Language Processing
PCB LO Personalised Component-Based Learning Object
PGL Personal Generative Library
PGLO Personalised Generative Learning Object
PL Personalised Learning
PLE Personalised Learning Environment
PSLO Personalised Smart Learning Object
QG Query Generator
SLE Smart Learning Environment
SLO Smart Learning Object
SPGL Student’s Personal Generative Library
SR Speech Recognition
SRS Speech Recognition System
ST Scientific Thinking
STEM Science, Technology, Engineering and Mathematics
TPACK Technological, Pedagogical And Content Knowledge
TPL Teacher’s Personal Library
VTT Voice-to-Text
Contents
1 Context and Model for Writing This Book:An Idea of Big
Concepts
1.1 Introduction
1.2 A Short Glance to Education Evolution
1.3 A Short Glance to Computing Evolution
1.4 A Short Glance to STEM Evolution
1.5 A Short Glance to Computational Thinking Skills
1.6 Context Model to Define Our Approach
1.7 Evolutionary Model for Change
1.8 The Topics This Book Addresses
1.9 Concluding Remarks
References
Part I Pedagogical Aspects of STEM-Driven CS Education Evolution:
Integrated STEM-CS Skills Model, Personalisation Aspects and
Collaborative Learning
2 Models for the Development and Assessment of Integrated STEM
(ISTEM) Skills:A Case Study
2.1 Introduction
2.2 The Aim and Motivation
2.3 Research Tasks and Methodology
2.4 Related Work
2.5 Defining Context and Functionality for STEM-CS Skills
2.6 Defining the Structure of STEM-CS Skills Model
2.7 Analysis of the Interdependencies Among Different Skills
2.8 Feature-Based STEM-CS Skills Model (RQ3)
2.9 Analysis of Metrics and Defining Metrics Model for Skills
Evaluation
2.10 Model for Evaluating and Describing of the ISTEM-CS Skills
2.11 Validation of the ISTEM-CS Skills Model Through Case
Study (RQ6)
2.12 ISTEM-CS Skills and Their Metrics Generating Tool
2.13 Summarising Discussion and Evaluation
2.14 Conclusion
Appendix
References
3 Enforcing STEM-Driven CS Education Through Personalisation
3.1 Introduction
3.2 Related Work
3.3 Requirements for Personalised STEM-Driven CS Learning
and Research Questions
3.4 Basic Idea and Methodology
3.5 Background
3.6 A Framework for Implementing Personalised STEM-Driven
CS Education
3.6.1 Structural Models of Personalised LOs
3.6.2 Personalised Processes and Activities Within the
Framework
3.6.3 Tools and Approaches to Implement the Proposed
Framework
3.7 Case Study
3.8 Discussion and Concluding Remarks
References
4 Personal Generative Libraries for Personalised Learning:A Case
Study
4.1 Introduction
4.2 Related Work
4.3 The Concept of the Personal Generative Library
4.4 Methodology and Background
4.5 Structure and Functionality of PGL
4.6 Integration of PGLs into the Framework of Personalised
Learning
4.7 Case Study and Results
4.8 Discussion and Evaluation
4.9 Conclusion
References
5 Enforcing STEM-Driven CS Education Through Collaborative
Learning
5.1 Introduction
5.2 Related Work
5.3 Basic Idea of the Approach and Methodology
5.4 The Concept ‘Real-World Task’ in STEM Research and Its
Complexity
5.4.1 Complexity Issues of Real-World Tasks
5.4.2 Conceptual Model for Solving Real-World Tasks
5.5 Framework for STEM-Driven Contest-Based Collaborative
Learning
5.6 Case Study
5.7 Discussion and Evaluation
5.8 Conclusion
Appendix
References
Part II Internet of Things (IoT) and Data Science (DS) Concepts in
K–12 STEM-Driven CS Education
6 Methodological Aspects of Educational Internet of Things
6.1 Introduction
6.2 Related Work
6.3 Research Strategy, Aim, and Requirements
6.4 Motivation and Basic Idea
6.5 Background:Conceptual Modelling of IoT
6.6 A Framework for Introducing IoT into STEM-CS Education
6.7 Interpretation of IoT Architecture for STEM-Driven CS
Education
6.8 Discussion on Proposed Methodology
6.9 Conclusion
References
7 Multi-stage Prototyping for Introducing IoT Concepts:A Case
Study
7.1 Introduction
7.2 Related Work
7.3 Methodology:Implementation Aspects Through Modelling
7.3.1 A Multi-stage Model for Introducing IoT into STEM-
Driven CS Education
7.3.2 A Framework for Solving Real-World Tasks Through
IoT Prototyping
7.3.3 A Detailed Specification of IoT Prototype Design
Processes
7.3.4 IoT Prototyping Task Solving Through Inquiry-Based
and Design-Oriented Collaborative Learning
7.4 Extending Smart Learning Environment with Tools for IoT
Prototyping
7.5 Case Study
7.6 Summarising Discussion and Evaluation
7.7 Conclusion
References
8 Introducing Data Science Concepts into STEM-Driven Computer
Science Education
8.1 Introduction
8.2 Related Work
8.3 Motivation and Research Methodology
8.4 Conceptual Model for Introducing DS Concepts into K–12
8.5 Implementation of the Methodology:A Three-Layered
Framework
8.6 Development of the DS Model
8.7 Extending Smart Learning Environment
8.8 Modelling for Developing the Task Solution System
8.9 Development of the Assessment Model
8.10 A Case Study and Experiments
8.11 Summarising Discussion and Evaluation
8.12 Conclusion
References
Part III Introduction to Artificial Intelligence
9 A Vision for Introducing AI Topics:A Case Study
9.1 Introduction
9.2 Related Work
9.3 Background and AI Key Concepts
9.4 A Framework for Introducing AI Topics
9.5 Methodology for Implementing the Proposed Framework
9.6 Generic Architecture for Introducing AI Tools into SLE
9.7 Adopted Generic Scenario for Delivery of the AI Content
9.8 Summarising Discussion and Conclusion
References
10 Speech Recognition Technology in K–12 STEM-Driven Computer
Science Education
10.1 Introduction
10.2 Related Work
10.3 Basic Idea with Motivating Scenario
10.4 Background
10.5 Research Methodology
10.6 Extending Smart Learning Environment for Speech
Recognition Tasks
10.7 Case Study to Support Task 1
10.8 Case Study to Support Task 3
10.9 Summarising Discussion and Conclusions
Appendix 1
Appendix 2
Appendix 3
References
11 Introduction to Artificial Neural Networks and Machine
Learning
11.1 Introduction
11.2 Related Work
11.3 Operating Tasks and Methodology
11.4 Background:Basic Concepts and Models of ANNs (RQ2)
11.5 Motivating Example:A Binary Classification (RQ3)
11.6 Case Study 1:Implementation of Single-Layered
Perceptron Model
11.7 Case Study 2:Implementation of Multi-Layered Perceptron
Model
11.8 Summarising Discussion and Evaluation
11.9 Conclusion
References
12 Overall Evaluation of This Book Concepts and Approaches
12.1 Aim and Structure of This Chapter
12.2 What Is the Contribution of This Book?
12.3 Difficulties and Drawbacks of the Proposed Approach
12.4 Rethinking of Discussed Approach
12.5 STEM-Driven Precision Education:A Vision Inspired by
Concepts Discussed in This Book
12.6 Topics for Future Work
References
Index
© The Author(s), under exclusive license to Springer Nature Switzerland AG 2024
V. Štuikys, R. Burbaitė, Evolution of STEM-Driven Computer Science Education
https://doi.org/10.1007/978-3-031-48235-9_1
1.1 Introduction
The title of this book begins with the term ‘evolution’ to define the
recent changes and innovations in the so-called STEM-driven computer
science (CS) education. By this paradigm, we mean concepts, ideas, and
approaches presented in the published book by Springer in 2018 under
the title “Smart STEM-Driven Computer Science Education: Theory,
Methodology and Robot-based Practices”. What is this book about?
Shortly, that book [ŠB18] is about how three fields, i.e., STEM, CS, and
Educational Robotics, are to be (or could be) combined seamlessly into
a coherent educational methodology to gain synergistic benefits from
all. More specifically, the basic idea we have discussed there focused on
(1) the high variability of STEM pedagogy, variability of CS-related
content, and variability of the available technology used and (2) how to
project these variabilities onto high school CS curriculum using modern
technologies of two types. The first type is educational robotics as
direct facilities for learning for introducing interdisciplinary knowledge
(i.e., scientific, engineering, and technological ones the robotics highly
promotes) in a high (secondary) school. The second type (borrowed
from software engineering) is supporting technologies for the
automated preparation, design, and use of the content and learning
processes. The supporting technologies include high-level modelling
aiming at defining and expressing the learning variabilities at a high
abstraction level through feature models and then providing
transformations of those into the executable code of the teaching
content using meta-programming techniques. Note that we have
integrated those technologies seamlessly using conventional software
only.
Since 2018, when the book [ŠB18] appeared, many innovations
have taken place in STEM and CS education research and practice
worldwide. We have also accumulated new knowledge and new
scientific results due to our intensive research in this field. The core of
that is the novel or extended approaches we have developed since that
time. Those rely on the concepts we have introduced in the title as Big
Concepts. Why do we use the term “Big Concepts” and what they are?
We will present a more extended motivation for using this term later.
Here, the reader should accept the following terms (Internet of Things
(IoT), Data Science (DS) and its sub-field Big Data (BD), Artificial
Intelligence (AI), Integrated STEM Thinking (ISTEMT), and Smart
Education) as Big Concepts (further BCs) either intuitively from the
context, or his/her previous knowledge. Note that we define ISTEMT as
a compound (ISTEMT = Computational Thinking (CT) + Design Thinking
(DT) + Data Thinking (DtT) + Scientific Thinking (ST)). In other words, it
aggregates all known so far movements, i.e., CT movement [Win06,
DT19, MRR+22], DT movement [LSG+19], and DtT movement
[BVW+21, MRR+22] supplemented by Scientific Thinking.
What is the role of STEM, CS, and robotics? Whether, or not they are
BCs? Of course, in our vision, they are BCs too; however, their role is
different. For developing the concept and content of this new book, we
put STEM-driven Computer Science Education at the centre. We treat this
paradigm (STEM, CS, and Robotics) as our object here for investigation,
research, and practice from the perspective of evolutionary changes
under the impact of the content taken from the field of BCs such as IoT,
DS, and AI. In other words, this book aims to show how BCs can
contribute to extending, enforcing, and enhancing STEM-driven CS
education at the K–12 level at least in this decade of the twenty-first
century. Note that regarding the term ‘Big Concept’, one can meet in the
literature the term ‘Big Ideas’ [RAE16, Boo19, MRR+22] or “Big
Computer Science” [Cer17] used in similar contexts.
At the very beginning, we need to motivate and explain yet two
important aspects. (1) What is the way for introducing BCs into this
educational paradigm? (2) How to measure and evaluate the
evolutionary changes based on those concepts? Regarding the first
aspect, it is worth reminding how it is difficult to introduce new
courses without governmental initiatives and support into the K–12
curriculum due to administrative and other reasons. A more reasonable
way is to change and extend the already existing relevant courses
because this largely depends on the teacher’s readiness and vision. Of
course, this way has own restrictions on the extent and scope of the
new content. We argue that it is impossible to cover a large portion of
the relevant content for any BC within the existing courses such as
Computer Programming, Robotics, etc. Therefore, for extending the
existing courses, typically we rely on introductive and more engaging
themes for the relevant BC. Those need to be adapted to the student’s
level and their previous knowledge. We admit the relevance of BCs to
STEM-CS education. Note that, in our case, some BCs (ISTEMT/CT,
Smart Education) do not have specific content. We argue that it is
possible to deliver these concepts by providing teaching content
regarding other BCs. In addition, we will describe the evolutionary
model in this chapter as a framework for systematically introducing
BCs throughout this book.
Regarding the second aspect, we have developed and researched the
integrated STEM-CS skills model [BŠK+22]. It covers multiple skills (CT
skills, design thinking, scientific thinking, and other skills) and
competencies integrated into a unified model needed for learners to
provide self-adaptation and self-preparation for managing and
overcoming the challenges in the twenty-first century. This model
serves as an instrument to evaluate the obtained new knowledge based
on BCs.
The aim of this chapter is (i) to outline the context of our research
and (ii) to motivate and develop the evolutionary model for change
based on BCs along with guidelines for implementing this model so that
it would be possible to formulate tasks and content of the entire book.
Tasks of this chapter are:
(i) Definition of education evolution. (ii) Characterisation of basic
attributes of computing evolution. (iii) Evolution of integrated STEM
education, (iv) Short glance to computational thinking skills. (v)
Development of the evolutionary model explaining the way we
introduce BCs into STEM-driven CS education. (vi) Formulation of
objectives, content, and contribution of our research.
learners
b Peeragogy is the pedagogy based on the management of collaborative
learning
c Cybergogy is the pedagogy based on using cyber technologies
d IoT—Internet of Things , AI—Artificial Intelligence, BD—Big Data, LA
—Learning Analytics
Adapted from [MNN+21]
(1.4)
In formula (1.4), the sign ‘*’ means the physical and logical
aggregation among indicated items. This model defines the core
components of the IoT in a very simplified manner. Note that we
presented a short vision of computing evolution from the perspective of
our previous [ŠD12, ŠB18] and current research. One can find more
extensive research on this topic in [DT19, TD16], for example, from the
perspectives of the evolution of computational thinking.
In summary, we conclude. (1) Computing/CS evolves with ever-
growing acceleration and impacts society. (2) This evolution has
resulted in the continuous complexity growth of computing systems.
(3) CS evolution can be characterised by the efforts to manage the
complexity growth by adding ever-higher abstraction levels. (4) All
citizens, especially the young generation, have to be prepared to
understand and accept those changes. (5) In this regard, STEM-driven
CS education is the relevant approach to learning for understanding,
accepting, and implementing the newest achievements in computing.
(6) Broadly, this is about skills and knowledge the young generation
needs in the twenty-first century for “unlocking a Reskilling
Revolution” [WEF21].
Tribe 2. Eryonidea.
Tribe 3. Peneidea.[135]
The third legs are chelate except in genera in which the legs are
much reduced. The third maxillipedes are seven-jointed, the second
maxillipedes have normal end-joints, and the first maxillipedes are
without a lobe on the base of the exopodite. The pleura of the first
abdominal segment are not overlapped by those of the second. The
abdomen is without a sharp bend. The branchiae are usually not
phyllobranchs.
Fam 1. Peneidae.—The last two pairs of legs are well developed,
and there is a nearly complete series of gills. Cerataspis,[136] a pelagic
form. Parapeneus, Peneus, Aristaeus, etc.
Fam. 2. Sergestidae.—The last or last two pairs of legs are
reduced or lost. The gill-series is incomplete or wanting. Sergestes
possesses gills, and the front end of the thorax is not greatly
elongated. Lucifer has no gills, and the front of the thorax is greatly
elongated, giving a very anomalous appearance to the animal. All the
members of this family are pelagic in habit.
Fam. 3. Stenopodidae.—One or both legs of the third pair are
longer and much stouter than those of the first two pairs. On a
number of small anatomical points this family, including the littoral
genus Stenopus from the Mediterranean and other warmer seas and
Spongicola commensal with Hexactinellid sponges from Japan, is
separated by some authors in a Tribe by itself.
Tribe 4. Caridea.
The third legs are not chelate. The third maxillipedes are 4–6
jointed, the end-joint of the second maxillipede nearly always lies as
a strip along the end of the joint before it, and the first maxillipedes
have a lobe on the base of the exopodites. The pleura of the second
abdominal segment overlap those of the first. The abdomen has a
sharp bend; the branchiae are phyllobranchs.
Fam. 1. Pasiphaeidae.—In this family the end-joint of the
second maxillipedes is normally formed, and exopodites are usually
present on all the thoracic limbs. Rostrum small or wanting. Rather
numerous genera are known, most of which inhabit the deep sea,
though a few come into the littoral zone. Pasiphaea chiefly in the
deep sea, Leptochela in the tropical littoral zone.
Fam. 2. Acanthephyridae.—The end-joint of the second
maxillipede is modified as in other Caridea, and the rostrum is very
strong and serrate, but in the presence of exopodites, and in the form
of the mouth-parts, this family agrees with the preceding. It is also a
characteristic deep-sea family. Acanthephyra, Hymenodora,
Nematocarcinus, etc.
Fam. 3. Atyidae.—This is an entirely fresh-water family,
especially characteristic of the rivers and lakes of the tropics, some of
the forms being exceedingly large and taking the place of the
Crayfishes in these waters. Characteristic of this family is the fact
that the fingers of the chelae are spoon-shaped, and carry peculiar
tufts of bristles. Exopodites are present on the thoracic limbs of some
of the genera (Troglocaris, Xiphocaris from Australia and the Malay
Islands, Atyephyra from S. and W. Europe), but are absent in others.
Caridina, widely spread and common in Indo-Malay and Africa;
Atya from West Indies, West Africa, and Pacific Islands.
Fam. 4. Alpheidae.[137]—The exopodites are absent, and the
rostrum is absent or very feeble. The chelae are powerful, and usually
very asymmetrically developed. Alpheus has an enormous number of
species which live chiefly in the tropical seas, where they haunt
especially the coral-reefs, making their homes among the coral or in
sponges, etc. Although occurring in the Mediterranean they
penetrate very rarely into colder seas.
Fam. 5 Psalidopodidae.—This family, characterised by the
absence of chelae on the second thoracic limbs, which carry instead a
terminal brush of hairs, and by the rudimentary condition of the
eyes, is represented by the genus Psalidopus from the deep waters of
the Indian Ocean.
Fam. 6. Pandalidae.—The first thoracic limb is without chelae,
only six-jointed. The rostrum is large and toothed. The genus
Pandalus has numerous representatives in the northern littoral, P.
annulicornis being one of the prawns most commonly met with in
the fish-markets.
Fam. 7. Hippolytidae.—The first and second thoracic limbs
bear chelae, the carpus of the second being divided into two or more
segments. The first pair of chelae are not distinctly stronger than the
second. Virbius has many species in the littoral zone of all seas, and
one species, V. acuminatus, is pelagic. Hippolyte also has numerous
littoral forms distributed all over the world, but chiefly in the arctic
or subarctic seas. H. varians, common on the English coasts, shows
interesting colour-reactions to its surroundings.[138]
Fam. 8. Palaemonidae.—
The first two pairs of legs are
chelate, the carpus of the second
not being subdivided. Palaemon
serratus, a very common prawn
in the British littoral.
Palaemonetes in the brackish and
fresh waters of Europe and N.
America.
Fam. 9.
Glyphocrangonidae.—The first
pair of legs are subchelate, the
carpus of the second pair is
subdivided, and the rostrum is
long. Glyphocrangon (Fig. 110)
with numerous species entirely
confined to deep water.
Fam. 10. Crangonidae.—
Fig. 110.—Glyphocrangon spinulosa,
from the right side, × 1. (From an
The first pair of legs are
original drawing prepared for Professor subchelate, the carpus of the
Weldon.) second pair is not subdivided,
and the rostrum is short.
Crangon vulgaris is the common
Shrimp of the North Sea.
Tribe 5. Loricata.
Tribe 6. Thalassinidea.
Sub-Order 2. Anomura
In this division are included the so-called Hermit-lobsters and
Hermit-crabs, in which the condition of the abdomen is roughly
intermediate between that of the Macrura and that of the Brachyura.
It is not much reduced in size, and the pleopods of the sixth pair are
fairly well developed, but it is usually carried flexed towards the
thorax, and is never a powerful locomotory organ as in the Macrura.
The antennal scale, if present at all, is a mere spine, not the large
leaf-like structure of the Macrura; and there is never a partition
between the two first antennae as in the Brachyura.
The last or last two pairs of pereiopods are reduced, and are turned
on to the dorsal surface or carried inside the branchial chamber; but
this curious character is met with again in certain Brachyura
(Dromiacea and Oxystomata).
Tribe 1. Galatheidea.[140]
Tribe 2. Hippidea.
Tribe 3. Paguridea.[142]
The ordinary Hermit-crabs, common on the English as on every
coast, are characterised by the fleshy asymmetrical abdomen from
which all the hard matter has disappeared, and which is carried
tucked away in an empty Gasteropod shell. The abdomen is spirally
wound in accordance with the shape of the shell, and a firm
attachment is effected by means of the sixth pair of pleopods,
especially that of the left side, which is fashioned into the form of a
hook and is curled round the columella of the shell; this attachment
is so secure that in trying to pull a Hermit-crab out of its shell the
body is torn apart before the hold gives way. The other pleopods are
in a much reduced condition, being generally altogether absent from
the right side of the abdomen, and often greatly reduced on the left
side, especially in the male, though in the female they are still used
for the attachment of the eggs.
The last two pereiopods are much reduced and are concealed
inside the shell, which they help to carry. The great chelae are usually
asymmetrically developed, that on the right side being much larger
than that on the left, and often serving the purpose of shutting the
entrance to the shell when the crab is withdrawn inside.
The constant association of a large group of animals like the
Hermit-crabs with the appropriated empty houses of another group
is sufficiently curious, but it does not stop there. In almost every case
there are present one or more Sea-anemones growing on the outside
of the shell, and each kind of Hermit-crab generally carries a special
kind of Anemone. Thus at Plymouth, Eupagurus bernhardus is
generally symbiotic with Sagartia parasitica, or else with a colony of
Hydractinia echinata, while E. prideauxii is usually associated with
Adamsia palliata. In the latter case the shell is frequently absorbed,
so that the Anemone comes to envelop the crab like a blanket.
Instead of Anemones carried turret-like and imposing aloft, or
enveloping the inmate of the shell like a blanket, some of the
Hermits have Sponges, an unexpected association; and it is a
common sight at Naples to find the little red round Sponge,
Suberites, running around animated by its Hermit within. It is held
that Anemone and crab mutually assist one another, that the
Anemone stings the crab’s enemies, and that the Hermit-crab carries
the Anemone to new feeding-grounds. It is also said that when a crab
grows too big for its shell, and is forced to seek another, it persuades
the Anemone to loosen its attachment to the deserted shell and to be
transplanted to the new one, and that there is something mesmeric
in its power, because nobody else can pull an Anemone off a shell
without either cutting it off at the base or tearing it to pieces. Other
animals as well sometimes enter into this partnership. At Plymouth a
Polychaet worm, Nereis fucata, frequently inhabits the Whelk’s shell,
together with Eupagurus bernhardus, and puts out its head for a
share of each meal; and at Naples the Amphipod Lysianax punctatus
is almost always present in the shells of Eupagurus prideauxii.
Besides the ordinary twisted Pagurids which inhabit Gasteropod
shells, there are a few which preserve the symmetry of the body. The
interesting Pylocheles miersii[143] (Fig. 118), taken by the
Investigator in the Andaman Sea at 185 fathoms, inhabits pieces of
bamboo; it is perfectly symmetrical, with well-developed pleopods
and symmetrical chelae, which, when the animal is withdrawn,
completely shut up the entrance to its house (Fig. 118, A).
It is doubtful whether this animal ever inhabited a spiral shell or
not in its past history; but there is no doubt that a number of peculiar
crabs, which caused the older systematists much trouble, are
Pagurids, derived from asymmetrical shell-haunting ancestors that
have secondarily taken to a different mode of life, and lost, or
partially lost those characteristics of ordinary Hermit-crabs which
are associated with life in a spiral shell. These are the Lithodidae and
the “Robber-crab,” Birgus latro, of tropical coral islands.
Although the Robber-crab and the Lithodidae bear a certain
superficial resemblance to one another in that they lead a free
existence, and have reacquired to a great extent their symmetry, yet
it is clear that they have been independently derived from different
groups of asymmetrical Hermit-crabs, and that their resemblance to
one another is due to convergence.
Birgus latro (Fig. 119), a gigantic crab, frequently over a foot in
length, lives on land, and inhabits the coasts of coral islands in the
Indian and Pacific Oceans where cocoa-nut trees grow. It feeds on
the pulp of the cocoa-nut, which it extracts by hammering with its
heavy chela on the “eye-hole” until room is made for the small chela
to enter and extract the pulp. There is not the slightest doubt that the
animal often ascends the cocoa-nut trees for the purpose of picking
the nuts, a fact illustrated by a fine photograph by Dr. Andrews,
exhibited in the Crustacean
Gallery in the Natural History
Departments of the British
Museum. It uses the husk of the
nut to line its burrow, and it is
said to have the habit of putting
its abdomen into the nut-shell for
protection and carrying it about
with it. Owing to its terrestrial
mode of life, the branchial
chamber is highly modified, being
divided into two portions—a
dorsal space, the lining of which
is thrown into vascular ridges and
folds for aerial respiration, and a
lower portion where the
rudimentary branchiae are
situated. Although the Robber-
crab lives ordinarily on land, it
must be supposed that these
branchiae are of some service; the
young are hatched out as ordinary
Zoaeas in the sea, and go through
a pelagic existence before seeking
the land. At the present time the Fig. 118.—Pylocheles miersii, × 1. A,
Robber-crab is confined to the End view of a piece of mangrove or
Pacific and the islands of the bamboo, the opening of which is closed
by the great chelae (c) of the Pagurid;
Indian Ocean, wherever the B, the animal removed from its house.
cocoa-nut grows. It seems, (After Alcock.)
however, that its association with
the cocoa-nut is a comparatively
modern one. Mr. C. Hedley, of Sydney, who has had great experience
of the Pacific Islands, informs me that the cocoa-nut is not, as is
usually supposed, a native of these coral islands, but has been
introduced, probably from Mexico, by the Polynesian mariners
before the discovery of America by Columbus. Before the
introduction of the cocoa-nut the Robber-crab must have fed on
some other tree, possibly the Screw Pine, Pandanus.